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Dorsal fin

The dorsal fin is an unpaired, median fin situated on the dorsal (upper) surface of the body in many aquatic vertebrates, most notably fish and cetaceans such as dolphins and whales, where it primarily functions to provide stability and prevent rolling during locomotion.^[1]^[2] In fish, it is typically supported by a series of internal pterygiophores attached to the vertebral column, with external structures consisting of either stiff spines or flexible rays.^[1] Structurally, dorsal fins in bony fish (teleosts) often exhibit two distinct forms: a spiny dorsal fin composed of unbranched, rigid spines that can lock into position for protection or anchoring, and a soft-rayed dorsal fin made of segmented, branched rays that aid in finer control of movement.^[1] These spines may be venomous in certain species, such as lionfish, enhancing defensive capabilities against predators.^[1] In cartilaginous fish like sharks, the dorsal fin lacks rays or spines but features a trailing edge that generates low-pressure zones to improve hydrodynamic efficiency and thrust from the tail.^[3] For cetaceans, the dorsal fin is a fleshy, boneless structure composed of dense connective tissue, varying in size and shape across species—for instance, tall and falcate in killer whales for enhanced stability in high-speed pursuits.^[4]^[5] Beyond stabilization, dorsal fins contribute to steering, balance, and even defensive displays; in fish, raising the fin can deter attackers or facilitate quick turns, while in dolphins, its unique shape and markings enable individual identification in research and social contexts.^[6]^[7]^[8] Not all aquatic species possess a dorsal fin—some fish have reduced or absent versions, and certain whales like sperm whales lack them entirely—highlighting evolutionary adaptations to specific aquatic lifestyles.^[1]^[9]

Introduction

Definition and Basic Characteristics

The dorsal fin is an unpaired, medial fin situated on the dorsal (back) side of aquatic vertebrates, such as fish, sharks, and certain marine mammals, typically extending along the midline from behind the head toward the caudal peduncle, the narrow region preceding the tail.^[1]^[10] This positioning distinguishes it as a key element in the median fin system, which contrasts with the laterally positioned paired fins.^[11] Basic characteristics of the dorsal fin include variability in size, shape, and configuration relative to the body axis. Shapes range from triangular or low-profile forms in many bony fish to tall, sail-like structures in species like sharks, while size can span from small and compact to elongated along much of the back.^[12]^[13] In some fish, such as perches and sunfish, the dorsal fin may consist of a single structure or multiple segments, often featuring a forward spiny portion and a rear soft-rayed section, though it remains unpaired overall.^[14]^[15] The term "dorsal fin" derives from the Latin dorsum, meaning "back," reflecting its anatomical placement, and entered ichthyological literature in the mid-18th century as systematic descriptions of fish anatomy emerged.^[16] It is generally distinguished from other fins by its unpaired, median nature, in contrast to the paired pectoral and ventral (pelvic) fins that occur symmetrically on the sides of the body, and alongside the similarly unpaired anal and caudal fins.^[1]

Occurrence Across Vertebrates

Dorsal fins are prevalent among aquatic vertebrates, particularly within the major fish clades. In actinopterygian fish (ray-finned fishes), which comprise the largest group of living vertebrates, a single dorsal fin is characteristic, supported by lepidotrichia (fin rays) that provide flexibility and stability during swimming.^[17] For example, salmon (Salmo salar) exhibit a prominent single dorsal fin positioned midway along the body. In contrast, many perciform fish like the perch (Perca fluviatilis) possess two distinct dorsal fins: an anterior spiny portion for defense and a posterior soft-rayed one.^[18] Chondrichthyan fish (sharks, rays, and chimaeras) typically feature two dorsal fins, often equipped with anterior spines for protection against predators, as seen in species like the great white shark (Carcharodon carcharias).^[19] Sarcopterygian fish (lobe-finned fishes) vary in dorsal fin structure; coelacanths have two separate dorsal fins with fleshy bases, while lungfish, such as the Australian lungfish (Neoceratodus forsteri), have a single low, continuous dorsal fin integrated with the tail.^[20]^[21] Among marine mammals, dorsal fins occur primarily in cetaceans but show significant variation. In odontocetes like dolphins (family Delphinidae), a prominent, falcate dorsal fin is present, aiding in maneuverability in open water, as observed in the bottlenose dolphin (Tursiops truncatus). However, in some mysticetes and certain odontocetes, the structure is reduced or vestigial; beluga whales (Delphinapterus leucas), for example, lack a true dorsal fin, instead possessing a low dorsal ridge to minimize drag and heat loss in icy Arctic environments.^[22] In extinct vertebrate groups, dorsal fins were also common in secondarily aquatic forms. Ichthyosaurs, Mesozoic marine reptiles convergent with modern cetaceans, possessed a dorsal fin in later species, evidenced by preserved soft tissue impressions showing a low, triangular structure along the midline, as in Ophthalmosaurus icenicus.^[23] Dorsal fins are absent or highly reduced in non-aquatic or semi-aquatic vertebrates outside of fully marine lineages. Most amphibians lack dorsal fins in their adult forms, relying instead on limbs for locomotion, though tadpoles possess a dorsal tail fin during larval stages. Lungfish, while aquatic sarcopterygians, show reductions in some species, with the dorsal fin often low and elongated rather than prominent. Terrestrial vertebrates, including reptiles, birds, and mammals, generally do not possess dorsal fins, as their transition to land-based locomotion eliminated the need for such aquatic stabilizers; exceptions are rare and limited to secondarily aquatic taxa like sea turtles, which have low dorsal scutes but no true fin.^[24] Taxonomically, dorsal fins are found in the vast majority of the approximately 37,000 described fish species (as of 2025), with variations linked to habitat. Open-water (pelagic) swimmers, such as tuna (Thunnus spp.), typically have taller, more rigid dorsal fins for enhanced stability at high speeds, while bottom-dwelling species like flatfish (Pleuronectiformes) often exhibit reduced or embedded dorsal fins to facilitate camouflage and benthic movement.^[25]^[26]

Anatomy and Development

Structural Components

The dorsal fin in vertebrates, particularly fish, is primarily composed of flexible, ray-like structures known as lepidotrichia in bony fishes (actinopterygians), which consist of paired, segmented hemitrichia formed from dermal bone and covered by a thin layer of skin often bearing scales or, in some cases, denticles for added protection.^[27] These fin rays are embedded in connective tissue that provides flexibility and strength, with muscle fibers attaching at their proximal bases to enable controlled movement and shaping of the fin.^[28] Support for the dorsal fin arises from internal skeletal elements that anchor it to the axial skeleton. In bony fishes, pterygiophores—bony or initially cartilaginous rods—serve as the primary supports, inserting between the neural spines of the vertebral column to transmit forces and maintain rigidity; these typically include proximal (basal), middle, and distal components, with radials extending distally to articulate with the lepidotrichia.^[29] In cartilaginous fishes such as sharks, the dorsal fin lacks ossified pterygiophores and is instead supported by a series of cartilaginous radials and basals that radiate from the body wall, providing a more flexible but rigid framework.^[30] Variations in these support structures occur across taxa, with some bony fishes exhibiting hardened spines integrated into the anterior dorsal fin for defensive purposes; for instance, in catfish (Siluriformes), the leading dorsal spine is sharp, serrated, and associated with venom glands at its base, deterring predators through puncture and toxin delivery.^[31] The dorsal fin receives a rich blood supply via branches from the dorsal aorta, forming a vascular network within the fin tissue that supports metabolic demands. In endothermic species like sailfish (Istiophorus platypterus), this vascularization contributes to thermoregulation by enhancing heat exchange across the fin's large surface area, aiding in retention or dissipation of body heat.^[32] Innervation of the dorsal fin includes both motor and sensory components, with spinal nerves forming plexuses at the fin base to control musculature. Sensory nerves, particularly mechanoreceptors embedded in the lepidotrichia, provide proprioceptive feedback on fin position and water flow, essential for coordinated locomotion.^[33] Regarding size, the dorsal fin varies widely but can be proportionally large; in sailfish, it extends nearly the full body length, often exceeding one-third of the total length in adults, which amplifies its structural and physiological roles.^[34]

Embryonic Origins and Growth

The dorsal fin in teleost fish originates embryonically from the median fin fold, a continuous epithelial structure that emerges along the dorsal midline shortly after gastrulation. In zebrafish (Danio rerio), this fin fold begins forming around 24 hours post-fertilization, during the late somitogenesis stage, and serves as the primordium for all median fins, including the dorsal, caudal, and anal fins. The fold consists of a thin layer of epidermis overlying mesenchymal cells derived from the lateral plate and paraxial mesoderm, providing the initial scaffold for fin development. This structure is transient and conserved across early vertebrate embryos, facilitating the outgrowth of unpaired appendages without paired fin buds. During growth, the dorsal fin skeleton differentiates from the sclerotome, a ventral compartment of somites in the paraxial mesoderm. Lineage tracing in zebrafish reveals that sclerotome cells migrate dorsally into the fin bud mesenchyme starting at approximately 5.6 mm standard length (around 2 weeks post-fertilization), contributing pterygiophores and proximal radials to support fin rays. Concurrently, programmed cell death via apoptosis in the inter-ray regions of the fin fold sculpts the distinct boundaries of the dorsal fin, reducing the continuous fold into separate modules by eliminating excess tissue between emerging fins. This process, marked by caspase-3 activation, ensures proper patterning and prevents fusion of median structures. Genetic regulation of dorsal fin position and ray formation relies on Hox gene clusters and Sonic hedgehog (shh) signaling pathways. Hox genes, particularly those in the hox13 paralog groups, establish anterior-posterior identity along the fin axis, coordinating ray segmentation and elongation. Meanwhile, shh expression in the notochord and floor plate diffuses to pattern the fin mesenchyme, promoting proximal-distal outgrowth and lepidotrichial ray differentiation through Gli transcription factor activation. These pathways operate independently of the larval fin fold's epidermal constraints, allowing autonomous median fin development. Post-embryonically, dorsal fin growth occurs through sequential addition of fin rays in larvae, with new lepidotrichia forming at the posterior margin via hedgehog-mediated proliferation. In zebrafish, this juvenile fin maturation spans 4-6 weeks post-hatching, involving ray bifurcation and extension to reach adult proportions. In metamorphosing flatfishes like the Japanese flounder (Paralichthys olivaceus), thyroid hormone-driven remodeling alters dorsal fin ray counts and shifts its position anteriorly, adapting to the benthic, asymmetric body plan during the larval-to-juvenile transition. Developmental anomalies, such as the dominant smoothback (smb) mutation in zebrafish, lead to complete absence of the dorsal fin due to sclerotome depletion. This 2024-identified insertion in sox10:Gal4 regulatory elements impairs paraxial mesoderm segmentation by 24 hours post-fertilization, preventing mesenchymal aggregation in the dorsal fin bud and resulting in truncated anal fins as well. Such mutants highlight the sclerotome's essential role in median fin induction, with no compensatory mechanisms from other mesodermal sources.

Functions and Physiology

Hydrodynamic Stabilization

The dorsal fin serves as a primary stabilizer during swimming, functioning like a keel to prevent rolling and maintain yaw stability by generating counter-torque against lateral forces induced by body undulations or external disturbances.^[35] This hydrodynamic role is evident in its ability to produce lateral forces that counteract rotational tendencies, ensuring the fish maintains a straight trajectory in steady locomotion.^[36] In elasmobranchs such as spiny dogfish, the first dorsal fin exhibits controlled oscillations out of phase with the body, enhancing this stabilizing effect during cruising speeds.^[37] Beyond basic equilibrium, the dorsal fin contributes to maneuverability by aiding pitch control and facilitating sudden turns through modulation of water flow around its surface. By adjusting its conformation, the fin alters hydrodynamic pressures to refine body orientation, improving agility during straight-line propulsion and evasive actions.^[36] This active control allows fish to respond dynamically to environmental cues without compromising forward momentum.^[35] Biomechanically, the dorsal fin's streamlined shape minimizes drag by allowing retraction or alignment with the body axis in fast swimmers, reducing frictional resistance during sustained locomotion.^[1] Simultaneously, it generates lift forces perpendicular to the body axis, which oppose roll and contribute to overall postural stability through vortex shedding and pressure differentials.^[38] Comparatively, pelagic species often feature larger dorsal fins to enhance open-water balance, where prolonged exposure to currents demands greater resistance to yaw perturbations.^[36]

Specialized Physiological Roles

The dorsal fin also serves defensive roles in several fish species, often through venomous or mechanically adaptive structures. In lionfish (Pterois spp.), the dorsal fin bears up to 13 long, venomous spines connected to glandular venom sacs, which deliver neurotoxic proteins upon penetration, deterring predators and causing intense pain and tissue damage in potential threats or handlers. Weeverfish (Trachinus spp.), such as the greater weever, feature a prominent first dorsal fin spine armed with venom that includes hemolytic and proteolytic enzymes, enabling rapid immobilization of attackers via injection during defensive postures when buried in sediment. Triggerfish (Balistidae family), exemplified by the gray triggerfish, employ an erectable first dorsal fin with three lockable spines that can be raised and secured to wedge into tight crevices, preventing extraction by predators, or displayed aggressively to signal threat.^[39]^[40]^[41] Beyond defense, the dorsal fin contributes to sensory and display functions in various species. In anglerfishes (Lophiiformes order), the first dorsal fin spine is modified into an illicium—a movable filament topped with a bioluminescent esca (lure)—which protrudes from the head to mimic prey and attract organisms into striking range, facilitating ambush predation in low-light deep-sea environments. During courtship, male guppies (Poecilia reticulata) intensify coloration and patterns on their dorsal and caudal fins through physiological changes, such as iridophore expansion, to signal fitness and stimulate female receptivity, with brighter displays correlating to higher mating success in varying light conditions.^[42]^[43]^[44] In some species, the dorsal fin aids propulsion in unconventional ways, supplementing or altering standard swimming. The ocean sunfish (Mola mola) uses its large, flexible dorsal fin in conjunction with the anal fin for sculling propulsion, but frequently adopts a lateral drifting posture at the surface where the dorsal fin acts like a sail, passively harnessing wind and currents for energy-efficient displacement over long distances while basking or recovering from dives. Recent drone observations of white sharks (Carcharodon carcharias) reveal the dorsal fin's high flexibility, allowing rotation at its base to probe or investigate surface objects—such as floating debris or potential prey—by directing sensory flow or tactile exploration without altering body orientation.^[45]^[46] In cetaceans, the dorsal fin aids thermoregulation by serving as a site for heat dissipation through specialized vascular networks, allowing controlled heat loss during dives or in varying water temperatures.^[47] Additionally, dorsal fin structures provide physiological markers for research and identification. In bluefin tuna (Thunnus thynnus), annual growth rings in the calcified first dorsal fin spine serve as reliable aging indicators, with recent comparative analyses confirming their accuracy against vertebral counts for estimating age and growth rates in Mediterranean populations up to 10 years old. For Rice's whales (Balaenoptera ricei), natural notches, lacerations, and nicks on the dorsal fin enable photo-identification of individuals, as demonstrated in a 2025 catalog of 31 whales where such markings facilitated tracking of population dynamics and site fidelity in the Gulf of Mexico.^[48]^[49]

Variations and Adaptations

In Fish Species

In ray-finned fishes (Actinopterygii), dorsal fins exhibit significant morphological diversity, often divided into spiny and soft-rayed types that serve distinct ecological roles. The spiny dorsal fin, characteristic of the superorder Acanthopterygii, features rigid spines that provide defense against predators by locking into place when threatened, as seen in species like the largemouth bass (Micropterus salmoides), which possesses a dual dorsal fin with the anterior portion spiny for protection and the posterior soft-rayed for propulsion.^[50] In contrast, soft dorsal fins predominate in more basal actinopterygians, offering flexibility for maneuvering in complex habitats, while some advanced percomorphs like parrotfishes (Scaridae) display a single, high-arched dorsal fin with numerous soft rays that aids in agile navigation among coral reefs.^[51] Among cartilaginous fishes (Chondrichthyes), dorsal fins are typically triangular and flexible, covered in dermal denticles that reduce drag and enhance hydrodynamic efficiency during swimming. In sharks such as the great white (Carcharodon carcharias), these fins stabilize the body against roll and contribute to precise turns in open water pursuits.^[52]^[53] Skates (Rajidae), however, often have reduced or vestigial dorsal fins, with many species lacking prominent structures to facilitate benthic gliding along the seafloor, minimizing interference with their flattened body form.^[54] Lobe-finned fishes (Sarcopterygii) feature fleshy dorsal fins supported by robust internal bones, representing primitive structures that foreshadowed tetrapod limb evolution. In coelacanths (Latimeria spp.), the two separate dorsal fins are thick and lobed, providing stability in deep-water environments where slow, deliberate movements predominate.^[55]^[56] Ecological pressures drive further variations in dorsal fin morphology across fish species, adapting structure to habitat and lifestyle. Fast-swimming pelagic predators like tunas (Thunnus spp.) possess tall, sickle-shaped dorsal fins that generate lift and reduce yaw during high-speed chases, enabling sustained velocities over 70 km/h.^[57] Bottom-dwelling flatfishes such as flounders (Paralichthys spp.) have low-profile dorsal fins that extend continuously along the body, blending seamlessly with the substrate to enhance camouflage and ambush hunting.^[58] In specialized cases, burrowing species like the newly discovered Listrura elongata from southern Brazil's Rio Camboriú basin exhibit severely reduced or absent dorsal fins, an adaptation for navigating silty burrows that prioritizes streamlining over surface stability.^[59] Dorsal fin patterns also play a key role in species identification and conservation efforts. A 2025 study on blacktip reef sharks (Carcharhinus melanopterus) demonstrated that unique pigmentation and shape variations in dorsal fins enable reliable individual and species-level discrimination, improving population monitoring in coral reef ecosystems.^[60] Additionally, the iSharkFin AI system, developed in 2021, achieves 59.1% accuracy at the species level and 85.3% at the genus level in recognizing wet dorsal fins from photographs, facilitating rapid identification of 39 shark species in fisheries enforcement.^[61]

In Marine Mammals and Reptiles

In cetaceans, dorsal fin morphology varies significantly across species, reflecting adaptations to their aquatic lifestyles. Dolphins and orcas typically possess tall, triangular dorsal fins that enhance hydrodynamic stability and facilitate high-speed swimming.^[62] In contrast, humpback whales feature a small, falcate (sickle-shaped) dorsal fin located posteriorly on the back, which supports agile maneuvers during foraging. Notably, beluga whales and narwhals lack a true dorsal fin, instead exhibiting a low dorsal ridge; this absence minimizes drag and heat loss in icy Arctic environments, allowing safer navigation under pack ice.^[22]^[62] Dorsal fins in cetaceans often bear scars and notches from human interactions, particularly with fisheries. In the critically endangered Rice's whale, photo-identification studies have documented deep triangular and round notches, as well as linear lacerations on dorsal fins in over 30% of cataloged individuals, attributes linked to entanglements in fishing gear such as lines and traps.^[49] Among pinnipeds, dorsal structures are generally less prominent than in cetaceans. True seals exhibit small, flexible dorsal ridges rather than distinct fins, aiding in streamlined swimming and haul-out behaviors.^[63] These features, along with natural markings like scars and pelage patterns, are utilized in photo-identification catalogs to track individual seals and monitor population dynamics.^[64] In extinct marine reptiles, dorsal fin-like structures provided stabilization during locomotion. Fossil evidence from mosasaurs reveals soft tissue preservation indicating a dorsal lobe associated with the tail fin, forming a wing-like stabilizing extension supported by neural spines, which likely enhanced maneuverability in open water.^[65] Plesiosaurs, with their paddle-like flippers, are reconstructed with low-profile dorsal ridges or soft tissue expansions along the vertebral column to counter roll and yaw, complementing their long-necked or short-necked body plans for efficient propulsion.^[66] Structurally, dorsal fins in marine mammals differ markedly from those in fish, consisting of dense fibrous connective tissue overlaying cartilaginous elements at the base, without the ray-like supports (lepidotrichia) typical of teleosts.^[67] In some dolphins, underlying muscles allow minor adjustments to fin rigidity, contributing to fine-tuned control during swimming.^[68] These fins serve specialized roles beyond basic stabilization, including aiding breaching in whales by providing leverage and balance during aerial leaps, which can exceed body length in height.^[69] In cold waters, dorsal fins facilitate thermoregulation through countercurrent heat exchange systems, where arteries and veins in the fin transfer heat to conserve core body temperature or dissipate excess warmth during exertion.^[70]^[62]

Evolutionary and Research Perspectives

Evolutionary Origins

The dorsal fin originated approximately 420 million years ago in early osteichthyans, evolving from a continuous median fin fold that extended along the dorsal midline of ancestral chordates.^[71] This primitive structure provided basic stabilization and propulsion, with fossil evidence from Devonian sarcopterygians like Eusthenopteron foordi (circa 375 million years ago) showing a well-developed dorsal fin composed of lepidotrichia (fin rays) supported by endoskeletal radials, marking an early diversification in median fin morphology.^[72] In more basal jawed vertebrates, such as the placoderm Dunkleosteus terrelli (circa 380 million years ago), a prominent dorsal fin is evident in fossil reconstructions, indicating its presence across early gnathostome lineages before the full radiation of osteichthyans.^[73] In sarcopterygian fishes, the dorsal fin co-evolved with paired fins like the pectorals, sharing developmental modules for endoskeletal support and ray formation, which facilitated enhanced maneuverability in aquatic environments.^[74] During the transition to tetrapods in the late Devonian (circa 370 million years ago), the dorsal fin was progressively reduced and ultimately lost as early tetrapods like Ichthyostega adapted to shallow-water and terrestrial locomotion, where midline fins became unnecessary for stability.^[75] Key evolutionary innovations include the appearance of spiny anterior rays in the dorsal fin of acanthomorph teleosts around 100 million years ago in the Early Cretaceous, enhancing defensive capabilities and contributing to their ecological dominance.^[76] Independent reductions occurred in lineages like batoids (rays and skates), where the dorsal fin is often diminutive or absent due to adaptations for benthic lifestyles and pectoral fin expansion.^[19] Comparative embryology reveals that dorsal fin patterning shares conserved Hox gene expression domains with ancestral chordates, where these transcription factors establish anteroposterior identity along the midline, a mechanism retained from early vertebrate fin folds to modern osteichthyan structures.^[77] This genetic continuity underscores the deep homology in median fin development across chordate evolution.^[78]

Modern Research Applications

Modern research on dorsal fins has advanced photo-identification techniques for tracking individual marine animals, leveraging unique shapes, notches, and markings on the fins. In 2025 studies of Rice's whales in the Gulf of Mexico, researchers developed a photo-identification catalog using dorsal fin attributes such as linear cuts and tissue loss to distinguish 25 genetically unique individuals, enabling non-invasive monitoring of this endangered population. Similarly, aerial drone-based photo-identification of humpback whales in 2025 improved re-sighting rates by capturing dorsal fin details, supporting long-term population assessments across 57 cetacean species. For sharks, dorsal fin patterns have proven reliable for individual identification; a 2025 study on blacktip reef sharks confirmed that unique fin markings allow for accurate tracking, enhancing understanding of population dynamics in coral reef ecosystems. Another 2025 analysis of oceanic whitetip sharks used dorsal fin photo-identification to assess demographics and fishery interactions, marking the first such application for this species. Dorsal fin spines serve as key structures for aging and population studies in commercially important fish like tuna, where annuli—annual growth rings—provide validated age estimates. A 2025 comparative study on Mediterranean Atlantic bluefin tuna analyzed annuli in dorsal fin spines and caudal vertebrae from reared specimens, estimating ages from 4 to 20 years and confirming the spines' reliability for growth modeling in overfished stocks. Biomechanical research employs advanced imaging to model dorsal fin function, revealing previously unobserved behaviors. Drone observations in 2025 demonstrated that white shark dorsal fins exhibit high flexibility, rotating toward objects in an investigatory manner, which informs hydrodynamic models of fin-mediated sensing and stability during hunting. Studies on swimming efficiency highlight interactions between dorsal and caudal fins, where coordinated motions enhance propulsion. A 2025 hydrodynamic analysis showed that undulating dorsal fins increase thrust and efficiency in bionic models by interacting with caudal fin beats, particularly at varying amplitudes that mimic fish locomotion. Another 2025 investigation found that opening dorsal and anal fins during caudal motion boosts overall swimming efficiency despite higher energy costs, optimizing performance in schooling fish. In conservation efforts, dorsal fin analysis detects fishery impacts, such as propeller-induced notches that signal human-whale interactions. For Rice's whales, 2025 photo-identification revealed deep triangular and round notches on dorsal fins attributable to fishing gear, underscoring threats to this critically endangered species with fewer than 100 individuals. AI-driven tools like iSharkFin, updated through 2025, use machine learning to identify shark species from dorsal fin images, achieving high accuracy for 39 species in wet fin trade samples and aiding enforcement of international protections. Developmental genetics research elucidates the embryonic origins of dorsal fin skeletons using mutant models. A 2024 zebrafish study identified the sclerotome—a somite-derived structure—as the primary source of dorsal and anal fin skeletal cells; in smoothback mutants, reduced sclerotome expansion led to complete dorsal fin loss and partial anal fin agenesis, highlighting its essential role in median fin development.

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Oct 26, 2015 · This study provides evidence that spiny dogfish control movements of the first dorsal fin during steady swimming to stabilize body position. In ...
[36]
Hydrodynamic function of dorsal fins in spiny dogfish and bamboo ...
These data suggest that the first dorsal fin in spiny dogfish has primarily a stabilizing function, while the second dorsal fin has a propulsive function.
[37]
Hydrodynamic function of dorsal and anal fins in brook trout ...
Jan 15, 2007 · Recent kinematic and hydrodynamic studies on fish median fins have shown that dorsal fins actively produce jets with large lateral forces.
[38]
Genomes of Two Billfishes Provide Insights into the Evolution of ...
Feb 3, 2021 · The countercurrent heat exchangers, which consist of blood vessels, are crucial for endothermic teleosts for heat conserving. They can transport ...
[39]
Evolution and Consequences of Endothermy in Fishes
The evidence for two hypothesized advantages of endothermy in fishes, thermal niche expansion and enhancement of aerobic swimming performance, is analyzed
[40]
Enhanced thermoregulation abilities of shortfin mako sharks as the ...
Aug 29, 2025 · We show that regionally endothermic shortfin mako sharks have enhanced thermoregulatory ability with an order higher warming rate than ...Missing: billfish | Show results with:billfish
[41]
Lionfish, Scorpionfish, and Stonefish Toxicity - StatPearls - NCBI
Summarize the frequency of encounters with poisonous fish. Describe ... Lionfish, venomous spines, dorsal, pelvic and anal fins Contributed by Karen ...Continuing Education Activity · Introduction · Etiology · Differential Diagnosis
[42]
The toxicology of greater weever (Trachinus draco): Clinical and ...
The greater weever (Trachinus draco) is a venomous benthic ray-finned fish belonging to the Trachinidae family, which comprises nine extant species of weever- ...
[43]
Gray Triggerfish – Discover Fishes - Florida Museum of Natural History
Feb 5, 2025 · The first dorsal fin has three spines that can be locked into an erect position for use as predator-defense and as an anchoring device. The ...Common Names · Importance To Humans · Biology<|separator|>
[44]
Deep-sea anglerfish - MBARI
This bulbous beast has a “fishing pole” projecting from its head. The first ray of its dorsal fin is modified into a filament (called an illicium) with a sac ...
[45]
Male courtship decisions are influenced by light environment ... - NIH
Sep 28, 2016 · Furthermore, light environments directly affect the perceived conspicuousness of male guppy multi-component colour sexual signals [7,21,22].Missing: dorsal | Show results with:dorsal
[46]
The ontogeny of courtship, colour and sperm production in male ...
Apr 1, 2005 · The onset of sexual behaviour in three age classes of male guppy Poecilia reticulata occurred in synchrony with the appearance of body ...<|control11|><|separator|>
[47]
The locomotor system of the ocean sunfish Mola mola (L.)
Propulsion is by unpaired dorsal and anal fins; a pseudocaudal fin ('clavus') acts as a rudder. Despite common perception, young sunfish are active predators ...Missing: sail drifting
[48]
Drone observations reveal white shark (Carcharodon carcharias ...
Sep 9, 2025 · Drone observations reveal white shark (Carcharodon carcharias) dorsal fins are highly flexible and possible investigatory structures. Carlos ...Missing: probing | Show results with:probing
[49]
Ageing Mediterranean Bluefin Tuna: A Comparative Study Between ...
This study provides a direct comparison of age estimates derived from dorsal fin spines and caudal vertebrae in Atlantic bluefin tuna.Missing: marker | Show results with:marker
[50]
Biological findings from a newly developed photo-identification ... - NIH
Sep 8, 2025 · Ten out of the 31 individually identified Rice's whales presented clean-cut lacerations or notches on the dorsal fin. Four whales had linear ...
[51]
Functional Innovations and the Conquest of the Oceans by ...
One of the characteristics that has strongly contributed to the ecological and evolutionary success of the spiny-rayed fishes is fin spines in dorsal and anal ...
[52]
Phylogenetic Classification of Living and Fossil Ray-Finned Fishes ...
Apr 18, 2024 · Here we provide an unranked phylogenetic classification for actinopterygian fishes based on a summary phylogeny of 830 lineages of ray-finned fishes.Missing: dual | Show results with:dual
[53]
Shark anatomy
Dec 19, 2016 · There are one or two fins present along the dorsal midline called the first and second dorsal fin. These are anti-roll stabilizing fins. These ...Missing: cartilaginous | Show results with:cartilaginous
[54]
Diversity of dermal denticle structure in sharks: Skin surface ...
Cartilaginous fishes (Chondrichthyes: sharks, skates, rays, and chimaeras) ... flexible dorsal fins, which result from a steady swimming trade off. J ...
[55]
Cartilaginous Fish – Skates, Rays and Chimaeras
Also, skates typically have a prominent dorsal fin while the dorsal fin is absent or greatly reduced in rays. Cartilaginous fish all breathe through 5 to 7 ...<|separator|>
[56]
BIO370-Lobe-finned Fishes - Udo Savalli
May 23, 2025 · Sarcopterygii Characteristics · Muscular lobed paired fins joined to body by single bone · Two dorsal fins with separate bases · Has a pulmonary ...
[57]
Development of the Pectoral Lobed Fin in the Australian Lungfish ...
Jul 21, 2021 · The evolutionary transition from paired fins to limbs involved the establishment of a set of limb muscles as an evolutionary novelty.
[58]
Phylogeny, biology, and evolution of acanthopterygian fish clades
Mar 13, 2025 · Most have two dorsal fins, slender caudal peduncles, and deeply forked caudal fins ... ray-finned fishes (Actinopterygii). Bull Peabody Mus Nat ...
[59]
Trophic resource partitioning among five flatfish species ...
Aug 7, 2025 · Five species of flatfishes in a tropical bay in south-eastern Brazil were studied to test the hypothesis that resource partitioning along ...
[60]
New burrowing catfish species found in Brazil river basin
Jul 4, 2025 · Researchers have discovered a long, slender new species of catfish from southern Brazil that, over time, has lost some of its fins.Missing: ecological variations tall tuna low flounder bottom- dwellers reduced
[61]
[PDF] A blacktip's black tip: The reliability of using dorsal‐fin patterns for ...
May 25, 2025 · By facilitating the identification of individual sharks based on their unique fin patterns, photo ID can improve our understanding of population.
[62]
Performance of iSharkFin in the identification of wet dorsal fins from ...
This study reports the current performance of the iSharkFin system, a machine learning technology which aims to allow users to identify the species of a wet ...
[63]
Dorsal Fins, From Killer Whales to Narwhals - Baleines en direct
The beluga has a dorsal ridge that, much like the previously-discussed cetaceans, is very useful. In this cold-water species, heat loss to the environment is a ...
[64]
Pinnipeds: Seals, Sea Lions, and Walruses | Smithsonian Ocean
The easiest way is to look at their ears—sea lions have small ear flaps while seal ears are nothing but small holes.Missing: identification | Show results with:identification
[65]
Photo-ID and Photogrammetry - SouWEST
Since 1912, hot brands have been used to give unique identifying markings to numerous pinniped species (e.g. northern fur seals, southern elephant seals, grey ...
[66]
Soft tissue preservation in a fossil marine lizard with a bilobed tail fin
Sep 10, 2013 · In mosasaurs, of course, the fin lobes are reversed, the ventral one being the primary thrust generator, but together with the dorsal fin lobe, ...<|separator|>
[67]
Plesiosaurs by the Tail | National Geographic
Dec 10, 2013 · Drawing from this body of evidence, Wilhelm identified skeletal features that are consistent with a tail fin in the skeletons of the plesiosaurs ...
[68]
What Is a Dolphin's Dorsal Fin Used For?
May 17, 2019 · The fin and flukes are composed of dense, fibrous connective tissue lacking bone, cartilage, or muscle.Missing: structure rays
[69]
Physiology - Dolphin Research Center
Inside the flukes, dorsal fin, and pectoral fins are a bunch of arteries surrounded by a network of veins. Along the surface of the skin, in these areas, are ...
[70]
Energetic and physical limitations on the breaching performance of ...
Mar 11, 2020 · We found that breaching whales use variable underwater trajectories, and that high-emergence breaches are faster and require more energy than predatory lunges.
[71]
All About Killer Whales - Adaptations | United Parks & Resorts
Thermoregulation · Some arteries of the flippers, flukes, and dorsal fin are surrounded by veins. · When a killer whale dives, circulation decreases at the skin, ...Missing: breaching | Show results with:breaching
[72]
Adipose fin development and its relation to the evolutionary origins ...
Jan 24, 2019 · The dorsal, anal and caudal fins of vertebrates are proposed to have originated by the partitioning and transformation of the continuous ...
[73]
How fish fins evolved just before the transition to land - UChicago ...
Dec 30, 2019 · Fin rays are actually formed by pairs of bones. In Eusthenopteron, for example, the dorsal, or top, fin ray was slightly larger and longer than ...
[74]
Dunkleosteus Reconstruction - Palaeontologia Electronica
The dorsal fin outline of Dunkleosteus is unknown. The few pachyosteomorphs with known dorsal fins (all aspinothoracidans) show highly diverse shapes.
[75]
Fin ray patterns at the fin-to-limb transition - PNAS
Dec 30, 2019 · Rays are usually segmented, branching, and more robust on the anterior, or leading edge, of the fin (25, 26).
[76]
The Fish–Tetrapod Transition: New Fossils and Interpretations
Mar 17, 2009 · Its tetrapod-like features include loss of all its midline fins—dorsal and anal—a flatter skull, a much longer snout, and larger and more ...
[77]
New insights on early evolution of spiny-rayed fishes (Teleostei
In this study, we provide new insights into the early evolution of the acanthomorphs and the euteleost allies based on the phylogenetic analysis of a newly ...
[78]
(PDF) Evidence that mechanisms of fin development evolved in the ...
Aug 9, 2025 · The dorsal (unpaired) fin fold develops from somite mesoderm (sclerotome and dermomyotome) and neural crest cells (Freitas et al., 2006) ...Missing: osteichthyans | Show results with:osteichthyans
[79]
Teleost Hox code defines regional identities competent for ... - PNAS
Jun 11, 2024 · We show that different hox genes regulate dorsal fin development between zebrafish and medaka and may have been transferred during evolution ...