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Javan rusa

The Javan rusa (Rusa timorensis), also known as the Sunda sambar, is a medium-sized deer species belonging to the family Cervidae, characterized by its robust build, grayish-brown coat, large rounded ears, and short legs.^[1]^[2] Males typically weigh 70–135 kg and stand 95–110 cm at the shoulder, featuring lyre-shaped, three-tined antlers up to 95 cm long, while females are smaller at 50–90 kg with no antlers.^[2] Native to the tropical islands of Indonesia—including Java, Bali, and the Lesser Sunda Islands—and East Timor, it has been introduced to regions such as Australia, New Zealand, and New Caledonia, where it sometimes impacts local ecosystems.^[2]^[3] This adaptable herbivore inhabits a range of environments, from deciduous forests and grasslands to shrublands, marshes, and agricultural plantations, often favoring forest edges and areas up to 900 m elevation.^[1]^[2] Its diet consists primarily of grasses, leaves, bark, fruits, and occasionally seaweed, obtained through browsing and grazing, with minimal need for free water as it derives hydration from vegetation.^[1]^[3] Behaviorally, the Javan rusa is largely nocturnal or crepuscular, living in loose herds of 5–25 individuals led by a dominant male, who becomes aggressive and vocal during the July–September rutting season, often decorating antlers with grass or twigs to attract mates.^[1]^[3] Reproduction involves a polygynous mating system, with an eight-month gestation period yielding typically one fawn (rarely two), which is weaned at 6–8 months and reaches sexual maturity at 18–24 months.^[1] Classified as Vulnerable on the IUCN Red List since 2008, the Javan rusa faces ongoing population decline, estimated at fewer than 10,000 mature individuals, primarily due to habitat loss from deforestation and agriculture, poaching for meat and traditional medicine, and human-wildlife conflicts.^[3] Despite local abundance in protected areas, its fragmented range and sensitivity to invasive species like Acacia further threaten survival, prompting conservation efforts focused on habitat restoration and anti-poaching measures in Indonesia.^[3]

Taxonomy and nomenclature

Scientific classification

The Javan rusa belongs to the order Artiodactyla, family Cervidae, subfamily Cervinae, and genus Rusa within the deer family. Its binomial name is Rusa timorensis (de Blainville, 1822), originally described from specimens on Timor Island in the Journal de Physique, de Chimie, d'Histoire Naturelle et des Arts.^[4] Previously, it was classified under the genus Cervus as Cervus timorensis or Cervus russa, reflecting early 19th-century taxonomic groupings that lumped various Asian deer species together based on superficial similarities in antler structure and body form.^[5] The transfer to the genus Rusa, established to distinguish Southeast Asian deer from Eurasian Cervus species, was formalized in the late 20th century through revisions emphasizing cranial morphology, pelage patterns, and biogeographic isolation. Taxonomic revisions in the 20th century clarified the distinction of R. timorensis as a separate species from the sambar deer (Rusa unicolor), which had been conflated due to overlapping habitats and similar ecological roles in Indo-Malayan forests. This separation was driven by morphological analyses revealing differences in antler configuration—lyre-shaped and three-tined in R. timorensis versus more complex and multi-tined in R. unicolor—and supported by genetic studies indicating divergence during the Pleistocene, approximately 1.8 million years ago.^[3] Key contributions include Grubb's 1990 list of deer taxa, which recognized R. timorensis as distinct, and subsequent works by Groves and Grubb (2005, 2011) that integrated morphometric data from museum specimens across the Indonesian archipelago to affirm its species status. These revisions also addressed introgression risks from human-mediated introductions, highlighting the need for precise delineation to inform conservation.^[6] Seven subspecies of R. timorensis are currently recognized, primarily differentiated by geographic isolation across the Indonesian islands and associated variations in body size, antler length, and coat coloration. For instance, the nominate subspecies R. t. timorensis inhabits Timor and adjacent areas, featuring relatively smaller antlers (up to 60 cm) and a more uniform grayish-brown pelage adapted to open grasslands.^[3] R. t. russa, found on Java and Bali, exhibits larger overall size (males up to 115 kg) and more robust antlers (up to 95 cm), reflecting richer forest habitats that support greater nutritional intake.^[7] Another example is R. t. kangeanensis from the Kangean Islands, characterized by intermediate antler sizes and lighter dorsal markings, which aid camouflage in mixed shrubland environments. Other subspecies, such as R. t. floresiensis on Flores, R. t. macassaricus on Sulawesi, R. t. djonga on Bawean Island, R. t. moluccensis in the Moluccas, and R. t. renschi on Sumba, show further clinal variations in horn curvature and mane development, though ongoing genetic assessments question the full validity of some due to historical translocations.^[3] These distinctions underscore the species' adaptive radiation across Wallacean islands, with morphological traits often correlating to local vegetation density and predation pressures.^[5]

Common names and etymology

The Javan rusa, scientifically known as Rusa timorensis, is referred to by several common names reflecting its native range across Indonesian islands. Primary English names include Javan rusa, Sunda sambar, Timor deer, and Javan deer, with "rusa deer" also commonly used in contexts emphasizing its genus.^[8]^[9]^[1] The genus name Rusa derives from the Malay and Indonesian word for "deer," a term historically applied to various deer species in the Indo-Malayan region. The specific epithet timorensis originates from the island of Timor, the type locality where the species was first described based on specimens collected in the early 19th century.^[10]^[5] In Indonesia, regional variations highlight local linguistic influences, such as "rusa hutan" (forest deer) in Bahasa Indonesia, denoting its habitat in wooded areas. Historically, English nomenclature included "Moluccan rusa" for populations or subspecies from the Moluccan Islands, reflecting early colonial observations of introduced groups.^[11]^[12] The species holds cultural significance in Indonesian nomenclature, notably featured on the obverse of the 1988 500-rupiah banknote alongside the national coat of arms, symbolizing the nation's biodiversity and heritage.^[13]^[14]

Physical characteristics

Body size and appearance

The Javan rusa exhibits notable sexual dimorphism in body size, with males generally larger than females. Adult males typically measure 142–185 cm in head-body length and weigh 70–135 kg, while females are smaller, with lengths of approximately 125–145 cm and weights of 50–90 kg. Shoulder height is about 95–110 cm overall, with males reaching 100–120 cm and females 90–105 cm, contributing to their medium-sized build relative to other deer species.^[4]^[15]^[10]^[1]^[16] In appearance, the Javan rusa has a robust, somewhat stocky form with relatively short legs, a long and narrow face, large rounded ears, and a short tail measuring 15–20 cm. The coat is coarse and grayish-brown overall, often appearing rough, with darker tones on the back and hindquarters; the forehead and throat are lighter, typically grayish, while the underparts and inner thighs show yellowish-brown hues. Fawns are born without spots, featuring a plain reddish-tan coat with white underparts. Seasonal variations are minimal in their tropical range, though the coat may appear slightly lighter and greyer during drier periods compared to the reddish-brown tones in wetter seasons. The hooves are cloven with two main toes, enabling adaptation to diverse terrains from forests to grasslands.^[1]^[7]^[15]^[16]^[17]

Antlers and sexual dimorphism

The Javan rusa exhibits pronounced sexual dimorphism, most notably in the presence of antlers, which are exclusive to males. These antlers are lyre-shaped and typically feature three tines: the brow tine at the base, the bez tine midway along the beam, and the main beam extending upward. Adult antlers measure 50–80 cm in length on average, with exceptional records reaching 95 cm, and weigh approximately 2.5 kg.^[1]^[18]^[1] Antler growth follows an annual cycle, with casting peaking from December to January, after which new antlers emerge from the pedicles. Growth is rapid during the initial phase, covered in velvet, and reaches full hardness by May to June when the velvet is shed. In the Javan subspecies (Rusa timorensis russa), antlers tend to be longer than in other populations, such as those from Timor or the Moluccas, often exceeding 91 cm in West Java under optimal conditions.^[18]^[19] Beyond antlers, males display thicker necks, often with a darker, rougher mane, and possess larger skulls compared to females, contributing to their overall bulkier build. Females, lacking antlers, have more slender bodies and lighter frames. Both sexes bear tarsal scent glands on the inner hind legs, used for marking territory, as well as preorbital glands near the eyes for additional olfactory communication.^[20]^[21]^[22] Antler quality and size peak during prime adulthood but decline after approximately 10 years of age, coinciding with reduced reproductive productivity. In the wild, Javan rusa have an overall lifespan of 15–20 years.^[23]^[1]

Distribution and habitat

Native range

The Javan rusa (Rusa timorensis) is native to the Indonesian islands of Java, Bali, and Timor, including the territory of East Timor.^[24] These core areas represent the species' indigenous distribution within the Indo-Malayan and Australasian realms, shaped by historical biogeographic patterns across the Wallace Line.^[25] Two subspecies are recognized within this native range: R. t. russa, which occurs on Java and Bali, and R. t. timorensis, restricted to Timor.^[18] Populations of R. t. russa show genetic divergence between western and eastern Java as well as Bali, indicating limited gene flow and adaptation to island-specific conditions.^[25] On Timor, R. t. timorensis occupies similar ecological niches but with evidence of human-mediated influences on its spread beyond the main island. Historically, the Javan rusa was widespread across much of Java prior to colonial-era habitat alterations, occupying extensive lowland grasslands and forest edges.^[26] Pre-colonial records suggest a broad distribution covering central and eastern regions, with possible early introductions to nearby islands like Lombok and Sumbawa, where local populations have since gone extinct due to habitat loss and hunting.^[18] Today, the range on Java is highly fragmented, confined largely to protected areas such as Baluran National Park and Alas Purwo National Park in East Java, where viable populations persist amid ongoing threats.^[26] On Bali, remnants occur in Bali Barat National Park, while Timor's populations remain more continuous in eastern lowlands. The species primarily inhabits elevations from sea level to about 900 meters, favoring open grasslands and savannas, though some groups in Java's mountainous regions, such as the Yang Plateau near Mount Argopuro, extend up to 2,400 meters.^[18]^[27] This elevational flexibility allows adaptation to varied terrains, from coastal plains to upland plateaus, but current distributions emphasize lowland strongholds within national parks.

Introduced populations

The Javan rusa (Rusa timorensis) has been introduced to several non-native regions, primarily through human-mediated dispersal during the colonial era, with the main purposes being game hunting, ornamental parks, and trials for agricultural or livestock use. Earliest documented introductions trace to the Dutch colonial period in the Indian Ocean and Pacific islands; for instance, the species was brought to Mauritius around 1639 by Dutch settlers as a potential protein source, where it subsequently established wild populations. Similarly, in the Pacific, introductions began in the 19th century, often facilitated by European acclimatisation societies seeking to diversify local fauna for sport and economic purposes.^[28]^[29] Key modern introductions include Australia, where the Javan rusa was first released in the 1860s in New South Wales for hunting, leading to feral populations that spread to Queensland by the early 20th century. In New Caledonia, approximately 12 individuals of the subspecies R. t. russa were imported from Java in the 1870s for hunting and farming, resulting in a self-sustaining population across Grande Terre. Papua New Guinea saw introductions around 1900, initially near Port Moresby, where sizeable herds developed in coastal and grassland areas. Vanuatu and other Pacific islands, such as parts of the Solomon Islands, received the species in the late 19th to early 20th centuries for similar hunting purposes, though records are sparser. These efforts reflect broader colonial patterns of species translocation from Southeast Asia to island ecosystems.^[30]^[31]^[32]^[33]^[34]^[28] Establishment has been highly successful in many sites, particularly in tropical and subtropical environments resembling the native range, though with notable ecological consequences as an invasive species. In Australia, feral populations thrive in eastern coastal regions, numbering in the tens to hundreds of thousands across New South Wales and Queensland, where they compete with native herbivores like the swamp wallaby for forage and contribute to habitat degradation through overgrazing and browsing. New Caledonia hosts one of the largest introduced populations, estimated at several hundred thousand individuals, exerting pressure on endemic plants and altering forest understories. In Papua New Guinea, populations remain viable in southern lowlands, supporting local hunting but posing risks to agricultural lands. Mauritius sustains a stable herd, primarily managed for trophy hunting, while Vanuatu's populations are smaller and more localized, with limited expansion due to rugged terrain. Overall, these groups have adapted well, exhibiting high reproductive rates in the absence of natural predators.^[15]^[35]^[36]^[28]^[34] Most introduced populations derive from Javan stock (R. t. russa), reflecting the species' primary origin in Java, which has led to low genetic diversity and potential subspecies mixing in feral herds due to small founder numbers and isolation. Genetic studies in Australia reveal bottleneck effects, with minimal variation compared to native populations, increasing vulnerability to diseases but facilitating rapid adaptation to new habitats. In New Caledonia, analyses confirm descent from the initial 1870s imports, with ongoing gene flow limited by geography, resulting in homogeneous groups across the island. This genetic uniformity underscores the anthropogenic nature of these introductions and informs management strategies to mitigate invasiveness.^[37]^[32]^[28]

Habitat preferences

The Javan rusa (Rusa timorensis) primarily inhabits open grasslands, savannas, and forest edges within its native range on the islands of Java, Bali, and Timor. It thrives in tropical and subtropical moist lowland forests, montane forests, deciduous woodlands, and shrublands, while also tolerating monsoon forests and dry scrublands. This flexibility allows it to occupy a variety of ecosystems, from marshes to production forests and non-protected areas influenced by seasonal precipitation patterns.^[38]^[39]^[3] In terms of microhabitat needs, the species prefers areas with dense vegetative cover for protection, such as tall grasses along ecotones, and maintains proximity to water sources despite rarely drinking directly, instead deriving most hydration from foliage. It actively avoids dense rainforests, favoring transitional zones that balance concealment and access to open foraging spaces.^[38]^[1] The Javan rusa demonstrates adaptations to environmental challenges, including tolerance for seasonal droughts through efficient moisture extraction from vegetation and resilience to periodic fires in grassland habitats. Its altitudinal range extends from sea level to elevations of up to 900 meters, enabling occupation of both lowland and highland terrains with rugged features.^[39]^[1]^[3] Over time, habitat use has shifted from primary forests to secondary growth due to human-induced landscape alterations, with populations increasingly relying on modified environments like regrowth areas and invasive-dominated scrub. For example, in Baluran National Park, the proliferation of acacia species has transformed open savannas, leading to greater utilization of these secondary habitats by the deer.^[38]^[3]

Behavior and ecology

The Javan rusa (Rusa timorensis) displays primarily crepuscular activity patterns, with peaks in feeding and movement occurring at dawn and dusk, though individuals may shift to diurnal behaviors in habitats with low predator pressure.^[38] In areas with higher predation risks, such as native Indonesian forests, they tend to be more nocturnal, foraging under cover of darkness to minimize encounters with predators like leopards and dholes.^[1] Observations in protected island populations, such as Panaitan Island, confirm predominantly daytime activity, with feeding comprising over 50-70% of diurnal time budgets across age and sex classes.^[40] Socially, Javan rusa are gregarious, forming herds of 5-10 individuals in forested habitats, though groups can expand to 25 or more in open grasslands or savannas where visibility aids vigilance.^[38] Females and their young typically organize into matriarchal units led by adult does, which provide anti-predator signals and maintain group cohesion, while subadults remain integrated until maturity.^[40] Adult males, in contrast, are largely solitary outside the breeding season or form small bachelor pairs, avoiding larger aggregations to reduce competition.^[1] Seasonal dynamics influence grouping, with larger aggregations forming during the dry season to congregate near limited water sources, enhancing mutual protection in resource-scarce environments.^[41] In the wet season, increased forage availability promotes dispersal into smaller units or solitary foraging, reducing group sizes overall.^[38] Social structure peaks in complexity during the July–September rut, when males join female herds temporarily, heightening territorial interactions.^[38] Communication among Javan rusa relies on a combination of vocal, chemical, and visual signals to coordinate groups and alert to threats. Alarm calls, often described as barking or honking barks, serve to warn herd members of approaching danger, with recordings confirming their use in response to human disturbance.^[42] Scent marking occurs via secretions from hind-leg tarsal glands, rubbed on vegetation to delineate territories and signal reproductive status, particularly by males during the rut.^[1] Visual displays include foot-stomping to create vibrations and signal alarm, alongside postures like ear twitching and head bobbing for intra-group coordination.^[38]

Diet and foraging

The Javan rusa (Rusa timorensis), an opportunistic grazer-browser, primarily consumes grasses, which form the bulk of its diet, supplemented by leaves, young shoots, herbs, fruits, bark, and occasionally seaweed in coastal areas. In West Papua habitats, common forage includes gramineous species such as field grass, Imperata cylindrica, and elephant grass (Pennisetum purpureum), underscoring the deer's reliance on available herbaceous vegetation. Introduced populations in New Caledonia show native plants comprising 40-60% of the diet, with grasses and forbs dominating alongside browse.^[43]^[44]^[15] Foraging occurs mainly at night in open grassy areas through grazing, with browsing in shrublands during safer daytime periods, adapting to habitat structure and predation risk. The deer derives most of its moisture from vegetation, exhibiting low free-water requirements that enable survival in arid or seasonal environments with minimal standing water; experimental studies confirm intake ratios as low as 3.3 liters of water per kilogram of dry matter consumed on forage-based diets.^[15]^[45] Dietary composition shifts seasonally in response to forage availability and quality, with grasses like Cynodon dactylon comprising up to 80% during wet periods for their palatability and abundance, while dry seasons prompt increased browsing on shrubs and dicots. Invasive plants, such as certain acacias in altered habitats, can modify these patterns by providing alternative but less nutritious options, potentially affecting nutritional intake.^[46]^[47] As a ruminant, the Javan rusa features a four-chambered stomach (rumen, reticulum, omasum, abomasum) where microbial fermentation breaks down fibrous plant material into volatile fatty acids for energy, supporting its mixed diet of tough forages. This system enables high digestibility of cellulose-rich grasses, with seasonal variations in intake and nutrient retention observed in tropical conditions. Selective feeding behaviors help avoid unpalatable or toxic plants, prioritizing nutrient-dense options.^[48]^[49]^[50]

Reproduction and life cycle

The Javan rusa (Rusa timorensis) employs a polyestrous breeding system, with mating occurring year-round but peaking between July and September in its native range.^[1] Males compete aggressively for access to females through vocalizations, displays, and physical clashes using their antlers.^[51] Gestation lasts approximately 8 months (around 240–253 days), after which a female typically gives birth to one calf, though twins occur rarely.^[1]^[52] Births take place throughout the year, but exhibit seasonality aligned with the breeding peak in native habitats. Newborn fawns are precocial, able to stand shortly after birth, and remain hidden in vegetation for the first few weeks while the mother forages nearby and returns periodically to nurse.^[4]^[51] Calves are weaned at 6–8 months of age, after which they begin to accompany the herd more actively.^[1] Sexual maturity is attained at 18–24 months, depending on nutrition and habitat conditions; full physical growth, including antler development in males, is reached by about 5 years.^[53]^[52]^[1] The life cycle includes a juvenile phase characterized by rapid growth and learning social behaviors, transitioning to an adult reproductive phase that dominates until senescence around 15 years of age; individuals may live up to 20 years in the wild or captivity.^[4]

Predators

The Javan rusa faces predation from several native carnivores across its range in Indonesia and Timor-Leste. The Javan leopard (Panthera pardus melas) is the primary predator, employing ambush tactics to target adults in forested habitats, where it relies on stealth and short bursts of speed to close in on unsuspecting deer.^[54] Packs of dholes (Cuon alpinus) hunt cooperatively in groups of three or more, using coordinated chases to exhaust and overwhelm prey, with a noted instance of three to four individuals killing an adult Javan rusa.^[55] On the islands of Flores and Rinca, Komodo dragons (Varanus komodoensis) pose a threat primarily to young and injured deer, attacking solitarily by biting legs to immobilize victims before tearing into flesh, often leading to death from blood loss or infection over hours or days.^[56] Reticulated pythons (Malayopython reticulatus) act as opportunistic predators, constricting and consuming fawns or weakened juveniles in dense vegetation or near water.^[1] Historically, the Javan tiger (Panthera tigris sondaica) served as a major apex predator on Java, preying on adult Javan rusa through stalking and ambush similar to the leopard; this subspecies was declared extinct in 2008, with the last confirmed sightings in the 1980s.^[57] Predation impacts are most severe on fawns, which face elevated risks from dholes, pythons, and Komodo dragons due to their smaller size and limited mobility, with observations indicating avoidance of predator-adjacent areas by females with young.^[58] Adults experience lower predation rates overall but remain vulnerable during foraging in open areas or when isolated from groups. Javan rusa employ several anti-predator strategies to mitigate these threats, including forming herds for collective vigilance, which enhances early detection of approaching predators.^[54] When danger is detected, individuals emit loud honking alarm calls to alert the group, prompting flight responses.^[59] Additionally, the deer rely on their agility, capable of rapid sprints to evade pursuits, particularly from leopards or dholes.^[1]

Conservation

Population status

The Javan rusa (Rusa timorensis) is classified as Vulnerable on the IUCN Red List, a status first assigned in 2008 and confirmed in the latest assessment as of October 2025.^[38] The global population of mature individuals is estimated at fewer than 10,000, reflecting ongoing declines primarily in native ranges.^[38] In native habitats, populations continue to decline due to habitat fragmentation and other pressures. Current estimates indicate around 5,000 individuals on Java, 1,500 on Bali, and a stable population of about 2,000 on Timor.^[38] Introduced populations are stable or increasing in several regions outside the native range. For example, in Australia, rusa deer have established populations across New South Wales, Queensland, and other areas, though these do not contribute to the conservation of native stocks due to their invasive status and genetic separation.^[30] Population monitoring in Indonesia relies on methods such as camera traps for detecting presence and abundance in protected areas, alongside periodic park censuses using line transects and fecal sampling to track trends and genetic health.

Major threats

The Javan rusa (Rusa timorensis) is primarily threatened by extensive habitat loss driven by deforestation for agricultural expansion and palm oil plantations, which have reduced Java's forest cover to approximately 20% of its land area from historically higher levels.^[60]^[39] This conversion fragments grasslands and forests essential for the deer's foraging and shelter, with oil palm development particularly impacting lowland areas across Java and Bali.^[26] Illegal poaching remains a major risk, with hunters targeting the species for meat and antlers despite national bans under Indonesian law, leading to significant population reductions in accessible areas.^[61]^[3] In regions like Timor, poaching intensity is heightened by limited enforcement resources and socioeconomic pressures, exacerbating declines in less monitored habitats.^[26] These threats have contributed to an overall population decrease, classifying the Javan rusa as vulnerable on the IUCN Red List.^[39] Human-wildlife conflicts intensify the pressures, as crop raiding by herds prompts retaliatory killings by farmers protecting agricultural fields.^[54] Proximity to livestock also raises risks of disease transmission, with wild deer potentially contracting or spreading pathogens to domestic animals in shared grazing areas.^[62] Climate change compounds these issues through prolonged droughts that diminish forage quality and availability in grasslands, while rising sea levels threaten coastal marshes and lowlands where the deer seek refuge.^[63]^[64]

Protection efforts

The Javan rusa (Rusa timorensis) is protected under Indonesia's Law No. 5/1990 on the Conservation of Living Natural Resources and Their Ecosystems, which establishes the legal framework for safeguarding biodiversity and prohibits the exploitation of protected species without permits. In 2018, it was officially designated as a fully protected species under amendments to this law, banning hunting, capture, and trade in the wild to address ongoing threats such as habitat loss and poaching.^[54] Key conservation programs include reintroduction efforts within Ujung Kulon National Park on Java, where 16 individuals (three males and 13 females) were translocated from Peucang Island to Panaitan Island between 1978 and 1982 to bolster population viability and restore ecological balance in the park's grasslands.^[58] This initiative has contributed to the species' persistence in a UNESCO World Heritage site, supporting broader biodiversity goals alongside the endangered Javan rhino. In Bali, community-based conservation initiatives integrate local participation through ecotourism and habitat management in areas like West Bali National Park, where residents monitor populations and promote sustainable land use to reduce encroachment while generating economic benefits.^[63] In August 2025, Bali Zoo released 12 additional Javan rusa into the park, where the wild population was estimated at around 1,014 individuals as of 2023.^[65] The Javan rusa holds cultural significance in Indonesia, symbolizing national wildlife heritage and featured on the obverse of the 1988 500-rupiah banknote alongside the Garuda Pancasila emblem, which has raised public awareness of its conservation needs.^[13] In introduced ranges like Australia, where the species is managed as a feral population, sustainable hunting programs emphasize ethical practices, regulated quotas, and habitat stewardship to control numbers while funding conservation through licensing fees.^[66] International efforts involve the IUCN Species Survival Commission (SSC), which coordinates assessments and action plans through its specialist groups, including monitoring the species' vulnerable status and recommending habitat connectivity measures. Transboundary protection plans for populations on Timor, spanning Indonesia and Timor-Leste, promote joint monitoring and anti-poaching patrols to safeguard shared habitats amid border challenges.

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Dec 27, 2017 · While R. unicolor is widespread throughout South and Southeast Asia (from India and Sri Lanka, Southern China and most of Indochina to Borneo ...
[28]
Geographical analysis of the Javan deer distribution in Indonesia ...
Feb 1, 2020 · Javan deer (Cervus timorensis) is a native animal of Indonesia whose range extends from Java, Bali and to the islands east of the Wallace ...
[29]
Yang Plateau: The last bastion of endangered Javan deer - jawawa
Apr 25, 2000 · ... range with the highest peak being Mount Argopuro (3088m), lies between Mt ... Ledeboer's great efforts to protect Javan deer in Yang Plateau ...
[30]
Cervus timorensis russa - GISD
May 17, 2007 · Rusa deer also represent an important resource for local people (hunting, farming). view this species on IUCN Red List. Pathway. Cervus ...Missing: facts | Show results with:facts
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Javan Rusa Deer - Heritage Bel Ombre |
Rusa deer originate from the islands of Java and Bali in Indonesia. They were introduced to Mauritius by Adrian Van der Stel, a Dutch governor who came to ...Missing: etymology | Show results with:etymology<|control11|><|separator|>
[32]
Feral rusa deer | Business Queensland
Aug 18, 2023 · Medium-sized deer, with Javan rusa stags standing up to 110cm at shoulder and weighing about 120kg, hinds standing up to 95cm at shoulder and ...
[33]
[PDF] Feral rusa deer - Queensland Government publications
Stags weigh about 120 kg, hinds up to 80 kg. Stags have three tined antlers with the beams forming a characteristic lyre shape. Life cycle. During the ...
[34]
[PDF] Population genetic structure of wild and farmed rusa deer (Cervus ...
Aug 14, 2009 · Abstract Historical records indicate that 12 rusa deer. (Cervus timorensis russa) were introduced in New-. Caledonia during the 1870s.
[35]
[PDF] Population Regulation of Rusa deer in New Caledonia
Rusa DEER ORIGIN & IMPACTS. Rusa timorensis russa (de Blainville, 1822). INTRODUCED IN 1870 (from Indonesia). Biological factors. Rapid acclimation. Food plant ...<|control11|><|separator|>
[36]
RUSA DEER (Cervus timorensis) AS A HOST FOR THE CATTLE ...
The rusa deer. (Ceri-tis timoretisis) was first introduced into. Papua. New. Guinea about. 1900. Today, sizeable populations occur in the. Port. Moresby area of ...
[37]
[PDF] Home ranges of rusa deer (Cervus timorensis) in a subtropical peri ...
Jul 11, 2022 · Rusa deer (Cervus timorensis) are one of those six species and have established numerous populations along the eastern seaboard of Australia ( ...
[38]
[PDF] Landscape structure does not hinder the dispersal of an invasive ...
Jan 24, 2024 · Javan rusa deer (Rusa timorensis) were introduced to New Caledonia in 1870 and the population consists of several hundred thousand individuals.
[39]
Low levels of genetic variation within introduced Javan rusa deer ...
Javan rusa deer (Cervus timorensis russa) is one of six introduced wild deer species in Australia. The largest population is located in the Royal National ...
[40]
IUCN Red List of Threatened Species
Current population trend · Number of mature individuals · Population severely fragmented · Continuing decline of mature individuals.
[41]
Geographical analysis of the Javan deer distribution in Indonesia ...
Javan deer was classified as Vulnerable in the IUCN Red List owing to its native wild population which are patchily distributed in a small geographical range ( ...Missing: facts | Show results with:facts
[42]
[PDF] The Time Budget of Javan Deer (Rusa timorensis, Blainville 1822) in ...
The activities pattern of Javan deer were divided into six categories: feeding, moving, resting, grooming, anti predator, and others. Feeding includes ...
[43]
Temporal and spatial dynamics of insular Rusa deer and wild pig ...
Aug 25, 2016 · A highly seasonal wet–dry cycle is dominant in the region, and there is considerable annual rainfall variation during the short summer monsoon ...
[44]
XC960845 Javan Deer (Rusa timorensis) - Xeno-Canto
Jan 13, 2025 · Javan Deer Rusa timorensis · alarm call. Simon Elliott. Remarks from the Recordist. alarm barks to recordist. Location. Rating. Rate the quality ...
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(PDF) Forage Food of Timor Deer (Cervus timorensis) in Manokwari ...
Aug 9, 2025 · The results indicated five most common forage species consumed in the study; they were field grass, Imperata (Imperata cylindrica), elephant ...
[46]
(PDF) The diets of introduced rusa deer (Cervus timorensis russa) in ...
Aug 6, 2025 · Seasonal variations in the proportions of plant types in the rumen (% dry weight) of rusa deer from Mépouiri (hot wet season: Mar–May 2001 ...Missing: social | Show results with:social
[47]
Water consumption by rusa deer (Cervus timorensis) stags as ...
Mar 9, 2007 · Stags ate 2·04 kg dry matter (DM) per day and drank 6·4 kg water per day, while the drinking water: food DM ratio was 3·3 l/kg. In experiment 2, ...
[48]
https://www.atlantis-press.com/article/126009996.pdf
[49]
Seasonal changes in the dietary compositions of rusa deer in ...
Diets shifted seasonally, possibly in response to availability and palatability. ... We observed an interspecies association between wild Javan lutungs ( ...
[50]
[PDF] Nutrient Digestibilities in Rusa Timor (Cervus timorensis de ...
Ruminants, for example, in Timor deer, would excrete energies which are digested as fermentation heat and gases such as CH4; in addition, not all things exist-.
[51]
Consumption and Digestion of Forages by Male Rusa (Cervus ...
Aug 5, 2025 · The tropical rusa deer exhibits seasonal variations in feed intake, food constituent digestibilities, VFA profile and N retention. Castration ...<|separator|>
[52]
[PDF] SHRUB PALATABILITY TO RUSA DEER (CERVUS ... - UKnowledge
The animal model was javan rusa deer (Cervus timorensis russa) because of its browsing behaviour and its commercial importance in New Caledonia (Le Bel,. 1993).Missing: habitat | Show results with:habitat
[53]
(PDF) Reproductive Behaviour Of Timor Deer (Rusa Timorensis)
Aug 5, 2025 · Aggressive behavior commonly leads mating. Parturition and maternal behavior of hinds are affected by limitation of space, therefore dividing ...
[54]
https://besjournals.onlinelibrary.wiley.com/doi/full/10.1002/2688-8319.12118
[55]
[PDF] Growth and Reproductive Performances of Farmed Timorensis Deer ...
There seemed to be a clear breeding season for timorensis deer in this study, which was within the moderate months of March to. April (average rainfall 236 mm) ...
[56]
The role of the introduced rusa deer Cervus timorensis for wildlife ...
Jan 7, 2022 · The rusa deer has been introduced to Merauke region and later to Vogelkop Peninsula in Indonesian New Guinea (West Papua) in 1928.
[57]
Exploring temporal activity of dholes, their prey, and competitors in ...
Jul 4, 2024 · Baluran NP is also one of only three protected areas on Java ... Observations on ajag (Cuon alpinus) hunting in Baluran and alas Purwo, east Java.
[58]
Knee deep in trouble: Rusa deer use an aquatic escape behaviour ...
Aug 6, 2025 · We document six observations of an aquatic behaviour used by rusa deer (Rusa timorensis) to delay an imminent attack from Komodo dragons ...
[59]
Is the Javan tiger Panthera tigris sondaica extant? DNA analysis of a ...
Mar 21, 2024 · The Javan and Bali tigers were categorized as Extinct on the IUCN Red List in both 2003 and 2008, leaving only the Sumatran subspecies extant.
[60]
Home range and habitat use of reintroduced Javan Deer in Panaitan ...
Sep 30, 2015 · The elevation range of Panaitan Island is also 0–320 m, but all age class of Javan deer in this island only used the elevation range between 0 m ...
[61]
Javan Deer (Rusa timorensis) :: xeno-canto
Javan Deer Rusa timorensis, 1:14, Simon Elliott, 2013-10-22, 16:00, Indonesia, Bali, Kabupaten Buleleng, Bali, 0, alarm call. alarm barks to recordist. [sono] ...Missing: anti- predator herding
[62]
Refining National Forest Cover Data Based on Fusion Optical ...
Aug 17, 2023 · The Java region had the lowest forest cover areas, with a total percentage of about 20% of the total area. The national forest extents for each ...
[63]
For Indonesia's Javan deer, non-protected areas play key ...
Feb 20, 2020 · A significant portion of the total population of Indonesia's Javan deer occurs outside protected areas, highlighting the role of such zones as wildlife refuges.
[64]
Serosurveillance and Molecular Investigation of Wild Deer in ...
Jul 13, 2020 · Therefore, the role wild deer might play in the spread of diseases to livestock remains unclear.
[65]
Effects of Javan rusa deer (Cervus timorensis) on native plant ...
Aug 5, 2025 · Javan deer was classified as Vulnerable ... They used preferentially the sclerophyll forest and flood plains, especially during the dry season.Missing: migration | Show results with:migration<|separator|>
[66]
Conference of the Parties to the Convention on International Trade ...
Dec 26, 2024 · The twentieth regular meeting of the Conference of the Parties to CITES (CoP20) is scheduled to be held in Uzbekistan, November 24-December 5, 2025.
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Rusa Deer Hunting Outfitters & Guides In Queensland Australia
... Javan rusa deer. ... Ethical Hunting Practices: High Country Safaris is committed to sustainable hunting, ensuring respect for wildlife and the environment.
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https://www.conservation.org/docs/default-source/gef-documents/timor-leste-wetlands-prodoc-final_25-06-25_clean.pdf