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Chasmosaurus

Chasmosaurus is a genus of chasmosaurine ceratopsid dinosaur that lived during the Late Cretaceous Campanian stage, approximately 76 to 75 million years ago, in western North America. The name derives from Greek words meaning "opening lizard", referring to the large openings in its frill.^[1] This quadrupedal herbivore, known from fossils primarily in the Dinosaur Park Formation of southern Alberta and Saskatchewan, Canada, measured roughly 5 meters in length and weighed about 1.5 tonnes.^[2]^[3] It is distinguished by its large, shield-like bony frill featuring prominent parietal fenestrae (openings), short triangular postorbital horns, and a small nasal horn, adaptations possibly related to display or defense.^[1] The genus was first described in 1914 based on a partial skull from the Belly River Group, with the type species C. belli named in honor of geologist Walter Bell.^[1] A second species, C. russelli, was named in 1940 by C.M. Sternberg, differentiated by the angle of the posterior parietal bar embayment on the frill.^[1] Additional specimens, including skulls up to 90 cm in rostral-to-epijugal length, indicate variation from juveniles to adults, with overlapping stratigraphic ranges in the formation.^[1] A related chasmosaurine, formerly known as C. mariscalensis but now classified as Agujaceratops mariscalensis, has been reported from the Aguja Formation in Texas, suggesting a broader distribution across Laramidia. As a megaherbivore, Chasmosaurus likely fed on low-lying vegetation using its beak-like mouth and shearing dental battery, inhabiting coastal plain environments with rivers and floodplains.^[4] Its frill, with large rounded fenestrae and triangular epiparietals, may have served in intraspecific signaling or thermoregulation, though interpretations vary.^[4] Over a dozen skulls and partial skeletons have been recovered, contributing to understanding ceratopsid diversity and evolution in the final millions of years before the Cretaceous-Paleogene extinction.^[1]

Etymology and Discovery

Naming and Etymology

The genus Chasmosaurus was formally established in 1914 by Canadian paleontologist Lawrence M. Lambe, who coined the name to replace the preoccupied Protorosaurus for material previously assigned to Monoclonius. The etymology derives from the Greek chasma (χάσμα), meaning "opening" or "cleft," combined with sauros (σαῦρος), meaning "lizard," directly referencing the prominent fenestrae—large openings—in the skull's parietal frill.^[5]^[6] The type species, C. belli, originated from Lambe's initial description in 1902, based on the holotype specimen CMN 491, a partial parietal bone recovered from the Belly River Formation in Alberta, Canada. This species was named in honor of its collector, Walter Bell, an assistant with the Geological Survey of Canada. Lambe noted the holotype's distinctive frill features, including a narrowed midline bar and indications of bilateral fenestrae, which set it apart from other ceratopsians known at the time. This naming occurred amid intensive early 20th-century paleontological expeditions by the Geological Survey of Canada, which systematically explored the Late Cretaceous exposures of western Canada following initial discoveries in the 1890s. These efforts, led by figures like Lambe, revealed a rich ceratopsian fauna and contributed to the rapid expansion of knowledge about North American dinosaurs during that era.^[6]

Fossil Discoveries and Localities

The first fossils attributed to Chasmosaurus were discovered in 1898 by paleontologist Lawrence Morris Lambe of the Geological Survey of Canada along the Red Deer River Valley in Alberta, Canada. These initial specimens, collected from sediments of the Belly River Formation (now considered part of the overlying Dinosaur Park Formation), included a partial parietal bone from the neck frill, which Lambe initially described as a new species of Monoclonius (M. belli) in 1902 before reclassifying the genus as Chasmosaurus in 1914. The holotype for C. belli, cataloged as CMN 491 at the Canadian Museum of Nature, remains a fragmentary but diagnostic element that highlights the distinctive fenestrations in the frill.^[7] Subsequent excavations in the early 20th century expanded knowledge of Chasmosaurus, with significant material recovered from major quarries in the Steveville area of what is now Dinosaur Provincial Park, Alberta. In 1913, digs by Charles H. Sternberg and his sons uncovered multiple individuals from a bonebed, while further finds in the 1920s by Barnum Brown of the American Museum of Natural History added postcranial elements and additional skull fragments to collections. A key specimen from these efforts is the holotype of C. russelli (ROM 1223, Royal Ontario Museum), a partial skull from the lower Dinosaur Park Formation that represents an earlier stratigraphic occurrence than most C. belli material. These quarries have yielded over a dozen referred specimens, including partial skeletons that provide insights into ontogenetic variation.^[7]^[8] Fossils of Chasmosaurus are primarily known from western Canada, with the majority unearthed in Alberta (e.g., Dinosaur Provincial Park, Manyberries) and Saskatchewan (e.g., Saskatchewan Landing Provincial Park), though rare referred material has been reported from the United States in Montana's Judith River Formation. Stratigraphically, these occurrences span the late Campanian stage of the Late Cretaceous, approximately 77 to 74 million years ago, within the Oldman Formation (lower equivalent) and the Dinosaur Park Formation (upper sections), where Chasmosaurus coexisted with diverse ornithischian and theropod assemblages.^[7]^[4] In the 2010s, renewed fieldwork in the Dinosaur Park Formation revealed additional bonebeds containing disarticulated remains of multiple Chasmosaurus individuals across age classes, reinforcing evidence for gregarious social structures similar to those inferred in other ceratopsids. These discoveries, including well-preserved juvenile specimens, have enhanced understanding of population dynamics without altering the core taxonomic framework. No major new localities or assemblages have been reported since 2020.^[8]^[7]

Description

Overall Morphology and Size

Chasmosaurus was a medium-sized chasmosaurine ceratopsid, with skeletal reconstructions indicating an estimated body length of 4.3–5 meters and a body mass of 1.5–3.5 tonnes.^[3]^[4] As a quadrupedal herbivore, it possessed a robust build characterized by a short, deep torso supported by powerful limbs suited for supporting its weight while browsing low vegetation in its Late Cretaceous habitat.^[9] The overall body plan reflected the typical ceratopsian form, with a bulky frame, a long tail for balance, and a low-slung posture that facilitated access to ground-level plants.^[4] Skin impressions preserved in association with Chasmosaurus specimens, such as the second Monoclonius skeleton described by Brown (now recognized as Chasmosaurus), reveal a covering of large, polygonal scales across the body, with the frill featuring osteoderms (epiparietals and episquamosals). A 2025 study suggests the enlarged feature scales on Chasmosaurus skin may have functioned in sensation or thermoregulation rather than display.^[10]^[11]^[12] These features suggest a textured integument similar to that in other mature chasmosaurines. Hypotheses regarding sexual dimorphism in Chasmosaurus propose potential differences in body size between sexes, based on postcranial variations observed in specimens like those attributed to C. belli, though such distinctions remain unconfirmed and may instead reflect ontogenetic or interspecific variation.^[3] In comparison to its later relative Triceratops, Chasmosaurus exhibited a more gracile morphology, with proportionally slimmer limbs and a less massive frame that likely enhanced agility within the forested coastal plain environments of the Dinosaur Park Formation.^[4]^[9] This build underscores its adaptation as a nimble browser among dense vegetation, distinct from the heavier, more robust form of Triceratops in open terrains.^[13]

Skull and Frill Features

The skull of Chasmosaurus is elongated, typically measuring up to 1 meter in length from the rostrum to the rear of the frill in adult specimens.^[14] This cranial structure features large parietal fenestrae, which are distinctive triangular openings in the frill unique to the genus, often exceeding 400 mm in length and comprising a significant portion of the parietal bone surface.^[14] These fenestrae contribute to the lightweight yet expansive frill, which spans approximately 1.5 meters in width.^[4] The horns of Chasmosaurus are relatively modest compared to other ceratopsids. Brow horns, projecting from the postorbital region, are short, measuring 35–63 cm in length, and curve posteriorly with rounded apices that show evidence of remodeling in mature individuals.^[4] In C. mariscalensis, a small nasal horn, typically 15 cm long and transversely compressed, arises from the nasals; in C. belli, it is longer, up to approximately 25 cm.^[15]^[14] The frill margin is ornamented with 12–16 epoccipitals, triangular bony projections that are partially co-ossified in adults and vary in number along the squamosal and parietal edges, with 6–10 per side commonly observed.^[14] The frill itself is vascularized, exhibiting extensive grooves indicative of blood vessel networks, potentially aiding in thermoregulation or display, though its primary structural role is protective.^[14] Species variations include a narrower frill and longer brow horns in C. russelli compared to C. belli.^[14] The jaw apparatus is adapted for herbivory, featuring a battery of shearing teeth arranged in functional rows for grinding plant material, with up to 40 teeth per maxillary side and a similar count in the dentary.^[14] A beak-like predentary bone at the front of the lower jaw facilitates cropping vegetation.^[14] In juveniles, skull features differ markedly from adults, with smaller fenestrae, smoother frill margins lacking prominent epoccipitals, and more curved, less developed horncores, as evidenced by growth series specimens showing progressive ossification and elongation.^[14]

Postcranial Skeleton

The postcranial skeleton of Chasmosaurus is characterized by a robust vertebral column adapted for supporting the animal's quadrupedal posture. The cervical series consists of nine vertebrae, with the first three fused into a syncervical unit, followed by six additional cervicals that are amphiplatyan and wider than tall. The dorsal vertebrae number 18, also amphiplatyan, contributing to a flexible yet sturdy presacral region. The sacral vertebrae form a fused block of seven elements, including two or more dorsosacrals and caudosacrals, providing enhanced stability for weight transfer to the hindlimbs during locomotion.^[16] The caudal series includes at least 20 vertebrae, forming a moderate-length tail that tapers distally.^[16] The limb elements reflect Chasmosaurus's graviportal build, with forelimbs noticeably shorter than the hindlimbs at a ratio of approximately 0.7:1, facilitating a lower anterior posture. The humerus measures 50–60 cm in length, featuring a prominent deltopectoral crest extending nearly half its shaft for major muscle attachments. The femur is longer, at 70–80 cm, with a proximal fourth trochanter positioned relatively high on the shaft. The manus follows a phalangeal formula of 2-3-4-3-2, with the terminal phalanges of digits I–III expanded and hoof-like, supporting weight-bearing on keratinous sheaths, while digits IV–V are reduced. The pelvic girdle includes a broad, flaring ilium that provides extensive attachment surfaces for powerful hindlimb musculature, aiding propulsion. The rib cage comprises numerous dorsal ribs, with anterior ones Y-shaped and posterior ones featuring shorter capitular processes, enclosing the thoracic cavity without ventral reinforcement. Notably, gastralia are absent, allowing greater flexibility in the abdominal region compared to taxa that possess them.^[16] The tail, supported by chevron bones along its length, likely functioned in maintaining balance during movement, countering shifts in the body's center of mass induced by the large skull and frill. Specimens from bonebed assemblages reveal evidence of pathologies, including healed fractures in limb bones such as the metacarpals and phalanges, indicative of stress from habitual quadrupedal locomotion or intraspecific interactions.^[16]^[17]

Classification

Historical Taxonomy

Chasmosaurus was initially described by Lawrence M. Lambe in 1902 as Monoclonius belli, based on fragmentary skull material from the Belly River Formation in Alberta, Canada, placing it within the ceratopsian genus Monoclonius alongside other horned dinosaurs known at the time.^[14] In 1914, Lambe recognized the distinctive frill fenestrae in additional specimens and erected the new genus Chasmosaurus for the species, formally establishing it as a distinct ceratopsid with M. belli as the type species.^[14] By 1915, Lambe further classified Chasmosaurus within the newly proposed subfamily Chasmosaurinae, emphasizing its elongate frill with large openings as a defining trait, distinguishing it from the shorter-frilled Centrosaurinae.^[14] During the 1930s and 1940s, ongoing excavations in Alberta's Belly River Group, including the Dinosaur Park Formation, yielded more complete specimens that expanded the genus. Barnum Brown named Chasmosaurus kaiseni in 1933 based on skull AMNH 5401 exhibiting elongate postorbital horns from the same formation, while Charles M. Sternberg named Chasmosaurus russelli in 1940 based on a skull with elongate postorbital horns from the same formation.^[14] These additions reflected a period of taxonomic proliferation driven by new quarry discoveries in the 1920s through 1970s, which often led to over-splitting of species due to observed intraspecific variation in frill shape and horn length, interpreted at the time as diagnostic interspecific differences.^[14] By the 1980s, the recognized species included C. belli, C. russelli, C. kaiseni, and others like C. brevirostris and C. canadensis, with Thomas Lehman proposing C. mariscalensis in 1989 from Texas material in the Aguja Formation, further broadening the genus's geographic scope.^[14] Key revisions in the late 20th century began to address this fragmentation. Lehman (1990) analyzed frill morphology across chasmosaurines, grouping Chasmosaurus species based on shared fenestration patterns and suggesting that variations in C. belli and C. russelli represented a morphological continuum possibly influenced by ontogeny or sexual dimorphism, rather than separate taxa.^[14] Early debates emerged on the monophyly of Chasmosaurus, with some researchers questioning whether the genus formed a natural clade given the gradational traits observed in specimens from stacked bonebeds, hinting at potential synonymy but stopping short of full resolution before cladistic methods gained prominence.^[14] Pre-2000 synonym lists typically retained C. belli as valid, with C. russelli and C. kaiseni considered dubious or junior synonyms by some, underscoring the challenges posed by the abundant but variable fossil record from mid-century quarries.^[14]

Phylogenetic Position and Species Validity

Chasmosaurus is recognized as a basal member of the chasmosaurine ceratopsids within Ceratopsidae, consistently recovered in phylogenetic analyses as a relatively early-diverging taxon in the subclade. Recent specimen-based cladistic studies position it as the sister group to Vagaceratops or, in some matrices, Mercuriceratops, based on shared derived traits such as the configuration of epiparietals along the frill margin and moderate development of parietal fenestrae.^[14] These analyses employ comprehensive character matrices exceeding 150 traits, including cranial features like horn orientation (e.g., postorbital horn projection angles) and frill fenestrae size and position, alongside postcranial elements such as sacral rib fusion patterns. Phylogenetic trees from these datasets indicate the divergence of Chasmosaurinae from Centrosaurinae around 80 million years ago during the early Campanian, marking the onset of the North American ceratopsid radiation.^[14]^[18] Regarding species validity, Chasmosaurus belli and C. russelli are upheld as distinct and valid based on diagnosable differences in frill morphology, particularly the depth and angle of the posterior parietal embayment (shallow in C. belli versus deeper in C. russelli). The genus Mojoceratops, erected in 2009 for material from the Dinosaur Park Formation, has been synonymized with C. russelli due to overlapping diagnostic traits in epiparietal arrangement and frill shape, rendering it a junior synonym.^[14] The taxonomic status of Vagaceratops irvinensis, described in 2010 from upper Dinosaur Park Formation strata, remains debated but is generally considered potentially distinct from Chasmosaurus, supported by unique features like a reduced median parietal bar and straighter lateral rami; however, some analyses suggest it represents a transitional form or close relative rather than a separate genus. Similarly, the holotype of C. kaiseni (AMNH 5401) is regarded as referable to Chasmosaurus sp. due to insufficient preservation for species-level diagnosis, with no support for assignment to more basal ceratopsians like Protoceratops.^[14] Phylogenetic refinements from 2019–2020 studies, including expanded stratigraphic and morphological data from the Dinosaur Park Formation, have clarified the sequential succession of chasmosaurines (e.g., C. belli preceding Vagaceratops-like forms) and reinforced the basal positioning of Chasmosaurus within the subclade, with no new species named since 2020.

Paleobiology

Diet and Feeding Mechanisms

Chasmosaurus was a herbivorous dinosaur adapted for low-level browsing, targeting vegetation in the understory of floodplain forests, such as ferns and early angiosperms.^[19] Its quadrupedal stance and body proportions limited maximum feeding height to approximately 1 meter above the ground, allowing access to low-growing herbaceous plants without the need for bipedal rearing.^[19] This foraging strategy facilitated niche partitioning with coexisting taller herbivores, such as hadrosaurs, which could reach shrubs and tree foliage up to 5 meters high when standing bipedally, thereby reducing dietary competition in the Dinosaur Park Formation ecosystem.^[19] The skull of Chasmosaurus featured a robust dental battery suited for processing tough, fibrous vegetation, with multiple successional teeth arranged in families within the upper and lower jaws. Microwear patterns indicate that the teeth were used for shearing and grinding abrasive plant material like leaves and twigs. Jaw mechanics in Chasmosaurus emphasized efficient cropping and mastication, with a sharp, toothless beak (rhamphotheca) at the jaw tip for grasping and clipping vegetation, followed by scissor-like shearing via the dental battery. The primary occlusal motion was orthopalinal (back-and-forth along the jaw axis), producing scoring on tooth surfaces, though limited propalinal (fore-aft) components may have aided in shredding tougher fibers; transverse motion was absent, distinguishing ceratopsid feeding from that of hadrosaurs. The frill and overall cranial architecture permitted unobstructed head positioning for ground-level browsing, integrating with postcranial morphology to support this specialized herbivory without interference from ornamentation.^[19] Inferences from dental microwear and morphology confirm an exclusively plant-based diet, with scratch-dominated patterns consistent with consumption of woody browse and no indicators of animal matter. The frill of Chasmosaurus is hypothesized to have served primarily as a structure for species recognition and sexual display, with its elaborate shape and fenestrae facilitating visual signaling among individuals. These features align with patterns of positive allometry in ceratopsian cranial ornaments, indicating socio-sexual selection as a driver of frill evolution. Recent analysis of epidermal scales in Chasmosaurus suggests morphological stasis in skin growth, potentially supporting display functions.^[20] In terms of defense, the short brow horns of Chasmosaurus imply limited use for direct intraspecific combat. The frill itself may have functioned as a shield against predators, with its broad, bony margin offering passive protection to the neck while the animal faced threats head-on using its beak or herd positioning. Fossil aggregations in the Dinosaur Park Formation suggest herding behavior in Chasmosaurus, with multiple individuals preserved in close proximity indicating social grouping for protection and coordination.^[21] Such bonebeds, combined with evidence consistent with intraspecific interactions, support interpretations of complex social dynamics, including possible cooperative defense. The fenestrae in the Chasmosaurus frill may have played a sensory role, potentially amplifying low-frequency sounds for communication within herds, though direct evidence remains circumstantial.^[4] The role in thermoregulation remains debated. Ontogenetic changes in the frill, including rapid elongation and textural shifts during subadult stages, likely enhanced its role in mate attraction, with growth spurts coinciding with sexual maturity to maximize display efficacy.^[22]

Growth, Reproduction, and Paleoecology

Chasmosaurus exhibited rapid growth during its early life stages, transitioning from juveniles with striated bone textures indicative of fast deposition to adults with smoother surfaces. Bone histology reveals fibrolamellar bone tissue in younger individuals, supporting a quick juvenile phase where body length increased from approximately 1 m at hatching to subadult sizes within a few years. Lines of arrested growth (LAGs) in long bones indicate annual pauses in deposition, with most specimens showing 5–15 LAGs, suggesting individuals reached skeletal maturity around 9–10 years and could live up to 20 years or more. Population age structures from multiple specimens demonstrate a bias toward younger adults, consistent with high mortality rates in early ontogeny.^[23] As an ornithischian dinosaur, Chasmosaurus was oviparous, laying eggs in clutches, though specific nest sites remain unknown for the genus. Inferences from related ceratopsians, such as Protoceratops, suggest possible parental care, evidenced by juvenile-only clusters that may indicate protection or group rearing post-hatching.^[24] Sexual maturity likely occurred around 5–7 years, coinciding with the development of cranial ornamentation for display, based on ontogenetic series showing frill expansion in subadults. Chasmosaurus inhabited the alluvial coastal plains of the Dinosaur Park Formation in what is now southern Alberta, Canada, during the late Campanian (~77–75 Ma), an environment featuring meandering rivers, floodplains, and conifer-dominated forests.^[25] The climate was warm and humid, with seasonal monsoons driving periodic flooding and supporting high plant productivity.^[26] It coexisted with abundant hadrosaurids such as Gryposaurus and Parasaurolophus, as well as centrosaurine ceratopsids, ankylosaurs like Euoplocephalus, and predators including the tyrannosaurid Gorgosaurus.^[25] Chasmosaurus was relatively common in this assemblage, comprising a significant portion of ceratopsid fossils, though its abundance declined toward the formation's upper levels around 74 Ma, possibly linked to rising sea levels and habitat shifts.^[27] Bonebeds containing multiple Chasmosaurus individuals, studied in the 2010s, provide evidence of social grouping and potential seasonal migrations, as suggested by taphonomic patterns and associated fluvial deposits indicating movement along river systems.^[28]

References

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A Re-Evaluation of the Chasmosaurine Ceratopsid Genus ...
Jan 4, 2016 · The chasmosaurine ceratopsid Chasmosaurus is known from the Upper Cretaceous (Campanian) Dinosaur Park Formation of southern Alberta and Saskatchewan.
[2]
[PDF] Creature Guide - Philip J. Currie Dinosaur Museum
Hooked, parrot-like beak with powerful jaw closing muscles. Chasmosaurus: (Caz-mo-soar-us). Ceratopsid from late Cretaceous Alberta. Roughly 5 meters long ...
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[PDF] The Canadian Field-Naturalist - Semantic Scholar
Chasmosaurus are modest-sized (~1500 kg) ceratop- sid dinosaurs known from the upper Campanian-aged beds (~75 Ma [million years ago]) of western North.
[4]
New insights into chasmosaurine (Dinosauria - NIH
Jul 3, 2018 · Chasmosaurine ceratopsids are well documented from the Upper Cretaceous (Campanian) Dinosaur Park Formation (DPF) of southern Alberta and ...Missing: facts | Show results with:facts
[5]
[PDF] THE OTTAWA NATURALIST - Utah Geological Survey
The present paper is descriptive of a skull representing a new genus and species of trachodont, and of that of Chasmosa.urus belli, both forming part of last ...
[6]
https://www.biodiversitylibrary.org/item/18024#page/159/mode/1up
[7]
A Re-Evaluation of the Chasmosaurine Ceratopsid Genus ...
Jan 4, 2016 · The chasmosaurine ceratopsid Chasmosaurus is known from the Upper Cretaceous (Campanian) Dinosaur Park Formation of southern Alberta and Saskatchewan.Missing: Steveville | Show results with:Steveville
[8]
Systematics of Chasmosaurus - new... - F1000Research
Dec 17, 2015 · For this paper, I have chosen to use Chasmosaurus. Other ... Lambe 1915. Chasmosaurus Lambe 1914. Euceratopsia is a node-based ...
[9]
Systematics of Chasmosaurus - new... | F1000Research
### Summary of Chasmosaurus Morphology from https://f1000research.com/articles/4-1468/v1
[10]
A complete skeleton of the horned dinosaur Monoclonius, and ...
A complete skeleton of the horned dinosaur Monoclonius, and description of a second skeleton showing skin impressions.
[11]
A juvenile chasmosaurine ceratopsid (Dinosauria, Ornithischia) from ...
Jan 11, 2016 · An articulated, 1.5 m long skeleton of a juvenile Chasmosaurus, lacking only the front limbs, pectoral girdles, and terminal caudal vertebrae, was collected ...Missing: bonebed | Show results with:bonebed
[12]
Spiclypeus shipporum gen. et sp. nov., a Boldly Audacious New ...
The tooth row (318 mm long) extends most of the length of the dentary, and although poorly preserved anteriorly, there is room to accommodate no less than 24 ...
[13]
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[14]
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None
### Summary of Skull and Frill Anatomy of Chasmosaurus mariscalensis (Forster et al., 1993)
[16]
https://doi.org/10.11646/zootaxa.2963.1.1
[17]
Habitual locomotor behavior inferred from manual pathology in two ...
Rega et al. (2010) described patterns in three Chasmosaurus specimens of bone injury and degeneration involving metacarpal I and phalangeal elements. These ...
[18]
Patterns of divergence in the morphology of ceratopsian dinosaurs
Mar 21, 2018 · Node 2 represents the clade Ceratopsoidea, encompassing Centrosaurinae (orange branch) and Chasmosaurinae (blue branch), within which the ...
[19]
https://doi.org/10.1186/1472-6785-13-14
[20]
Redescription of UALVP 40, an unusual specimen of Chasmosaurus ...
Aug 5, 2015 · Redescription of UALVP 40, an unusual specimen of Chasmosaurus Lambe, 1914 ... Ottawa Naturalist, 27: 145–155. Go to Citation. Google ...
[21]
Sexual selection in Ceratopsia - Palaeontologia Electronica
Positive allometry for exaggerated structures in the ceratopsian dinosaur Protoceratops andrewsi supports socio-sexual signaling.
[22]
Bone resorption, bone lesions, and extracranial fenestrae in ...
Centrosaurines and chasmosaurines occasionally have large extra fenestrae and shallow, 'punched out lesions' on the frill (Tanke and Farke 2007) . These ...
[23]
Evidence of Combat in Triceratops | PLOS One - Research journals
This pattern is consistent with Triceratops using its horns in combat and the frill being adapted as a protective structure for this taxon.
[24]
[PDF] Systematics of Chasmosaurus - F1000Research
Dec 17, 2015 · Chasmosaurus takes this feature to an extreme7, resulting in an elongate rostrum, with the rostrum depth being 50% or less the skull's ...
[25]
The thermoregulatory functions of the Triceratops frill and horns
The results suggest very high and uniform heat flow through the parietal frill, maintaining mean frill temperatures between 0–4°C below the body core.Missing: Chasmosaurus | Show results with:Chasmosaurus
[26]
The evolution of 'bizarre structures' in dinosaurs: biomechanics ...
Dec 22, 2010 · The two general classes of explanation of bizarre structures in dinosaurs relate to function and display (including sexual selection, social ...Testing Mechanical... · Intraspecific · Dinosaurs And Sexual...
[27]
(PDF) A Re-Evaluation of the Chasmosaurine Ceratopsid Genus ...
Aug 6, 2025 · Background: The chasmosaurine ceratopsid Chasmosaurus is known from the Upper Cretaceous (Campanian) Dinosaur Park Formation of southern Alberta ...
[28]
A New Mass Mortality of Juvenile Protoceratops and Size ...
The juvenile group fills a key gap in the available information on aggregations in younger ceratopsians. Although we support the general hypothesis that many ...
[29]
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Oct 28, 2019 · Modern megaherbivore community richness is limited by bottom-up controls, such as resource limitation and resultant dietary competition.
[30]
(PDF) Competition structured a Late Cretaceous megaherbivorous ...
Oct 7, 2019 · Modern megaherbivore community richness is limited by bottom-up controls, such as resource. limitation and resultant dietary competition.
[31]
Calibrating geologic strata, dinosaurs, and other fossils at Dinosaur ...
An increase in rate above the Oldman–Dinosaur Park formational contact conforms to a regionally expressed pattern of increased accommodation at ∼76.3 Ma across ...
[32]
From Bonebeds to Paleobiology: Applications of Bonebed Data
Bonebeds are of interest to paleobiologists because they yield large numbers of fossils and, thus, provide important morphologic and taxonomic data sets ...