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Lesser scaup

The Lesser scaup (Aythya affinis) is a medium-sized diving duck native to North America, recognized for its black-and-white plumage in males and brown tones in females, with a distinctive peaked crown that sets it apart from similar species.^[1]^[2] Measuring 15–18 inches (38–46 cm) in length, it inhabits wetlands and open waters, breeding across boreal and prairie regions before migrating south for winter.^[2] With a global breeding population estimated at 3–3.8 million individuals (approximately 3.3–3.4 million for lesser scaup as part of a 2025 combined scaup estimate of 3.7 million), it is the most abundant diving duck in North America, though populations have experienced fluctuations and recent declines.^[1]^[2]^[3]^[4] Males exhibit iridescent purple gloss on their dark heads and upper bodies during breeding season, while females and non-breeding males display mottled brown plumage with a white patch at the base of the bill for camouflage.^[2] The species is often confused with the Greater scaup (Aythya marila), but the Lesser scaup can be distinguished by its smaller, more peaked head shape, narrower bill, and shorter white wing stripe.^[1]^[2] Juveniles resemble females but acquire adult-like feathering by their first winter.^[5] Breeding occurs primarily in the Prairie Pothole Region and boreal forests from central Alaska through Manitoba and into the northern United States, where pairs nest on dry ground near shallow wetlands with emergent vegetation, laying 9–11 olive-buff eggs that incubate for 21–27 days.^[5]^[2] Ducklings are precocial, diving shortly after hatching and fledging after 47–54 days, often forming large creches.^[1]^[2] In winter, flocks numbering in the thousands congregate on large lakes, reservoirs, rivers, and sheltered coastal bays from the southern United States to Central America and the Caribbean, preferring freshwater habitats inland.^[1]^[5] As a popular game bird, the Lesser scaup faces hunting pressure, particularly in the north-central U.S. and central Canada, but its overall conservation status is rated as Least Concern by the IUCN due to its large and widespread population despite fluctuations and recent declines linked to wetland loss.^[2]^[6]^[3] Habitat conservation efforts in key breeding areas continue to support its populations.^[6]

Physical description

Plumage and morphology

The lesser scaup (Aythya affinis) is a medium-sized diving duck measuring 39–46 cm in length, with a wingspan of 68–78 cm and a weight ranging from 454–1,200 g.^[7]^[8] Males average 820 g and females 730 g.^[7] Its compact body form, with legs positioned farther rearward on the torso and larger, lobed hind toes, facilitates efficient underwater propulsion and maneuverability during dives.^[9] Plumage in adult lesser scaup features a mix of iridescent and barred patterns adapted for aquatic environments. The head is rounded, often with a slight peak at the rear in males and a flatter profile in females, contributing to a streamlined silhouette.^[7] The bill is broad, blue-gray, and tipped with a distinct black nail, aiding in foraging for submerged prey.^[10] Wing morphology includes 10 full-length primaries and 14–15 secondaries, with a white stripe primarily on the inner secondaries that extends only halfway along the wing during flight.^[11] This structure supports agile diving and rapid ascents to the surface. Sexual dimorphism in plumage is pronounced, with males displaying more vibrant colors during breeding, though detailed variations are addressed separately.^[7]

Sexual dimorphism and measurements

The lesser scaup exhibits pronounced sexual dimorphism in plumage, with adult males displaying a striking contrast of iridescent black head and breast against white sides and flanks, while the back is finely vermiculated in gray.^[7] The male's head shows a purplish to greenish gloss, and the bill is slaty blue.^[12] In contrast, adult females are overall dark brown, with a paler face and throat, a distinctive white patch at the base of the bill (sometimes broken or absent), and darker upperparts; the bill is dark gray.^[7]^[12] Eye color also differs between sexes and varies with age in females. Adult males have brilliant yellow irises.^[7] In females, eye color progresses from dark brown or olive brown in yearlings to olive yellow or yellow by three years of age, with juveniles showing grayish to olive tones.^[13] Juveniles resemble females but are duller overall, with less distinct markings; juvenile males may show slightly darker body plumage and glossier wing coverts than females, transitioning to adult plumage during their first winter.^[7]^[8] Males are slightly larger and heavier than females, with total length averaging 431 mm in adult males versus 418 mm in females, and wing length 208 mm versus 203 mm; overall, males are 2-4% larger in linear measurements.^[14] Body mass ranges from 454-1,200 g across both sexes, with males typically heavier (up to 1200 g) than females (up to 1100 g).^[7]^[8] Both sexes experience seasonal weight fluctuations, gaining lipid and protein reserves during spring migration to support breeding, with body mass increasing along migration routes from locations like the upper Midwest to breeding grounds.^[15]

Identification

Distinguishing features

The lesser scaup (Aythya affinis) is often challenging to distinguish from the greater scaup (Aythya marila) in the field, but key morphological traits provide reliable identification cues, particularly in head shape and bill profile. Males exhibit a less rounded head with a more peaked crown, creating an angular appearance, especially noticeable when the bird is at rest or preening.^[7]^[16] In contrast, the greater scaup's head is smoothly rounded. The bill of the lesser scaup is narrower and straighter overall, with less bulging at the base and a subtle curve toward the tip, aiding differentiation at close range.^[16]^[17] In flight, lesser scaup show distinct behavioral and plumage differences that facilitate identification from a distance. They typically form tighter, more compact flocks compared to the looser groups of greater scaup, and their wingbeats are faster and more rapid.^[16] The white wing stripe on lesser scaup is limited to the secondaries, extending only halfway along the wing and rarely reaching the primaries, whereas greater scaup display more extensive white covering both primaries and secondaries.^[4]^[7] During the eclipse plumage period in summer, male lesser scaup closely resemble females in their mottled brown-gray body and darker head, but they retain the characteristic white wing stripe on the secondaries, providing a consistent field mark across seasons.^[7]^[16] Quick sexing and aging in the field rely on subtle plumage and soft-part features. Adult males possess bright yellow eyes, while females and juveniles have dark brown eyes; this eye color difference is particularly useful in non-breeding seasons.^[7]^[16] Females often show a small white patch or facial crescent at the base of the bill, absent in males, offering a reliable indicator for sex determination even at moderate distances.^[17]^[7]

Similar species and hybridization

The lesser scaup is most frequently confused with the greater scaup due to their similar overall plumage, but the greater scaup is approximately 20% larger, with a rounder head lacking the small peak at the rear that is typical of the lesser scaup.^[10]^[7] In flight, the greater scaup shows more extensive white in the wings, with the white stripe extending into the primaries, whereas the lesser scaup's white is confined to the secondaries.^[10] Vocalizations also differ; male greater scaup produce a discordant "scaup-scaup" call, while male lesser scaup give a softer purring sound.^[18] The ring-necked duck is another species often mistaken for the lesser scaup, particularly in mixed flocks during migration and winter, as both have dark heads and gray sides in males.^[7] However, male lesser scaup have a paler gray back compared to the blackish back of the ring-necked duck, and they lack the white spur on the flanks present in the latter.^[19] The ring-necked duck also features a subtle chestnut neck ring (often hard to see) that is absent in scaup, along with a more peaked crown and sloping forehead versus the lesser scaup's flatter head profile.^[20] Hybridization occurs frequently among Aythya species, with the lesser scaup documented to interbreed with the greater scaup, ring-necked duck, and canvasback in the wild.^[21] These hybrids exhibit intermediate characteristics, such as mottled plumage blending the parental patterns and bill shapes that combine features like the broader base of the scaup with the canvasback's wedge-like form.^[22] Genetic studies using mitochondrial DNA and single nucleotide polymorphisms have confirmed admixed individuals in both lesser and greater scaup populations, identifying backcrossed generations through varying levels of introgression.^[23] Identification of hybrids often relies on aberrant traits like faint neck rings or side spurs in scaup-ring-necked duck crosses, or unusual bill patterns in canvasback hybrids, with genetic analysis providing definitive confirmation in research settings.^[24] Hybrids with the redhead or tufted duck are rarer, with occasional records noted in North America, typically showing subtle crests or reddish tones in plumage not seen in pure lesser scaup.^[25]^[26]

Taxonomy

Etymology and naming

The common name "lesser scaup" distinguishes this species from the greater scaup (Aythya marila), with "scaup" originating from the Scottish term "scalp," referring to a bed of mussels, clams, or shellfish, which reflects the bird's preference for foraging on such substrates.^[4]^[27] Other colloquial names include "little blackhead," alluding to the male's glossy black head, and "bluebill" or "little bluebill," due to the bluish-gray coloration of the bill.^[28] In North America, it is regionally known as "broadbill" because of the wide, flattened shape of its bill.^[29] The scientific name Aythya affinis comprises the genus Aythya, derived from the Ancient Greek "aithuia," a term used by classical authors like Hesychius to describe an unidentified seabird, possibly a diving duck or auklet.^[30]^[29] The specific epithet "affinis" comes from Latin, meaning "related" or "neighboring," indicating its close similarity to the greater scaup.^[27]^[31] The lesser scaup was first scientifically described in 1838 by British naturalist Thomas Campbell Eyton in his Monograph on the Anatidae (or Duck Tribe), where it was named Fuligula affinis as the "American scaup."^[32] It was later reclassified into the genus Aythya to better reflect its systematic relationships within the diving ducks.^[21]

Classification and phylogeny

The lesser scaup (Aythya affinis) belongs to the order Anseriformes, the family Anatidae (which encompasses ducks, geese, and swans), and the genus Aythya (diving ducks or pochards). It is a monotypic species, with no recognized subspecies, reflecting its relatively uniform morphology and genetic structure across its North American range. This classification aligns with the tribe Aythyini, a group of specialized diving ducks characterized by adaptations for underwater foraging, such as dense plumage and lobed feet.^[1]^[21]^[33] Phylogenetically, the lesser scaup forms a superspecies with the greater scaup (Aythya marila), its closest relative, based on shared morphological and genetic traits indicating a recent common ancestry. A comprehensive morphological analysis using 99 osteological, tracheal, and plumage characters positioned A. affinis as sister to A. marila within the genus Aythya, with this North American-Holarctic pair branching early in the pochard radiation. This sister relationship was corroborated by mitochondrial DNA (mtDNA) sequencing of the control region, which resolved Aythya as a monophyletic clade and confirmed minimal genetic divergence between the scaups despite occasional hybridization. Recent genomic analyses using thousands of loci further support this close relationship, with molecular clock estimates suggesting their divergence occurred less than 1 million years ago, during the Pleistocene, amid climatic fluctuations that promoted speciation in Nearctic waterfowl.^[34]^[35]^[36]^[37] mtDNA studies further underscore the lesser scaup's endemism to North America, revealing low levels of introgression from A. marila (e.g., only one shared haplotype among 111 sampled individuals) and supporting its isolation as a distinct lineage adapted to continental wetlands. The fossil record provides broader context for Aythya's evolutionary history, with Miocene relatives like Aythya shihuibas from late Miocene deposits in China (~10–5 million years ago) indicating early diversification of diving ducks, though no direct fossils of A. affinis have been identified due to its relatively recent origin. These genetic and paleontological insights highlight the lesser scaup's position as a basal member of the New World Aythya clade, shaped by Pleistocene glacial cycles.^[38]^[35]^[39]

Distribution and habitat

Breeding range

The lesser scaup (Aythya affinis) primarily breeds across northern and central North America, with its core range encompassing prairie pothole wetlands, boreal forests, and subarctic tundra habitats. This distribution extends from central Alaska eastward through the Yukon and Mackenzie Delta regions to northern Manitoba and western Ontario in Canada, and southward into the United States through the Prairie Pothole Region to include areas in Montana, North Dakota, Minnesota, and northern Wyoming.^[40]^[5]^[41] Breeding occurs predominantly in shallow, seasonal, and semipermanent wetlands and lakes that feature emergent vegetation such as cattails, bulrushes, and sedges, providing cover for nesting and foraging while avoiding deeper open waters. Pairs and broods associate with fresh to moderately brackish sites ranging from 0.85 to 6.1 hectares in size, often with 20–75% vegetative cover and shorelines partially bordered by shrubs or trees; nests are typically placed on the ground in adjacent uplands or over water within 10 meters of the edge.^[42]^[40] Breeding densities are highest in the Canadian parklands and boreal forests, where approximately 70% of the population occurs in the western boreal regions of Alaska and Canada. In key areas like North Dakota's semipermanent and permanent wetlands, densities can reach 4.7–6.1 pairs per square kilometer. The species breeds from sea level up to elevations around 2,000 meters in the Rocky Mountains, with peak abundance often between 500 and 1,500 meters in regions like British Columbia.^[43]^[40]^[44] Surveys indicate a slight increase in breeding activity within the Prairie Pothole Region over the past several decades, potentially influenced by climatic variations affecting wetland availability. As of 2025, the continental scaup population is estimated at 3.7 million, a 10% decline from 2024 and 25% below the long-term average.^[5]^[3]

Non-breeding range and habitat preferences

The non-breeding range of the lesser scaup (Aythya affinis) encompasses the southern United States, Mexico, Central America, and extends southward to Colombia, including Puerto Rico and the Caribbean islands.^[45] Wintering concentrations occur along the Gulf and Pacific coasts, in estuaries, marine habitats, and even offshore waters of the Gulf of Mexico, with birds utilizing coastal bays and inland lakes for extended periods from September through March.^[1] These populations originate primarily from breeding grounds in northern North America, shifting to lower latitudes to avoid harsh winters.^[5] During the non-breeding season, lesser scaup exhibit a strong preference for large open water bodies such as reservoirs, rivers, lakes, and large wetlands, which provide ample space for foraging and resting in flocks.^[42] They favor estuaries and coastal bays rich in aquatic invertebrates, including mollusks, over smaller or more enclosed habitats, and are less likely to use small ponds compared to other duck species.^[7] Inland freshwater lakes are commonly selected in the southern U.S., contrasting with the greater scaup's tendency toward marine environments.^[1] Key stopover sites during migration include areas along the Mississippi and Central Flyways, where birds pause on rivers, lakes, and large wetlands en route to wintering grounds.^[46] The Great Lakes serve as important staging areas, particularly for fall migrants and during post-breeding molt.^[47] In the southern Prairie Pothole Region, small flocks use mid-May stopovers during spring movements.^[46] Altitudinal distribution in winter is generally at lower elevations, though birds occur up to 1,500 m in Mexico.^[42] Vagrant records of lesser scaup outside the typical non-breeding range are infrequent but documented in Europe, including the United Kingdom in the 1980s, as well as in Hawaii.^[45]

Migration

Patterns and routes

The lesser scaup (Aythya affinis) undertakes extensive migrations primarily along the Central and Mississippi Flyways, with western populations utilizing the Pacific Flyway for portions of their journeys. These pathways facilitate movement from breeding grounds in the prairie potholes of the northern Great Plains and Canadian prairies southward to wintering areas along the Gulf Coast and in the eastern United States. Banding studies indicate an average migration distance of 2,000–3,000 km, with routes generally avoiding major mountain ranges such as the Rockies by following river valleys and lowland corridors.^[46] During migration, lesser scaup exhibit distinctive flock dynamics, forming large rafts of thousands of individuals on open water bodies for resting and foraging, which provide safety in numbers against predators. In flight, they travel in V-formations to optimize energy efficiency, and migrations are predominantly nocturnal, with flocks using daytime for foraging and resting. These flocks often mix interspecifically with greater scaup (Aythya marila) and other diving ducks like ring-necked ducks (Aythya collaris), creating mixed-species groups that enhance collective vigilance.^[10]^[48]

Timing and environmental influences

The northward breeding migration of lesser scaup typically occurs from mid-April to mid-May, with most individuals arriving at breeding grounds by late May and peak nesting activity commencing in June.^[10]^[49] Departures from wintering areas often begin as early as mid-February but intensify in March to early April, particularly as ice melt opens access to northern wetlands and staging areas.^[46]^[50] In fall, southward migration generally spans September to October, with adults and broods departing breeding areas after molt and family dispersal, though juveniles often begin dispersing earlier in late summer to nearby wetlands before joining larger flocks.^[10]^[46] This timing aligns with post-breeding resource availability, as scaup are among the later waterfowl to migrate south, often continuing into November.^[48] Environmental conditions significantly influence these schedules. Droughts in prairie breeding regions reduce wetland availability, delaying arrival and breeding migration by limiting foraging opportunities en route, as observed during prolonged dry periods in the 1980s and 2010s.^[51]^[52] Hurricanes disrupt fall routes by altering coastal and lake habitats, leading to temporary declines in wintering abundance, such as post-landfall reductions on Lake Pontchartrain following events like Hurricane Katrina. Warmer temperatures and reduced ice cover associated with climate change can advance spring migration timing and rates.^[50] As of the 2025 U.S. Fish & Wildlife Service Waterfowl Breeding Population and Habitat Survey, stable populations amid variable 2024 precipitation suggest continued resilience in migration patterns despite climatic pressures.^[53] Weather patterns also drive departures; cold fronts with northerly tailwinds trigger mass nocturnal movements, as documented through radar studies showing synchronized flock takeoffs following frontal passages over the Great Lakes and Mississippi Flyway.^[10]^[54] Greater spring precipitation further accelerates migration rates by enhancing wetland conditions for refueling.^[50]

Ecology and behavior

Foraging and diet

Lesser scaup are diving ducks that forage primarily by performing head-down dives, propelling themselves with powerful strokes of their webbed feet while keeping their wings folded against their bodies.^[55] These dives typically reach depths of 1-3 meters, though they can extend to 6 meters in some habitats, and last 10-30 seconds on average, allowing them to access benthic prey in shallow waters.^[56]^[9] The diet of lesser scaup is omnivorous but dominated by animal matter, with aquatic invertebrates such as mollusks (including zebra mussels Dreissena polymorpha), insects (e.g., chironomid larvae), and crustaceans (e.g., amphipods like Hyalella azteca) comprising the majority of intake in many regions.^[57]^[58] Plant material, including seeds and tubers of aquatic plants like pondweeds (Potamogeton spp.) and bulrushes (Scirpus spp.), makes up a substantial but lesser portion, often around 15-46% depending on location and season.^[57]^[56] Dietary composition shifts seasonally, with a greater emphasis on protein-rich invertebrates during the breeding period to support reproduction and a higher proportion of plant seeds and tubers in winter when invertebrate availability declines.^[59] In spring migration stopover areas, amphipods and insects often predominate, but recent studies indicate a shift toward less nutritious prey like snails and plants in some regions, potentially affecting body condition.^[57] Lesser scaup preferentially forage in shallow bays, wetlands, and lake margins with soft substrates that facilitate diving access to prey, as seen in the Great Lakes where they exploit invasive zebra mussels that proliferated since the 1990s.^[58] In Lake Erie, for example, zebra mussels constituted up to 98.6% of their diet during certain periods, reflecting the mussels' high density in shallow nearshore areas following their invasion in the late 1980s.^[60] To meet energetic demands, including building fat reserves for migration, adult lesser scaup consume approximately 220 grams of food per day during fall and winter, with foraging bouts distributed throughout the 24-hour cycle to fulfill nutritional needs efficiently.^[56]^[61] This intake supports daily energy expenditure and premigratory fattening, particularly during stopovers where high-quality foods like amphipods enhance lipid accumulation.^[62]

Reproduction and breeding biology

The lesser scaup exhibits seasonal monogamy, with pairs forming primarily during late spring migration from mid-March to early May, and new mates selected annually.^[49] Courtship involves males performing visual displays, including head-throwing where the head is rapidly jerked backward and forward, often accompanied by wing and tail flicks to attract females.^[2] Mate-switching and extra-pair copulations occur frequently during the breeding season.^[8] Nesting typically begins in late May through June on the breeding grounds, with females selecting sites near water bodies in emergent vegetation such as sedges, bulrushes, or cattails, often on dry land or floating mats in prairie potholes and marshes.^[48] The nest is a shallow ground scrape that the female gradually lines with grasses, other plant material, and her own down feathers, forming a saucer-shaped structure while laying eggs.^[8] Clutches average 9–11 pale greenish eggs, though sizes range from 6 to 14, with one egg laid daily; clutch size increases with female age, from about 9 eggs in yearlings to 12 in older birds.^[48]^[63] Incubation, performed solely by the female, lasts 21–28 days and begins 1–2 days before the last egg is laid; males typically depart shortly after clutch completion or midway through incubation in mid- to late June.^[8] Ducklings are precocial, hatching synchronously with eyes open and covered in down, and leave the nest within 24 hours to follow the female to water, where they feed themselves primarily on aquatic invertebrates.^[48] Females provide care for 2–5 weeks, leading broods and offering protection, while ducklings begin diving at about 2 weeks and fledge at 45–61 days old.^[8] First-year mortality is high, with survival rates around 38–41% for juveniles due to predation, cold stress, and other factors.^[64] Vocalizations play a role in breeding; females produce a sharp "scaup" or rasping alarm call similar to that of dabbling ducks when disturbed or signaling predators, and softer "arrr" or purring notes during courtship or to solicit aid.^[65]^[8] Males are mostly silent but emit soft grunts or whistles during courtship displays.^[8] If the first nest fails, females may renest up to two times in a season, though success rates are lower for yearlings (about 9%) compared to older females (18–27%), with overall renesting proportions varying by year and environmental conditions.^[63]

Conservation

Population trends and status

The continental population of lesser scaup (Aythya affinis), often assessed in combination with greater scaup (A. marila) due to identification challenges during surveys, peaked at approximately 6.3 million birds during the 1970s before declining steadily.^[66] By the 2010s, combined scaup numbers had fallen to around 3-4 million breeding birds, reflecting broader trends influenced by habitat availability.^[66] In 2025, the U.S. Fish and Wildlife Service (USFWS) estimated the combined scaup breeding population at 3.7 ± 0.2 million birds in the traditional survey area, similar to the 2024 figure but 25% below the long-term average of 4.9 million (1955–2024).^[3] This estimate falls short of the North American Waterfowl Management Plan (NAWMP) objective of approximately 6.25 million, based on 1970s-era benchmarks for population resilience.^[66] Overall, the population appears stable in the short term but remains below historical highs. The International Union for Conservation of Nature (IUCN) classifies the lesser scaup as Least Concern, with a 2016 assessment unchanged as of 2025 due to its large range and numbers exceeding vulnerability thresholds.^[67] Globally, NatureServe ranks it as G5 (secure), indicating no substantial risk despite localized pressures.^[41] Regionally, combined scaup populations show declines in prairie breeding areas, such as the Dakotas (65-71% below 2024 estimates in 2025) and southern Saskatchewan (18% below 2024), attributed to reduced wetland availability, while numbers in Alaska and the Yukon remained relatively stable (4% above 2024).^[3] Population monitoring relies on the USFWS Waterfowl Breeding Population and Habitat Survey (WBPHS), conducted via fixed-wing aerial transects since 1955 and corrected for visibility bias using ground or helicopter validation.^[3] Banding efforts, coordinated through the Bird Banding Laboratory, provide data on annual survival rates, estimated at 75-80% for adults based on long-term recovery analyses (1951–2011).^[68]

Threats and management efforts

The primary threats to lesser scaup populations stem from extensive habitat loss, particularly the drainage and degradation of wetlands in the Prairie Pothole Region (PPR), where agricultural expansion has resulted in over 50% of historical wetlands being lost or converted to cropland.^[69] This habitat fragmentation reduces breeding and foraging areas, exacerbating vulnerability during critical life stages. Climate change further amplifies these risks by intensifying droughts, which can dry up remaining wetlands and alter hydrologic cycles in the PPR, leading to decreased waterfowl productivity.^[70] Additional risks include residual lead poisoning from ingested spent shotgun pellets, despite the U.S. ban on lead shot for waterfowl hunting in 1991, as legacy contamination persists in sediments and affects diving ducks like lesser scaup that forage on the bottom.^[71] Contaminants accumulated in invasive zebra mussels, a common prey item for lesser scaup in the Great Lakes region, pose another concern, with elevated levels of selenium, cadmium, and polychlorinated biphenyls transferring through the food chain and potentially impairing health and reproduction.^[72] Hybridization with closely related species, such as greater scaup, may also contribute to genetic dilution in overlapping ranges, though its population-level impact remains understudied.^[73] Diseases represent a growing threat, with avian cholera (caused by Pasteurella multocida) causing recurrent outbreaks among waterfowl congregations in the 2020s, leading to high mortality in staging areas.^[74] Highly pathogenic avian influenza (HPAI H5N1) has similarly impacted lesser scaup, with 2025 surveillance data revealing infections in tracked individuals during migration, highlighting the virus's role in altering movement patterns and survival rates.^[75] Conservation efforts focus on habitat restoration and regulatory measures through the North American Waterfowl Management Plan (NAWMP), which has secured over 23 million acres of wetland and upland habitat across North America since 1986, benefiting lesser scaup breeding grounds in the PPR.^[76] Hunting regulations, enforced by the U.S. Fish and Wildlife Service, include restrictive bag limits—typically 2 scaup per day in many zones—to maintain sustainable harvest levels.^[77] Recent research addresses emerging threats, including research on dam-induced changes in the Mississippi River that disrupt scaup migration stopover sites, and climate models projecting potential northward range shifts in breeding areas due to warming temperatures.^[78]^[2]

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Oct 2, 2023 · Redhead is known to have hybridized with each of the other four North American Aythya in the wild, Canvasback, Lesser Scaup, Greater Scaup, and Ring-necked ...
[25]
Tufted Duck x Lesser Scaup (hybrid) - Birds of Nebraska – Online
Tufted Duck hybridizes with other species of Aythya (Carboneras and Kirwan 2020). There are numerous records shown in eBird.
[26]
LESSER SCAUP - BIRDS OF HAWAII
NAME: The English name 'Lesser' implies that there is a 'Greater' Scaup. The name 'Scaup' would origin from Sottish 'scalp' and refers to a ledge in the water, ...
[27]
Lesser Scaup (Aythya affinis) - iNaturalist
The lesser scaup (Aythya affinis) is a small North American diving duck that ... It is colloquially known as the little bluebill or broadbill because of its ...
[28]
Lesser Scaup Bird Facts - Aythya affinis - A-Z Animals
Their name comes from the female's call, which sounds like scaup, scaup. Migration Pattern and Timing. Lesser scaups are some of the last ducks to migrate each ...Missing: vocalizations | Show results with:vocalizations
[29]
Scaups, Pochards, and Allies (Genus Aythya) - iNaturalist
It has twelve described species. The name Aythya comes from the Ancient Greek word αυθυια, aithuia, which may have referred to a sea-dwelling duck or an auklet.Missing: etymology | Show results with:etymology
[30]
Greater and Lesser Scaup - Eastside Audubon Society
Sep 4, 2019 · The species name of the LESC, affinis (aff-EYE-nis), is Latin meaning adjacent, neighboring, allied, or related to, apparently in reference to ...<|control11|><|separator|>
[31]
Even John James Audubon Couldn't Tell the Difference Between ...
Feb 22, 2017 · ... Scaup Duck” is actually two different birds, Greater Scaup and Lesser Scaup. The latter species wasn't even described to science until 1838 ...
[32]
[PDF] A Phylogenetic Analysis of Modern Pochards (anatidae: Aythyini)
Sep 8, 2024 · Sexual size dimorphism was evaluated us- ing the ratio of the mean body mass of males divided by the mean body mass of females. Relative ...Missing: measurements | Show results with:measurements
[33]
A Phylogenetic Analysis of Modern Pochards (Anatidae: Aythyini)
101:410-435. LIvEZEY, B. C. 1991. A phylogenetic analysis and classification of Recent dabbling ducks (Tribe. Anatini) based on comparative morphology. Auk.
[34]
Multiple independent transpositions of mitochondrial DNA ... - PNAS
We provide evidence of nuclear sequences homologous to the mtDNA control region in seven species of diving ducks (tribe Aythyini). Phylogenetic analysis places ...
[35]
Rapid and recent diversification patterns in Anseriformes birds - NIH
Sep 11, 2017 · In the present study, we obtained similar divergence time values in the major nodes using a molecular clock based on mito-genomes and two ...
[36]
Molecular Ecology | Molecular Genetics Journal | Wiley Online Library
Nov 24, 2015 · Among 111 individuals, we found one introgressed mitochondrial DNA haplotype in lesser scaup and four in greater scaup. Likewise, based on the ...Missing: studies | Show results with:studies
[37]
[PDF] Re-description and phylogenetic assessment of the Late Miocene ...
It appears that Aythya shihuibas is distinct from other Miocene Asian anatids, and that the cf. Anas fossil could be the same species as one from Mongolia. The ...
[38]
[PDF] Habitat Suitability Index Models: Lesser Scaup (Breeding). - DTIC
The primary breeding range of the lesser scaup generally extends southeast from central Alaska to western Ontario and south to northern Wyoming and central ...
[39]
Aythya affinis | NatureServe Explorer
Habitat Comments: BREEDING: Marshes, ponds, and small lakes (AOU 1998). Usually nests near small ponds and lakes, sedge meadows, creeks; in cover 1-2 ft high, ...
[40]
Habitat - Lesser Scaup - Aythya affinis - Birds of the World
Pairs and broods are typically associated with fresh to moderately brackish, seasonal and semipermanent wetlands and lakes with emergent vegetation such as ...
[41]
Lesser Scaup Breeding Probability and Female Survival ... - BioOne
Aug 1, 2009 · Seventy percent of lesser scaup breed in the western boreal forest of Alaska (USA) and Canada, which is where scaup populations have seen ...Methods · Statistical Analysis · Discussion<|control11|><|separator|>
[42]
Lesser Scaup - BC Breeding Bird Atlas
The elevation range confirms that given by Campbell et al. (1990): highest abundance occurs in the 500-1,500 m range, although the Atlas sample size for ...Missing: rocky | Show results with:rocky
[43]
[PDF] Waterfowl Population Status, 2024 - U.S. Fish and Wildlife Service
Aug 20, 2024 · Scaup (greater and lesser combined) breeding population estimates (in thousands) for ... Lesser Canada geese nest throughout inte- rior and ...
[44]
Distribution - Lesser Scaup - Aythya affinis - Birds of the World
Species is common on the Great Lakes and along the upper Atlantic Coast of the U.S. from New Jersey south. Also found during winter in nearly any part of the ...Missing: preferences | Show results with:preferences
[45]
Movements and Migration - Lesser Scaup - Aythya affinis
Major migration routes for most Alaskan and Canadian breeders are through Canada south via the Mississippi Flyway to the Gulf Coast (Central and Mississippi ...
[46]
References - Lesser Scaup - Aythya affinis - Birds of the World
Afton, A. D. (1993). Post-hatch brood amalgamation in Lesser Scaup: female behavior and return rates, and duckling survival. Prairie Naturalist 25:227-235.
[47]
Breeding - Lesser Scaup - Aythya affinis - Birds of the World
Minnesota in late April (MJA); 68% of males and 99.8% of females are paired by early May on Delta Marsh, Manitoba (Siegfried 1974b ...Missing: percentage | Show results with:percentage
[48]
Environmental factors influence lesser scaup migration chronology ...
Aug 22, 2016 · We hypothesized that annual variation in spring migration chronology of scaup may be influenced by ambient temperature, snow water equivalency ( ...
[49]
Lesser Scaup Life History, All About Birds, Cornell Lab of Ornithology
Lesser Scaup breed in the prairie pothole region in the United States and Canada north throughout the boreal forest and tundra zone.
[50]
DU Special Report: 2024 Status of Waterfowl - Ducks Unlimited
Oct 29, 2024 · Although drought persisted across the prairies in 2023, this year's results suggest the decline may have slowed for most duck species, with ...Pacific Flyway · Atlantic Flyway · Goose Outlook
[51]
Lesser Scaup | North Dakota Game and Fish
Scientific Name, Aythya affinis. General Description, L 16.5”, WS 25”, 1.8lb ... Lesser Scaup have increased in the Prairie Pothole Region over the past ...Missing: first | Show results with:first
[52]
Radar observations of bird migration over the Great Lakes
Aug 8, 2025 · Nocturnal migrants seem generally less sensitive to a land/sea transition and rather cross water bodies in broad front migration (Myres 1964; ...
[53]
Diving energetics in lesser scaup (Aythyta affinis, Eyton)
May 1, 1994 · Durations of dives (td=13.5±1.4 s) and surface intervals (ti=16.3±2.2 s) were within the normal range for ducks diving to 1.5 m depth.
[54]
[PDF] Habitat suitability index models: Lesser scaup (wintering) - GovInfo
Lesser scaup prefer diving in water 1 to 3 m (3.3 to 9.8 ft) deep (Palmer ... Water depth (V3) affects the availability of food resources to lesser scaup.<|separator|>
[55]
[PDF] Diets of Lesser Scaup during Spring Migration throughout the Upper ...
Nutrient reserves acquired during spring migration are important determinants of sur- vival and reproductive success in Lesser Scaup. (hereafter scaup), and ...
[56]
Food habits of diving ducks in the Great Lakes after the zebra ...
The aggregate percent of zebra mussels in the diet of Lesser Scaups was higher in Lake Erie (98.6%) than in Lake St. Clair (54.4%).Missing: foraging | Show results with:foraging
[57]
[PDF] Lesser Scaup - Alaska Center for Conservation Science
Diet is fexible and changes in response to spatial and temporal availability (Anteau et al. 2014;. DuBour 2019). In the summer, lesser scaup eat aquatic ...
[58]
[PDF] Food Habits of Diving Ducks in the Great Lakes After the Zebra ...
Jan 1, 1996 · Lesser Scaups from Lake Erie ate larger zebra mussels than. Lesser Scaups from Lake St. Clair (Table 2, F•,•0 = 16.19, P < 0.001). Using Lake ...
[59]
FOOD CONSUMPTION BY CANVASBACKS AND SCAUPS - jstor
mean daily food intake per canvasback was 0.78 pound (wet weight) or 0.05 pound (dry we. The mean daily food intake per lesser scaup was 0.49 pound (wet ...
[60]
True metabolizable energy of foods consumed by lesser scaup ...
Sep 27, 2024 · Wild millet is consumed by scaup less frequently than the other foods (Moore et al. 1998) but was chosen for a comparison with dabbling duck ...
[61]
[PDF] Influence of Age and Time on Reproductive Performance of Female ...
Lesser Scaup are needed to examine the rela- tive importance of experience and age varia- tion in RE more critically. Comparisons of clutch size of wild first- ...Missing: dimorphism | Show results with:dimorphism
[62]
Demography and Populations - Lesser Scaup - Aythya affinis - Birds ...
... survival over the past 50 yr was highest for adult males (0.83), followed by adult females (0.78), first-year females (0.41), and first-year males (0.38; DNK).
[63]
[PDF] Observations on sound production in the Anatidae
Calls of female pochards are acoustic- ally much like those of female dabbling ducks. Thus, the alarm call of a female. Lesser Scaup Aythya affinis has the same.
[64]
The Great Scaup Mystery | Ducks Unlimited
Nov 1, 2007 · Some biologists have speculated that environmental contaminants may be a factor in the scaup population decline. However, researchers needed to ...Missing: northward | Show results with:northward
[65]
[PDF] Waterfowl Population Status, 2025 - U.S. Fish and Wildlife Service
Sep 2, 2025 · The combined estimate of lesser and greater scaup (A. affinis and A. marila; 3.7 소 0.2 million) was similar to the 2024 estimate and 25% lower ...
[66]
Lesser Scaup Aythya Affinis Species Factsheet | BirdLife DataZone
Habitat classification ; Artificial/Aquatic & Marine, Artificial/Aquatic - Water Storage Areas (over 8ha), Non-breeding ; Marine Coastal/Supratidal, Coastal ...<|control11|><|separator|>
[67]
Temporal variation in survival and recovery rates of lesser scaup
Apr 13, 2016 · Annual estimates of survival (S) and recovery probabilities (f) for lesser scaup banded and recovered in North America, 1951-2011, based on ...
[68]
Continued Decline of Wetlands Documented in New U.S. Fish and ...
Mar 22, 2024 · More than half of wetlands in the lower 48 states are gone, and losses continue, mostly in the Southeast, Great Lakes and Prairie Pothole regions.
[69]
Prairie Pothole Wetlands: Small Basins, Big Impact - SDSU Extension
Aug 28, 2025 · ... areas, losses exceed 90 percent. The region's fertile soils and relatively flat topography make it ideal for farming, but wetland drainage ...
[70]
[PDF] LEAD POISONING OF NORTH AMERICAN WILDLIFE FROM LEAD ...
Havera et al. (1992) found a 15% imbedded shot rate for Canvasbacks and 9% rate for Lesser Scaups (Aythya affinis). Some 27% of ...
[71]
UMESC - Study site locations: Michigan - Lesser scaup
Study site locations: Michigan - Zebra mussels are a vector for increased contaminant exposure - Lesser scaup. Selenium in lesser scaup liver tissue was ...
[72]
Division of Forestry and Wildlife: Wildlife Program | Lesser Scaup
As with many ducks, lesser scaups have difference in plumage depending on sex, with breeding males showing a dramatic contrast between their white body and ...<|control11|><|separator|>
[73]
Avian Cholera | Game Commission | Commonwealth of Pennsylvania
Avian cholera is the most significant infectious disease of wild waterfowl in North America. Single outbreaks can kill thousands of birds, and outbreaks occur ...Missing: scaup 2020s
[74]
Telemetry data of a Lesser Scaup (Aythya affinis) positive for 2.3.4.4 ...
Telemetry data of a Lesser Scaup (Aythya affinis) positive for 2.3.4.4 Highly Pathogenic H5N1. Metadata Updated: October 8, 2025. Wild lesser scaup from the ...
[75]
[PDF] 2020 Canadian NAWMP Report
For NAWMP partners, restoration means recovering the health of wetlands and upland habitats that waterfowl and other wildlife need to thrive. A restored wetland ...
[76]
Seasons and Bag Limits - Commonwealth of Pennsylvania
Daily limit for scaup varies; see below. Possession limits are three times the daily limits. SCAUP: North Zone: 1 scaup daily, Oct. 11-25 and Nov. 18-Dec ...
[77]
Lesser Scaup Highlight Ecological Changes Caused by Dams
Jun 11, 2025 · Lesser scaup use the Mississippi River as a migratory path. Pool 19, a critical resting point for the ducks, is in need of restoration to ...<|control11|><|separator|>
[78]
Modeling Wetland Resources for Spring Migratory Waterbirds Under ...
Apr 30, 2025 · ... lesser scaup during spring migration across the upper Midwest, USA. ... climate-change impacts on northern prairie wetlands. Wetlands 36 ...