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Turonian

The Turonian is a geologic stage and chronozone in the Upper Cretaceous Series of the international geologic timescale, representing a period of approximately 4.1 million years from 93.9 ± 0.2 Ma to 89.8 ± 0.3 Ma.^[1] It follows the Cenomanian and precedes the Coniacian, encompassing a time of elevated global temperatures, high sea levels, and significant marine transgression that expanded epicontinental seas across much of the world's continents.^[2] The base of the Turonian is formally defined by the first occurrence of the ammonite species Watinoceras devonense at the Global Boundary Stratotype Section and Point (GSSP) located in Bed 86 of the Bridge Creek Limestone Member, near Pueblo, Colorado, USA (38°16’56″N, 104°43’39″W).^[3] This boundary coincides with the Cenomanian-Turonian Oceanic Anoxic Event 2 (OAE2), a major perturbation in the global carbon cycle characterized by widespread marine anoxia, a positive carbon isotope excursion, and enhanced organic carbon burial, which marks the Cenomanian-Turonian extinction event, a significant perturbation affecting planktonic foraminifera, ammonites, and other marine taxa.^[4] The event, dated to around 93.9 Ma, was driven by volcanic outgassing, eutrophication, and sea-level rise, leading to stratified oceans and black shale deposition in basins worldwide.^[5] During the Turonian, diverse marine ecosystems flourished, with key index fossils including ammonites such as Collignoniceras woollgari for the middle substage, inoceramid bivalves like Mytiloides species, and microfossils such as the foraminifer Helvetoglobotruncana helvetica and nannofossil Microstaurus chiastius (marking its last occurrence near the base).^[3] On land, the stage saw the evolution and diversification of dinosaurs, including early ceratopsians and hadrosauromorphs in North America, amid a greenhouse climate that supported lush coastal vegetation and angiosperm dominance.^[2] The Turonian is subdivided into lower, middle, and upper substages, with boundaries often defined by ammonite biozonations and chemostratigraphic markers, reflecting dynamic paleoenvironments from shallow shelves to deep basins.^[6]

Stratigraphy

Definition and Naming

The Turonian is a stage in the geologic timescale, defined within the Upper Cretaceous series of the Cretaceous system. It represents the second chronostratigraphic stage of the Late Cretaceous epoch, succeeding the Cenomanian stage and preceding the Coniacian stage.^[7] The name "Turonian" was coined by the French paleontologist Alcide d'Orbigny in 1842, derived from Turonia, the ancient Roman designation for the Touraine region in central France, centered around the city of Tours, where characteristic Cretaceous strata, including the Craie Chloritée, were studied.^[8] d'Orbigny's initial classification divided the Upper Cretaceous into two broad stages—the Turonian below and the Senonian above—based primarily on assemblages of ammonites and other fossils observed in French sections.^[8] By 1851, he refined the Turonian to encompass beds with specific ammonite faunas, such as those dominated by species of the genus Mammites, and associated rudistid bivalves, establishing it as a distinct biostratigraphic unit.^[8] The concept of the Turonian stage evolved further in the mid-19th century through the work of German stratigrapher Albert Oppel, who built on d'Orbigny's framework by introducing zonal divisions based on the vertical ranges of index fossils, enhancing the precision of stage correlations across Europe.^[9] This zonal approach, initially applied to Jurassic sequences, was extended to the Cretaceous, including the Turonian, facilitating international standardization during the late 19th and 20th centuries.^[9] In the current International Chronostratigraphic Chart, the Turonian spans from 93.9 ± 0.2 million years ago to 89.8 ± 0.3 million years ago, reflecting radiometric calibrations integrated with biostratigraphy and cyclostratigraphy.^[7] The base of the stage is formally defined by the Global Stratotype Section and Point (GSSP) at Pueblo, Colorado, USA.^[8]

Boundaries

The lower boundary of the Turonian stage is defined by the first appearance datum (FAD) of the ammonite Watinoceras devonense (Wright & Kennedy, 1981).^[8] This boundary coincides with the base of Bed 86 in the Bridge Creek Limestone Member of the Greenhorn Limestone Formation.^[8] The Global Stratotype Section and Point (GSSP) for the base of the Turonian is located at the western end of the Denver and Rio Grande Western Railroad cut, near the north boundary of the Pueblo Reservoir State Recreation Area, west of Pueblo, Colorado, USA (coordinates: 38°16'56"N, 104°43'39"W).^[8] This GSSP was ratified by the International Commission on Stratigraphy (ICS) and approved by the International Union of Geological Sciences in September 2003.^[8] The upper boundary of the Turonian stage corresponds to the base of the Coniacian stage and is defined by the FAD of the inoceramid bivalve Cremnoceramus deformis erectus (Meek, 1871), which is synonymous with Cremnoceramus rotundatus (sensu Tröger non Fiege).^[10] This marks the transition from the uppermost Turonian Inoceramus labiatus Zone to the basal Coniacian C. deformis erectus Zone.^[10] The GSSP for this boundary, ratified in 2021, is situated at the base of Bed 46 in the Salzgitter-Salder quarry, Lower Saxony, Germany (coordinates: 52°07'28"N, 10°19'46"E), within the upper part of the Grauweisse Wechselfolge succession.^[10] Global correlation of the Turonian boundaries relies on integrated biostratigraphy, chemostratigraphy, and magnetostratigraphy.^[11] The δ¹³C isotope stratigraphy, particularly positive excursions such as the Cenomanian-Turonian event at the base and the 'i5' and 'i6' events near the top, provides a robust chemostratigraphic framework for synchronization across sections.^[10] Magnetostratigraphy ties the stage to polarity Chron C34n, with the upper boundary occurring within the later part of this long normal chron.^[12] Biostratigraphic markers, including ammonites (e.g., Forresteria petrocoriensis at the top) and foraminifera (e.g., Dicarinella concavata), further aid in precise alignment.^[10] Recognition of Turonian boundaries faces challenges due to regional facies variations, particularly in non-Tethyan areas where marine regressions and disconformities obscure biostratigraphic markers.^[13] In such realms, tectonic influences and local hiatuses can shift the apparent position of index fossils, necessitating auxiliary correlation tools like isotopes to resolve discrepancies.^[13]

Subdivision

The Turonian stage is informally divided into three substages based primarily on ammonite biostratigraphy from European reference sections: the Lower, Middle, and Upper Turonian. These divisions facilitate finer stratigraphic resolution within the overall stage duration of about 4.1 million years. The lower-middle substage boundary is defined by the first occurrence of the ammonite Collignoniceras woollgari, with a proposed Global Stratotype Section and Point (GSSP) at Bed 120 of the Rock Canyon Anticline section near Pueblo, Colorado. The middle-upper boundary lacks a formal definition.^[6]^[14] In the Tethyan domain, the stage is delineated by 10–12 ammonite biozones, which provide a standard for correlation; notable examples include the basal Watinoceras coloradoense Zone, the early Mammites nodosoides Zone, and the mid-stage Collignoniceras woollgari Zone.^[15] Complementary inoceramid bivalve biozonation enhances global applicability, with key intervals such as the Mytiloides labiatus Zone in the lower Turonian and the Inoceramus perplexus Zone spanning the middle to upper parts. Regional variations in zonation reflect local faunal assemblages; in North America, particularly the Western Interior Basin, subdivisions incorporate rudist bivalves alongside foraminifera and ammonites for carbonate platform correlations.^[16] Faunal provincialism between the cooler Boreal and warmer Tethyan realms leads to distinct ammonite and bivalve assemblages, complicating direct inter-realm matching without auxiliary methods.^[17] Precise age assignments within substages rely on integrated correlation tools, including cyclostratigraphy that identifies Milankovitch cycles in sedimentary rhythms and chemostratigraphy using carbon isotope profiles to align sections globally.^[2]

Paleoenvironment

Paleogeography

During the Turonian stage of the Late Cretaceous, approximately 93.9 to 89.8 million years ago, the global continental configuration featured ongoing fragmentation of the southern supercontinent Gondwana, with South America progressively separating from Africa as the South Atlantic Ocean basin began to open wider.^[18] Laurasia remained relatively stable as a northern landmass comprising North America, Europe, and Asia north of the Tethys, though rifting initiated in the proto-North Atlantic between Greenland and Eurasia, marking the early stages of continental breakup.^[19] Meanwhile, the Indian subcontinent continued its northward drift from its Gondwanan position toward the Eurasian margin, traversing the widening Tethys Ocean at rates contributing to the eventual Himalayan orogeny.^[20] The proto-Atlantic, or Central Atlantic Ocean, expanded as a result of seafloor spreading between the Americas and Africa, while the Tethys Ocean dominated the eastern hemisphere as a broad equatorial seaway connecting the Atlantic to the Pacific.^[19] Subduction along the Pacific margins, particularly the Farallon Plate beneath South America, drove the Andean orogeny, with compressive tectonics building proto-Andean highlands and influencing sediment dispersal across western Gondwana.^[18] In the Caribbean region, minor transpressional events deformed island-arc terranes, associated with oblique convergence and the early emplacement of the Caribbean plateau.^[21] Overall plate motions during this interval averaged 2–5 cm/year, reflecting a period of moderate tectonic activity following earlier Jurassic-Cenomanian reorganizations.^[22] High global sea levels facilitated extensive epicontinental flooding of continental interiors, creating shallow marine seaways that subdivided landmasses.^[23] In North America, the Western Interior Seaway reached its peak extent, stretching over 2,000 km from the Gulf of Mexico to the Arctic and bisecting the continent into eastern and western highlands within a subsiding foreland basin.^[23] Europe's Anglo-Paris Basin hosted a broad, chalk-dominated shelf sea connected to the Tethys, with marine transgressions depositing fine-grained carbonates across southern England and northern France.^[24] In Africa, the Saharan platforms experienced widespread inundation, forming shallow epicontinental seas that linked the Tethys to southern Gondwana and supported carbonate platform development in regions like Tunisia and Algeria.^[25]

Climate and Sea Levels

The Turonian stage represented a peak of the mid-Cretaceous greenhouse climate, characterized by elevated atmospheric CO₂ concentrations estimated at 1450–2690 ppmv based on stable isotope analysis of paleosol calcite from mid-Turonian deposits in Israel.^[26] Tropical sea surface temperatures (SSTs) reached averages near 35°C in the equatorial Atlantic, as reconstructed from the TEX₈₆ organic geochemical proxy in sediments from Demerara Rise.^[27] Polar regions remained ice-free, with evidence from stable oxygen isotope (δ¹⁸O) records in exquisitely preserved Tanzanian foraminifera showing no glacial excursions and indicating sustained high-latitude warmth. This warm regime contributed to global thermal expansion and minimal ice volume, fostering a world without perennial polar ice caps.^[28] Temperature reconstructions during the Turonian relied heavily on oxygen isotope proxies from marine microfossils and bivalves. Planktonic foraminiferal δ¹⁸O data from mid- to late Turonian sediments in the Yezo Basin (paleolatitude ~44°N) indicate mean annual SSTs of 26–29°C, with seasonal variations up to 7°C derived from bivalve shell analyses.^[29] Rudist bivalve shells from shallow-water Tethyan environments provide evidence of even higher temperatures, with Early Turonian SSTs reaching 41–45°C in low-latitude settings based on δ¹⁸O sclerochemistry.^[30] These proxies collectively point to a thermal maximum in the Early Turonian, transitioning to slightly cooler conditions by mid-stage, consistent with broader mid-Cretaceous warming trends.^[31] Sea levels during the Turonian initiated at a global highstand of approximately 200–250 m above present-day mean level, driven primarily by thermal expansion of seawater and the absence of significant polar ice volumes.^[32] This eustatic highstand facilitated widespread epicontinental flooding, particularly in tectonically stable regions. Mid-Turonian dynamics included a notable regression, with a eustatic fall exceeding 75 m linked to the Round Down carbon isotope excursion around 92.3 Ma, resulting in regional hiatuses and lowstands.^[2] This was followed by a transgression in the late Turonian, restoring higher sea levels and promoting renewed marine inundation.^[2] Ocean circulation in the Turonian featured restricted deep-water ventilation, particularly in the Tethys Ocean, where tectonic configuration and high temperatures promoted water column stratification.^[33] In the northwestern Tethys, stable thermoclines and prolonged thermal stratification led to reduced mixing between surface and deeper waters, as inferred from nannofossil assemblages and geochemical indicators in Algerian sections.^[34] Elevated pCO₂ levels implied potential ocean acidification, with hints from carbonate dissolution patterns and reduced calcification in marine records, though direct pH proxies remain limited.^[35]

Biodiversity

Marine Life

The Turonian stage of the Late Cretaceous witnessed a rich diversity of marine life, characterized by high faunal turnover and adaptation to varying oceanic conditions. Dominant among nektonic organisms were ammonites, which exhibited significant generic diversity across global basins, with over 100 genera recorded in Turonian assemblages, including prominent forms like Acanthoceras and Neocardioceras.^[36]^[37] These cephalopods served as key index fossils for biostratigraphy, occupying diverse habitats from epicontinental seas to deeper slopes, and their coiled shells facilitated buoyancy control in the water column.^[38] Benthic communities were heavily influenced by bivalves, particularly inoceramids such as Mytiloides and Inoceramus, which formed extensive shell beds in chalk and marl deposits, often dominating the seafloor in nutrient-rich environments.^[39]^[40] These large, thin-shelled mollusks thrived in suspension-feeding niches, contributing to sediment accumulation and serving as substrates for epifauna, with their prismatic calcite shells preserving well in dysaerobic settings.^[41] In shallow tropical platforms, rudist bivalves emerged as primary reef constructors, supplanting scleractinian corals which experienced a marked decline in abundance and framework-building roles during this interval.^[42] Species like Durania formed dense thickets and bioherms in warm, clear waters, creating wave-resistant structures through their conical, cemented valves that promoted carbonate deposition.^[43] Rudist-dominated reefs supported associated communities of algae and smaller invertebrates, reflecting a shift toward bivalve-centric ecosystems in the Tethyan realm.^[44] Planktonic foraminifera, exemplified by Helvetoglobotruncana helvetica, were abundant in open marine settings, often concentrated in greensand formations indicative of high productivity.^[45] Among nekton, early mosasaurs appeared in nearshore habitats, preying on fish and ammonites, while sharks like Ptychodus exploited durophagous niches by crushing thick-shelled prey such as inoceramids.^[46]^[47] Fish diversity included predatory enchodontids, which filled mid-trophic roles in food webs across epicontinental seas.^[48] Ecologically, Turonian oceans featured elevated productivity in upwelling zones, fueling phytoplankton blooms that supported diverse trophic levels, while benthic assemblages adapted to dysaerobic bottom waters through low-oxygen tolerant taxa like opportunistic foraminifera and infaunal bivalves.^[49]^[50] These conditions promoted opportunistic recolonization following transient anoxic episodes, enhancing overall marine resilience.^[51]

Terrestrial Life

During the Turonian, terrestrial ecosystems witnessed a significant radiation of angiosperms, which began to dominate landscapes previously held by gymnosperms and ferns. Fossil evidence from North American and European deposits shows that early angiosperms, including magnoliid-like flowers such as Cronquistiflora and Detrusandra, exhibited cupulate structures and tricolporate pollen, indicating adaptations for insect pollination and dispersal in forested environments.^[52] Palms, represented by stem fossils in French localities, appeared as understory or riparian elements, contributing to increasingly diverse woodland canopies.^[53] Pollen records from the Raritan Formation in New Jersey reveal a marked diversification of angiosperm taxa, with over 20 genera including laurels and plane trees, signaling a shift toward angiosperm-dominated vegetation in floodplain and coastal settings.^[54] Conifers, such as cupressaceae with Taxodium-like affinities, persisted in swampy and cooler habitats, while ferns like matoniaceous and dicksoniaceous species formed the understory, gradually declining as angiosperms encroached.^[55] Among vertebrates, dinosaurs were prominent components of Turonian terrestrial and semi-aquatic habitats. Ornithischian dinosaurs included early ceratopsians such as Zuniceratops from New Mexico and early hadrosauroids, such as Jeyawati rugosa from the Zuni Basin of New Mexico, which featured robust limbs and dental batteries precursor to later duck-billed forms, suggesting herbivorous grazing in floodplain meadows.^[56] Theropod dinosaurs encompassed spinosaurids like Spinosaurus, known from North African formations spanning the Cenomanian-Turonian boundary, with elongated snouts and sail-backed structures adapted for piscivory in riverine systems. Sauropod dinosaurs, once dominant, continued to decline in diversity and abundance, with only scattered titanosaur remains in European and Asian localities, reflecting a broader Late Cretaceous trend toward ornithischian dominance in herbivore guilds.^[57] Other vertebrates diversified in aerial and small terrestrial niches. Pterosaurs, including early members of Pteranodontia such as Cimoliopterus relatives from European chalk deposits, soared over coastal plains with wingspans exceeding 5 meters, preying on fish and insects.^[58] Early birds, represented by ornithurine taxa like Tingmiatornis arctica from high-latitude Arctic sediments, displayed mosaic features including keeled sterna and strong flight capabilities, occupying arboreal and shoreline foraging roles.^[59] Mammals remained small and inconspicuous, limited to multituberculates such as Bryceomys from Utah's Dakota Formation, which exhibited specialized teeth for gnawing seeds and insects in underbrush habitats.^[60] Insects and other invertebrates thrived in Turonian forests and freshwater systems, supporting ecosystem dynamics. Diverse beetles and early ants, preserved in Burmese and French ambers, included herbivorous and predatory forms that interacted with burgeoning angiosperm flora. Termites constructed mounds in wooded areas, as evidenced by coprolites from western France, facilitating soil aeration and nutrient cycling in humid environments.^[61] Freshwater ecosystems hosted baenid turtles with robust shells suited to riverine predation, alongside crocodilians such as goniopholidids from Japanese localities, which ambushed prey in deltas and lakes.^[62] These groups underscored the interconnectedness of terrestrial and aquatic biomes during this stage.

Significant Events

Oceanic Anoxic Event 2

The Oceanic Anoxic Event 2 (OAE2) transpired at the Cenomanian-Turonian boundary, marking the onset of the Turonian stage around 93.9 million years ago. This event spanned approximately 0.5 to 1 million years, from roughly 94.5 to 93.9 Ma, and is characterized by widespread marine anoxia that disrupted global ocean oxygenation.^[63] A prominent positive carbon isotope excursion in δ¹³C values, reaching up to +4‰, defines the event and is prominently recorded in organic-rich sediments known as the Bonarelli Level.^[63] The primary drivers of OAE2 included intense volcanic activity from large igneous provinces, such as the Caribbean Large Igneous Province, Ontong Java Plateau, and Kerguelen LIP, which released vast quantities of CO₂, inducing rapid global warming and enhanced greenhouse conditions.^[63]^[64] Orbital forcing, particularly through eccentricity and precession cycles, contributed by promoting periods of low seasonal insolation variation, which fostered water column stratification and reduced vertical mixing in oceans.^[65] Additionally, eutrophication arose from an accelerated hydrological cycle that increased nutrient delivery to marine settings, boosting primary productivity and organic matter flux, ultimately leading to oxygen depletion and stratified, anoxic bottom waters across vast oceanic realms.^[66] Globally, OAE2 is evidenced by the deposition of organic-rich black shales in multiple ocean basins, including the Tethys, Atlantic, and Pacific, with total organic carbon contents often exceeding 5-10% in these layers.^[63] The Livello Bonarelli in the Umbria-Marche region of Italy exemplifies this signature as the type section, featuring a ~1-meter-thick interval of laminated black shales that reflect the peak of anoxic conditions.^[65] OAE2 triggered substantial biotic turnover, with approximately 25-30% species extinction among planktonic foraminifera, particularly affecting deeper-dwelling, K-selected taxa like Rotalipora, while opportunistic, r-selected species such as Heterohelix exhibited preferential survival and dominance in post-event assemblages. Similarly, ammonites experienced significant genus-level declines linked to expanded oxygen minimum zones that disrupted their habitats, though regional variations in severity were noted across basins like the Tethys and Western Interior Seaway.^[67]

Eustatic Fluctuations

The Turonian stage records a series of third-order eustatic cycles that shaped its stratigraphic architecture, with depositional sequences typically comprising lowstand, transgressive, and highstand systems tracts. These cycles began with an early Turonian highstand, reflecting peak global sea levels during the initial phase of the stage, followed by a prominent mid-Turonian lowstand known as the Turonian regression, centered around sequence boundary KTu 4 at approximately 91.8 Ma. This regression marked a significant relative sea-level fall, after which a late Turonian transgression re-established deeper marine conditions. The amplitudes of these third-order fluctuations varied regionally but generally ranged from 50 to 100 meters, as evidenced by backstripping analyses and sequence boundary correlations across multiple basins.^[68]^[69] The primary drivers of these eustatic variations were non-glacial in nature, consistent with the prevailing hot greenhouse climate that minimized glacio-eustasy due to limited continental ice volumes. Thermal subsidence of ocean basins and fluctuations in sediment supply to continental margins played key roles in modulating relative sea levels, with additional contributions from dynamic topography and possible groundwater unloading during lowstands. Long-term ocean basin volume changes provided a backdrop for these shorter-term oscillations, while regional tectonics amplified local expressions without dominating the global signal.^[68] Stratigraphic records of these fluctuations are prominent in the Western Interior Basin, where multiple unconformities and hiatuses interrupt middle to upper Turonian deposits, such as those within the Carlile Shale Formation, reflecting erosion and non-deposition during the mid-Turonian lowstand. In the Middle East, the Arabian Carbonate Platform exhibits drowning events during the early Turonian transgression, characterized by abrupt shifts from shallow-water reefal and lagoonal facies to pelagic marls and shales, driven by rapid sea-level rise and subsidence that submerged platforms below optimal carbonate production depths.^[69] These eustatic cycles profoundly influenced depositional environments, causing facies belts to migrate laterally and vertically; lowstands facilitated progradation of terrestrial clastics and shallow marine sands onto shelves, while transgressions promoted retrogradation and the establishment of open marine shales and carbonates. The rhythmic patterning of these changes aligns with Milankovitch orbital forcing, particularly through eccentricity cycles, where bundling of ~100-kyr short-eccentricity oscillations within ~405-kyr long-eccentricity modulations drove periodic sea-level variability and associated carbon cycle perturbations.^[69]

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Aug 7, 2025 · Bed 22 is the most inoceramid-rich interval. It yielded Mytiloides incertus (Jimbo), Mytiloides labi-. atoidiformis (Tröger), Mytiloides ...
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ammonites, inoceramid bivalves and stable carbon isotopes
juddii) and three Turonian (W. devonense, F. catinus, M. nodosoides) zones. The inoceramid study enables recognition of the Mytiloides puebloensis Zone, and ...
[41]
Shell composition, cryptic costae, complex composite molds, and ...
May 20, 2016 · In one such bivalve, Mytiloides ipuanus (Wellman, 1959) (Inoceramidae), the mineralized shell comprises an outer prismatic calcite layer and an ...<|separator|>
[42]
[PDF] Evolutionary trends in the epithecate scleractinian corals
Beginning with the Turonian, corals ... Cenozoic, deep-water. Development and decline of scleractinian epithecate corals. Transformations of scleractinian faunas.
[43]
Cenomanian - Turonian rudist assemblages and sequence ...
Jan 4, 2019 · The presence of the rudist rudstone, with in situ preserved shells, intercalated with lime mudstone and foraminiferal bioclastic wackestone ...
[44]
The Rise and Fall of Rudist Reefs | American Scientist
Reefs of the dinosaur era were dominated not by corals but by odd mollusks, which died off at the end of the Cretaceous from causes yet to be discovered.
[45]
Integrated foraminifera and δ13C stratigraphy across the ...
Feb 9, 2018 · Two foraminiferal zones were distinguished: Rotalipora ... Turonian epochs in the foraminiferal associations in northern temperate latitudes.
[46]
Mosasauridae) from the Turonian (Upper Cretaceous) of Kansas ...
Mosasaurs are extinct marine lizards that are commonly found in Coniacian–Maastrichtian (Late Cretaceous) deposits nearly worldwide but are rare in older rocks.Missing: enchodontids | Show results with:enchodontids
[47]
The Italian record of the Cretaceous shark, Ptychodus latissimus ...
We hypothesize that P. latissimus was a third-level predator occupying habitats with abundant thick-shelled prey, such as inoceramid bivalves and ammonites.
[48]
Field Guide, Part 2; Sharks and Bony Fish
Jul 22, 2013 · Enchodontids: Fish in the genus Enchodus were small to medium sized predators. Several species are well represented throughout the chalk and are ...
[49]
[PDF] The Cenomanian-Turonian Oceanic Anoxic Event, II ... - ResearchGate
Cenomanian-. Turonian beds occur within predicted areas of upwelling; however, most of the same areas that were marine before and after the Cenomanian-.
[50]
Microfossil Assemblages and the Cenomanian-Turonian (late ...
As a result, increasingly dysaerobic bottom waters developed within the Chalk Sea, and were responsible for progressive disappearances in the benthonic ...
[51]
Sea-level changes and geochemical anomalies across the ...
Relationships between geochemical anomalies, sea-level change and other events were studied in the Cenomanian–Turonian boundary interval in the Pecı́nov quarry ...Abstract · Introduction · Molluscan AssemblagesMissing: Challenges recognition
[52]
Two new fossil flowers of magnoliid affinity from the Late Cretaceous ...
Sep 1, 1998 · Two taxa of cupulate magnoliid fossil flowers, Cronquistiflora and Detrusandra, are described from the Late Cretaceous (Turonian, ...
[53]
Palms of the past: can morphometric phytolith analysis inform deep ...
Palm fossils are often used as evidence for warm and wet palaeoenvironments ... Similarly, palm stems, reported as early as the Turonian of France ...Materials And Methods · Results · Discussion<|control11|><|separator|>
[54]
TURONIAN FLORA OF NEW JERSEY, USA - DOAJ
This flora encompasses mosses, ferns, gymnosperms and angiosperms, and a ... as well as the oldest unequivocal monocot. Although the relationships of ...
[55]
Rise to dominance of angiosperm pioneers in European Cretaceous ...
Early angiosperm diversification: The diversity of pollen associated with angiosperm reproductive structures in Early Cretaceous floras from Portugal. Ann ...
[56]
A new basal hadrosauroid (Dinosauria: Ornithopoda) from the ...
The fossil record of Turonian dinosaurs from North America is extremely sparse, largely due to high sea levels and the inundation of much of the continent ...Missing: hadrosauroids | Show results with:hadrosauroids
[57]
The end of the sauropod dinosaur hiatus in North America
Sauropod dinosaurs were the most diverse and abundant herbivores in Late Jurassic North American faunas, but they declined in diversity in the Early Cretaceous, ...
[58]
The first theropod dinosaur (Coelurosauria, Theropoda ... - Nature
Jul 10, 2020 · Here we describe a new coelurosaur, which is the first tetrapod recovered from the basal layers of this stratigraphic unit that consist of dark shales.Missing: examples ornithischians
[59]
A Large Ornithurine Bird (Tingmiatornis arctica) from the Turonian ...
Dec 19, 2016 · Bird fossils from Turonian (ca. 90 Ma) sediments of Axel Heiberg Island (High Canadian Arctic) are among the earliest North American records ...
[60]
Cenomanian and Turonian (Early Late Cretaceous) multituberculate ...
The difference in composition of mammalian faunas from the Cenomanian to Turonian may reflect ecologic changes causing geographic shifts in mammalian faunas ...
[61]
Termite coprolites (Insecta: Isoptera) from the Cretaceous of western ...
Aug 6, 2025 · In other Cretaceous localities, termite coprolites are frequently found in amber, suggesting that resin producers, most probably araucarians, ...<|control11|><|separator|>
[62]
Crocodyliform remains from the Upper Cretaceous (Turonian ...
Here we describe recently recovered crocodyliform remains from the Tamagawa Formation of Kuji Group, Kuji city, northeastern Japan.
[63]
Large igneous province activity drives oceanic anoxic event 2 ...
Feb 19, 2024 · The global expansion of ocean anoxia resulted in the massive burial of organic carbon, leading to a decrease in pCO2 that could have contributed ...
[64]
Oceanic anoxic cycles? Orbital prelude to the Bonarelli Level (OAE 2)
Aug 6, 2025 · The prelude to the second global oceanic–anoxic event of the Cretaceous (OAE 2), as recorded in the classic sections of the Scaglia Bianca ...
[65]
A perturbed hydrological cycle during Oceanic Anoxic Event 2
Feb 1, 2014 · We suggest that an acceleration of the hydrological cycle during OAE2 played a key role in supplying nutrients to coastal waters and enhancing ...Missing: eutrophication | Show results with:eutrophication
[66]
Late Cretaceous ammonoids show that drivers of diversification are ...
Jun 27, 2024 · Ammonite diversity also displayed recurring significant negative relationships with origination rates at the species level and the genus level ...
[67]
https://www.nature.com/articles/s41467-024-49462-z
[68]
https://doi.org/10.1007/s11430-016-0166-y