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Linnaea

Linnaea is a monotypic genus of flowering plants in the family Caprifoliaceae, comprising the single species Linnaea borealis, commonly known as twinflower.^[1]^[2] This evergreen subshrub features slender, trailing stems up to 1 m (3 ft) long, with opposite, ovate to orbicular leaves that are 5–16 mm long and toothed along the edges.^[3]^[4] The plant produces paired, nodding, bell-shaped flowers that are typically pink (occasionally white), fragrant, and 10–15 mm long, borne on erect stalks rising 1–6 cm above the foliage; these bloom from May to August.^[3]^[2] Fruits are small, dry, nut-like capsules covered in glandular hairs.^[3] Native to the subarctic and temperate regions of the Northern Hemisphere, Linnaea borealis has a circumboreal distribution extending from Eurasia and North America (including Alaska, Canada, and northern United States) to high-elevation sites in Uganda's Ruwenzori Mountains.^[5] It thrives in cool, moist, acidic, humus-rich soils within coniferous or mixed forests, alpine/subalpine zones, talus slopes, and peatlands, often associating with mosses and shade-tolerant vegetation, though it requires good drainage and cannot tolerate drought.^[3]^[4] The species is perennial and cannot self-pollinate, relying on cross-pollination from unrelated individuals for seed production.^[3] The genus name Linnaea honors the Swedish botanist Carl Linnaeus (1707–1778), known as the father of modern taxonomy, who reportedly considered L. borealis his favorite plant; it was proposed by his friend and collaborator, Dutch botanist Jan Frederik Gronovius, in 1753.^[4] The specific epithet borealis derives from Latin, meaning "northern," reflecting its predominantly boreal habitat.^[4] Three subspecies are recognized—ssp. americana in eastern North America, ssp. borealis in Eurasia, and ssp. longiflora in western North America—though the genus remains monotypic overall.^[5]^[4]

Taxonomy and etymology

Taxonomic classification

Linnaea is a monotypic genus within the subfamily Linnaeoideae of the family Caprifoliaceae, order Dipsacales, class Magnoliopsida, phylum Tracheophyta, and kingdom Plantae.^[6]^[5] The sole species, Linnaea borealis L., represents the entire genus, which is characterized by its placement in the core Eudicots based on molecular and morphological evidence.^[7] Historically, the genus was classified in its own family, Linnaeaceae, in older taxonomic systems due to its distinct morphology, such as paired flowers and trailing habit.^[8] Modern molecular phylogenetics, including analyses of nuclear ribosomal ITS and chloroplast DNA sequences, have integrated Linnaea into the expanded Caprifoliaceae sensu lato, confirming its monophyly within the subfamily Linnaeoideae alongside genera like Abelia, Diabelia, Dipelta, Kolkwitzia, and Vesalea.^[9]^[10] This reclassification, supported by studies from 2013 onward, reflects the close evolutionary relationships revealed through Bayesian inference and maximum likelihood methods, diverging from eastern Asian lineages around 50.86 million years ago.^[9] The genus Linnaea has no direct synonyms, though it is sometimes confused with related genera in Linnaeoideae, such as Abelia and Zabelia (now often treated as Diabelia), due to overlapping floral traits like tubular corollas.^[10] For the species L. borealis, synonyms include Linnaea americana Forbes, which pertains specifically to North American variants.^[7]^[3] Three subspecies of Linnaea borealis are widely recognized: L. borealis subsp. borealis, native to Europe and parts of Asia; subsp. americana (Forbes) Hultén, occurring in eastern North America; and subsp. longiflora (Torr.) Hultén, found in western North America and Asia.^[7] These subspecies differ primarily in corolla length and geographic isolation, with subsp. longiflora exhibiting longer flowers adapted to Pacific coastal regions.^[7] Some authorities recognize only two subspecies (borealis and longiflora), lumping subsp. americana with the latter, though three are widely accepted based on morphological differences.^[7]

Etymology and history

The genus Linnaea derives its name from the Swedish botanist Carl Linnaeus (1707–1778), reflecting his deep personal affection for the plant, which he affectionately termed planta nostra ("our plant") due to its significance in his early explorations in Lapland. The name was proposed around 1737 by Linnaeus's colleague, the Dutch botanist Jan Frederik Gronovius (1690–1762), who suggested it as a tribute; Linnaeus first used it in print in his Critica Botanica (1737), and formally described the species as Linnaea borealis in Species Plantarum (1753), marking it as the type species and initially treating the genus as monotypic, with L. borealis as its sole representative. Gronovius selected it specifically because Linnaea borealis was Linnaeus's favorite species, symbolizing the twin flowers that mirrored the collaborative spirit of scientific discovery.^[11]^[12] Linnaeus initially expressed reluctance to accept the honor, viewing the modest, unassuming nature of the plant as akin to his own humble origins, but he ultimately embraced it with characteristic humility, as noted in his Critica Botanica (1737), where he described Linnaea as "a plant of Lapland, lowly, insignificant, disregarded, flowering but for a brief space—after Linnaeus." This personal connection underscored Linnaeus's philosophy of naming, prioritizing plants that evoked simplicity and northern resilience over grandeur.^[13]^[12] Subsequent taxonomic history saw Linnaea integrated into the family Caprifoliaceae by Antoine Laurent de Jussieu in his Genera Plantarum (1789), where it was classified among honeysuckle relatives based on floral and fruit characteristics, solidifying its place in early natural classification systems. Throughout the 19th and 20th centuries, the genus retained its monotypic status centered on L. borealis, though debates arose regarding potential mergers with closely related genera such as Abelia, particularly as morphological similarities prompted discussions on whether to expand Linnaea to encompass broader clades; these considerations highlighted ongoing refinements in understanding its evolutionary affinities without altering its historical core.

Description and biology

Physical characteristics

Linnaea is a trailing evergreen subshrub that forms loose mats through slender, pubescent stolons and stems typically measuring 20–40 cm in length, with the stems becoming semi-woody and up to 0.3 cm in diameter with age. The plant rarely exceeds 10 cm in height, featuring creeping horizontal growth that allows it to spread over the ground surface via rooting at nodes. These stems are often glandular-hairy, contributing to the plant's adaptation for vegetative propagation in shaded forest floors.^[7]^[14] The leaves are opposite, simple, and evergreen, persisting for about two years, with blades that are ovate to elliptic or orbicular, measuring 5–16 mm long and 4–10 mm wide, and featuring shallowly toothed margins along the upper half. They have a leathery texture, are glabrous to sparsely hairy on the upper surface, and are supported by short petioles of 2–3 mm. This arrangement gives the foliage a glossy, bright green appearance that remains year-round.^[3]^[7] Flowers occur in pairs on arching, terminal inflorescences with peduncles 45–80 mm long, presenting as pendulous, bell-shaped blooms that are pale pink to white and tubular, 8–15 mm long, formed by five fused petals with a narrow basal tube flaring into rounded lobes. The corolla is often hairy inside, with four exserted stamens and a style protruding beyond the tube; blooming typically spans June to August in northern latitudes. Fruits develop as small, dry, one-seeded capsules, approximately 3 mm long, that are unequally three-celled and hairy with glandular tips, containing tiny seeds dispersed passively.^[3]^[15]^[7] Subspecies exhibit subtle morphological variations, particularly in corolla tube length relative to the limb; for instance, L. borealis subsp. longiflora has longer tubes (10–16 mm overall) that equal or exceed the limb width, distinguishing it from the shorter-tubed subsp. borealis and subsp. americana. These differences are most evident in western North American populations.^[14]^[16]

Reproduction and life cycle

Linnaea borealis, a perennial evergreen subshrub, exhibits a life cycle characterized by both sexual and vegetative reproduction, with the latter dominating population persistence in natural habitats. As a trailing plant, it overwinters through its persistent green leaves, which remain viable for 12–14 months, allowing continuous photosynthesis and survival in boreal and temperate environments.^[17]^[18] The species typically reaches reproductive maturity slowly, with seedlings requiring approximately 13 years to produce their first flowers, contributing to its long-lived clonal nature.^[19] Flowering occurs during the summer months, primarily from early June to mid-August in northern ranges, aligning with peak pollinator activity in forested and subalpine settings.^[18] Each pair of nodding, bell-shaped flowers lasts about one week, releasing a fragrant scent reminiscent of lilac or almond to attract insects at dusk.^[20]^[19] Pollination is entomophilous and strictly cross-pollination dependent due to self-incompatibility, which prevents inbreeding and ensures genetic diversity; primary pollinators include bumblebees (Bombus spp.) and small flies such as syrphids (Syrphidae), though isolated populations often experience low pollination success leading to reduced seed set of 24–60%.^[17]^[3]^[18] Following pollination, fertilized flowers develop into small, dry, nut-like fruits containing a single seed each, which mature over approximately 36 days from late July to September.^[20]^[19] Seed production is generally low, with viability around 50% under optimal conditions and seedlings rarely observed in the wild due to poor establishment; the non-dehiscent fruits feature sticky surfaces that facilitate limited zoochorous dispersal by adhering to animals or substrates, though no specialized long-distance mechanisms exist.^[19]^[17]^[3] Vegetative reproduction via stolons is the primary mode of spread, enabling the formation of extensive clonal colonies that expand up to 30–48 cm annually beneath forest litter.^[19]^[20] These horizontal stems produce new ramets (rejuvenating shoots) at low rates, with only 1–2% of ramets flowering each year, supporting long-term population stability over sexual recruitment.^[17]^[18] In cultivation, Linnaea borealis presents challenges for ex situ conservation due to its slow growth and low seed viability, with only about 33% survival after five years in outplantings.^[19] Vegetative propagation via stem cuttings (3–5 cm long) or layering succeeds in perlite-sand media under mist and bottom heat (21°C), rooting in 6–8 weeks, though full establishment takes 5–10 years.^[19] Recent studies on subsp. borealis have advanced in vitro techniques using Murashige and Skoog medium supplemented with cytokinins (e.g., BAP 1.0 mg/L), auxins (e.g., IAA 0.1 mg/L), and gibberellins (GA₃ 1.0 mg/L) for shoot multiplication, achieving up to 50 new shoots per explant in liquid cultures and 100% rooting with IBA-IAA combinations; these methods support biomass production for rare populations while minimizing habitat disturbance.^[21] Seed propagation requires 60-day cold stratification but yields low germination (1:30 ratio), underscoring the preference for clonal approaches in conservation efforts.^[19]

Distribution and habitat

Geographic range

Linnaea borealis exhibits a circumboreal distribution across the Northern Hemisphere, primarily in subarctic to temperate zones, spanning Europe, Asia, and North America.^[5] This range reflects its adaptation to cooler climates, with populations often concentrated in boreal forests and higher elevations where conditions remain suitable.^[21] In Europe, the species is widespread in northern regions, including Scandinavia (Norway, Sweden, Finland), Scotland, and the Baltic states, extending southward into the Alps (Austria, Switzerland, France, Italy, Germany) and the mountains of Poland, the Czech Republic, and Slovakia.^[5] It is notably absent from much of central Europe, where warmer post-glacial conditions have led to extirpations, confining it to relic populations in mountainous or northern refugia.^[21] The Asian range encompasses Siberia (including Krasnoyarsk, Irkutsk, Yakutiya, and Chita), the Russian Far East (Kamchatka, Khabarovsk, Primorye, Sakhalin, and Kuril Islands), Japan, Korea, Mongolia, northern China (Manchuria, Inner Mongolia, Xinjiang), Kazakhstan, and the Caucasus (North Caucasus and Transcaucasus).^[5] Here, L. borealis subsp. borealis predominates, forming extensive stands in subarctic woodlands.^[7] In North America, the plant occurs from Alaska southward to northern California on the west coast, across Canada to Newfoundland, and into the eastern and central United States, reaching the Appalachians, northern Arizona, New Mexico, and as far south as West Virginia and New Jersey.^[7] Subspecies distribution includes L. borealis subsp. americana in the eastern U.S. and Canada (from interior Alaska to the Atlantic coast), and subsp. longiflora along the Pacific Northwest from southern Alaska to northern California and the Rocky Mountains.^[7] Subsp. borealis appears in eastern Alaska.^[7] Historically, L. borealis expanded post-glacially from Late Glacial refugia, recolonizing northern latitudes as ice sheets retreated, but southern populations have since declined due to climatic warming, resulting in fragmented distributions at range edges like central Europe and the southern Appalachians.^[21]^[22] A disjunct population occurs at high elevations in the Ruwenzori Mountains of Uganda.^[5]

Preferred habitats

Linnaea borealis thrives in cool, moist climates characteristic of boreal and subarctic regions, where temperatures remain moderate and humidity is consistently high, tolerating partial to full shade but avoiding direct sunlight and dry conditions.^[7] It prefers acidic soils with a pH range of approximately 4.5 to 6.0, which support its growth in environments rich in organic matter and humus.^[23] Well-drained yet moist soils are essential, preventing waterlogging while maintaining adequate moisture levels, and the plant is commonly found in terrains such as forests, bogs, and talus slopes.^[24] In associated ecosystems, Linnaea borealis is a prominent understory species in coniferous forests of the taiga, along tundra edges, and in montane woodlands, often growing alongside mosses, ferns, and other shade-tolerant vegetation.^[4] It favors microhabitats that provide cool, stable conditions, such as algific talus slopes in North America, where north- or northeast-facing bluffs create perpetually moist and chilled environments due to cold air drainage and ice persistence.^[25] Elevations up to 3,000 meters are suitable in mountainous areas, allowing it to occupy higher-altitude woodlands and rocky slopes.^[7] The trailing growth habit of Linnaea borealis enables it to form extensive ground cover in shaded, humid forest floors, effectively colonizing and stabilizing organic-rich substrates while competing minimally for light.^[26] This adaptation suits its preference for undisturbed, moist understories where it can spread via rhizomes in well-aerated, acidic soils.^[27]

Ecology and conservation

Ecological interactions

Linnaea borealis engages in pollination ecology primarily through entomophily, with key pollinators including bumblebees (Bombus spp.), hoverflies (Syrphidae), muscid flies, and empid flies, which are attracted to the nectar rewards in its small, pendulous flowers. These insects, often small-bodied, facilitate cross-pollination in this self-incompatible species, though fragmented populations may experience reduced reproductive success due to limited pollinator movement.^[28]^[29] Seed dispersal in L. borealis occurs mainly via epizoochory, where nutlets adhere to the fur of mammals or feathers of birds, enabling transport over short distances. Vegetative propagation through stolons dominates its spread, forming extensive clonal patches that can persist for centuries and restrict gene flow by promoting intraclonal reproduction over sexual recombination. While wind may contribute minimally to dispersal, myrmecochory has not been documented.^[7]^[28] The species forms symbiotic arbuscular mycorrhizal associations with soil fungi. In food webs, its evergreen foliage is browsed by herbivores such as caribou and bighorn sheep, while seeds provide sustenance for birds like ruffed grouse; flowers provide nectar for insects.^[30]^[7] As a mat-forming perennial, L. borealis delivers ecosystem services by acting as ground cover that mitigates soil erosion on forested slopes and contributes to understory stability. It serves as an indicator species for old-growth coniferous forests, where its presence signals mature, undisturbed habitats with intact microclimates and biodiversity.^[7]^[19]

Conservation status and threats

Linnaea borealis is not formally assessed on the IUCN Red List globally but is ranked as G5 (globally secure) by NatureServe in North America, indicating no immediate threats to its overall persistence.^[31] However, vulnerability increases at its southern distributional edges, where populations face heightened risks from environmental changes. In Switzerland, it is assessed as Least Concern (LC) nationally.^[32] Regionally, the species is nationally scarce in the United Kingdom, confined primarily to fragmented pinewood habitats in the Scottish Highlands, with ongoing declines noted since the 1930s.^[33] In the southern United States, it holds S1 status (critically imperiled) in states like West Virginia due to limited occurrences in specialized microhabitats such as cool talus slopes.^[31] It is considered extirpated in portions of central Europe, including some lowland areas of Poland, where it persists only as a glacial relict in isolated refugia.^[34] Primary threats include climate change, which is projected to shift suitable boreal habitats northward through warming and drying conditions, potentially displacing southern populations.^[26] Habitat loss from logging, development, and altered woodland management has fragmented remaining stands, particularly in Europe and southern North America.^[35] Competition from invasive species, such as understory invasives in forest edges, poses additional risks in select locations, though monitoring is emphasized over direct impacts.^[18] Population trends remain stable in core boreal regions but show declines at range margins, driven by habitat fragmentation and reduced reproductive success.^[36] Isolated populations often exhibit genetic bottlenecks, with low within-population diversity and high differentiation between sites, limiting long-term viability due to the species' self-incompatibility.^[37] As a characteristic understory species of boreal forests, Linnaea borealis serves as an indicator for ecosystem health, with monitoring programs tracking its response to threats like fragmentation.^[38]

Cultural significance

Association with Carl Linnaeus

Carl Linnaeus first encountered Linnaea borealis during his botanical expedition to Lapland in 1732, where the plant's delicate, paired flowers growing amid the northern landscape left a lasting impression on him.^[39] This experience shaped his appreciation for the species, which he later described as embodying simplicity and understated beauty, reflecting his ideals in botanical observation.^[13] In his 1737 publication Flora Lapponica, Linnaeus declared Linnaea borealis his favorite flower, praising its humble form as a symbol of northern heritage and personal modesty amid the rugged terrains of his ancestral region.^[4] The plant's modest stature and brief bloom period resonated with Linnaeus's own self-perception, as he noted its unassuming nature in contrast to more showy species.^[13] The genus Linnaea was named in Linnaeus's honor by his mentor and collaborator, Dutch botanist Jan Frederik Gronovius, who proposed it in 1737 as a tribute to Linnaeus's contributions to botany.^[12] Initially, Linnaeus demurred, adhering to his early principle against eponymous naming in taxonomy, but he accepted the gesture and incorporated it into his works, using it in Critica Botanica (1737) to advocate for such commemorative names while quoting: "Linnaea was named by the celebrated Gronovius and is a plant of Lapland, lowly, insignificant, disregarded, flowering but for a brief space—after Linnaeus who resembles it."^[13] This association extended to Linnaeus's later publications, emphasizing the value of personal tributes in scientific naming to honor key figures in the field.^[40]

Symbolism and uses

Linnaea borealis serves as the provincial flower of Småland in Sweden, Linnaeus's home region, symbolizing the delicate beauty of northern flora and evoking themes of humility and resilience in boreal landscapes.^[41] This association underscores its representation of northern wilderness, where its paired, nodding pink flowers mirror the subtle elegance of subarctic ecosystems.^[4] It was depicted on older Swedish banknotes, such as the 50 krona note (1965–1990) showing Linnaeus holding the flower, underscoring its cultural importance.^[42] In horticulture, Linnaea borealis is valued as an ornamental groundcover for woodland gardens, rock gardens, and shaded borders, forming mats up to 3 feet wide with its trailing stems and evergreen foliage.^[24] It thrives in moderately fertile, humus-rich, acidic soils that remain reliably moist, preferring partial shade to mimic its natural forest understory, with blooms appearing from June to September.^[24] Propagation occurs via stem cuttings or division of young runners, though transplanting can be challenging due to its shallow roots and dependence on mycorrhizal fungi for nutrient uptake, often requiring careful soil handling to preserve these associations.^[43]^[29] Traditional medicinal uses of Linnaea borealis include preparations for skin conditions and respiratory ailments, with Norwegian folk medicine employing infusions or poultices from the whole plant to treat shingles, eczema, and rashes.^[44] Indigenous groups in North America, such as the Algonquin and Tanana, used teas from the leaves or mashed plant material for colds, coughs, headaches, and as a tonic during pregnancy.^[45]^[24] Modern ethnobotanical interest remains limited, focusing on its historical role rather than widespread contemporary applications. The plant appears in 18th-century botanical illustrations, such as those in Linnaeus's own works and contemporary floras, highlighting its aesthetic and scientific appeal through detailed engravings of its paired flowers.^[13] It has inspired literary references in poetry celebrating northern nature, including nods to its fragile charm in Romantic-era writings that evoke woodland serenity.^[46] As a circumboreal species, Linnaea borealis symbolizes vulnerability to climate change in conservation efforts, representing boreal plants at risk from habitat shifts and warming temperatures, with projects in the UK and Scandinavia using it to raise awareness about protecting ancient woodlands.^[38]^[35] Studies project potential range losses of 35-52% by 2070 under moderate emissions scenarios, emphasizing its role in campaigns for ecosystem preservation.^[22]

References

[1]
Linnaea Gronov. ex L. | Plants of the World Online | Kew Science
Linnaea Gronov. ex L. ... The native range of this genus is Subarctic & Temp. Northern Hemisphere, Uganda (Ruwenzori Mountains).
[2]
USDA Plants Database
**Summary of Linnaea borealis (USDA Plants Database):**
[3]
Linnaea borealis (American twinflower) - Go Botany
Habitat. Alpine or subalpine zones, forests, talus and rocky slopes ... Flower description: the flower has an inferior ovary, with or without a ...
[4]
Twinflower - USDA Forest Service
Linnaea borealis was reported to be Linnaeus favorite plant, and was named by his close friend and teacher Jan Frederik Gronovious in honor of Linnaeaus.Missing: etymology | Show results with:etymology
[5]
Linnaea borealis L. | Plants of the World Online | Kew Science
The native range of this species is Subarctic & Subalpine Eurasia to Alaska, Uganda (Ruwenzori Mountains). It is a subshrub and grows primarily in the temperate ...
[6]
Taxonomy browser (Linnaea borealis) - NCBI
Lineage (full): cellular organisms; Eukaryota; Viridiplantae; Streptophyta; Streptophytina; Embryophyta; Tracheophyta; Euphyllophyta; Spermatophyta; ...
[7]
Linnaea borealis - USDA Forest Service
The currently accepted scientific name of twinflower is Linnaea borealis L. [28,33,55,62,62]. It is a monotypic genus [28].
[8]
Linnaea borealis L. var. borealis—In Vitro Cultures and ... - MDPI
This taxon was formerly assigned to the Caprifoliaceae family; however, it was transferred to the family of Linnaeaceae [1,2]. A more recent classification has ...Missing: phylogenetics | Show results with:phylogenetics<|separator|>
[9]
Molecular Phylogeny and Biogeographic Diversification of ...
Mar 10, 2015 · In this study, we conducted phylogenetic and biogeographic analyses for Linnaeoideae and its close relatives based on sequences of the nuclear ...
[10]
Twins are not alone: a recircumscription of Linnaea (Caprifoliaceae)
Aug 21, 2013 · The genus Linnaea is reviewed and expanded to include the genera Abelia (excluding section Zabelia), Diabelia, Dipelta, Kolkwitzia and Vesalea, making it ...
[11]
Linnaea borealis - Oxford University Plants 400
The name Linnaea was first proposed by the Dutch botanist, Jan Frederik Gronovius, to honour Carl von Linnaeus, the Swedish botanist who devised the modern ...Missing: etymology | Show results with:etymology
[12]
Floral scent chemistry within the genus Linnaea (Caprifoliaceae)
Nov 27, 2017 · Linnaeus was particularly fond of the species and early referred to it as 'planta nostra' (my plant), deciding that it should carry the Latin ...Missing: etymology | Show results with:etymology
[13]
[PDF] In Search Of Linnaea… - Gustavus Adolphus College
In Critica Botanica (1737), Linnaeus writes, “Linnaea was named by the celebrated Gronovius [Dutch botanist Johan Frederick Gronovius] and is a plant of ...Missing: reluctance | Show results with:reluctance
[14]
Twinflower - Linnaea borealis - Alaska Wildflowers - Lee Petersen
Jul 23, 2020 · It was Linnaeus that developed the standard Latin binomial species name, and he went on to name over 8000 species of plants and animals. ...<|separator|>
[15]
Twinflower - Montana Field Guide
Evergreen subshrub. Stems slender, trailing; short erect shoots 1–6 cm. Herbage glabrate to pubescent, glandular above. Leaf blades ovate to suborbicular, ...Missing: morphology | Show results with:morphology
[16]
Variation in Linnaea borealis (Linnaeaceae) across North America ...
Jul 24, 2017 · Linnaea borealis subsp. longiflora is represented in eastern and southeastern Alaska, south to California, and east to the Atlantic and ...
[17]
[PDF] Life History Strategies in Linnaea borealis - DiVA portal
Figure 3. Life cycle graph for Linnaea borealis. Broken lines indicate the formation of a new apex (i.e. fecundity), and solid arrows indicate the continued ...
[18]
American Twinflower | Mass.gov
Apr 30, 2025 · American twinflower is typically associated with cool, moist, boreal forest habitats such as spruce-fir forests and high-elevation communities.Missing: morphology | Show results with:morphology<|separator|>
[19]
[PDF] Plant Propagation Protocol for Linnaea borealis L. ESRM 412
Apr 25, 2009 · Flowers from June to Sept., and seeds mature in 36 days. Seed germination (needs dormancy breaking?) L. borealis does not set very much seed, ...Missing: cycle | Show results with:cycle
[20]
Twinflower - Cairngorms National Park
The bloom lasts seven days, after which, if it is pollinated, the flower develops a small, one-seeded fruit that ripens over 36 days.
[21]
Linnaea borealis L. var. borealis—In Vitro Cultures and ... - NIH
Nov 11, 2021 · Linnaea borealis L. (Twinflower), a creeping dwarf shrub, was Linnaeus's favorite plant and became his symbol. This taxon was formerly assigned ...
[22]
From past to future: impact of climate change on range shifts and ...
Aug 14, 2017 · We found that circumboreal plants could potentially lose their geographic ranges in the future (2070; 35–52% in RCP 4.5 (representative concentration pathways),
[23]
Linnaea borealis Twinflower
Very slow to establish; it takes seedlings about thirteen years to bloom. Vegetative reproduction by stolons is the primary method of regeneration. First ...Missing: cycle | Show results with:cycle
[24]
Linnaea borealis (Twinflower) | Native Plants of North America
Jul 16, 2025 · Soil Description: Cool, moist, acidic, humus-rich soils. Conditions Comments: Requires good drainage but will not tolerate drought. Benefit.Missing: morphology | Show results with:morphology
[25]
[PDF] CTs46 Southern Algific Talus factsheet - files
Southern Algific Talus. Cool, moist, open plant communities on steep northwest‑ to northeast‑facing bluffs in karst landscapes of southeastern Minnesota.
[26]
Linnaea borealis - Trees and Shrubs Online
Habitat Moist but well drained acid soils in a broad range of woodland communities, and occasionally in grasslands and scrub. Woodland habitats are often ...
[27]
https://pfaf.org/user/Plant.aspx?LatinName=Linnaea%20borealis
[28]
Limited mate availability decreases reproductive success of ...
Limited mate availability and isolation from pollen exchange compromise the reproductive success of fragmented populations of L. borealis in Scotland.
[29]
Linnaea borealis ssp. longiflora - Washington Native Plant Society
Description. A creeping evergreen herb with trailing and flowering stems found in open forests, forest edges, and peatlands.Missing: morphology | Show results with:morphology
[30]
[PDF] Scientific Name: Rosa acicularis Lindl
Form arbuscular mycorrhizal associations. (Kranabetter and MacKenzie 2010). ... CYSIP: Botany, n.d. Linnaea borealis: Twinflower. ... twinflower (Linnaea borealis L ...
[31]
Door to Nature: The Swedish Twinflower
Jun 18, 2021 · Its Latin name is Linnaea borealis, in honor of the great Swedish botanist Carolus Linnaeus (1707-78), who was more commonly known in the early ...Missing: etymology reluctance
[32]
Linnaea borealis L. - Infoflora
Oct 29, 2025 · Reaction Value R. 1: Very acid (pH 2.5-5.5); 2: acid (pH 3.5-6.5); 3: lightly acid to neutral (pH 4.5-7.5); 4: neutral to basic (pH 5.5-8.5); 5 ...
[33]
Linnaea borealis | NatureServe Explorer
Classification ; Phylum: Anthophyta ; Class: Dicotyledoneae ; Order: Dipsacales ; Family: Caprifoliaceae ; Genus: Linnaea.
[34]
Twinflower - Plantlife
The isolation of the remaining sites of Twinflower leads to poor seed production and thus contributes to its continued decline. Other threats include; ...
[35]
(PDF) The Twinflower (Linnaea borealis L.) in the northern part of ...
Aug 9, 2025 · Distribution of Linnaea borealis stations in the northern part of the Południowopodlaska Lowland mapped on ATPOL grid squares (10 x 10 km, ...
[36]
Estates in the National Park providing a boost for twinflower…
Mar 23, 2023 · Habitat loss and changes in the way that woodlands are managed are thought to have resulted in a reduction in twinflower since the 1930s, so ...
[37]
Linnaea borealis L. in BSBI Online Plant Atlas 2020
The 10 km square distribution of L. borealis is stable within its core range, centred on the Cairngorms National Park. Here, since 2000, there has been a 38% ...
[38]
Genetic Variability of Isolated Eastern Alpine Populations of Linnaea ...
Very low genetic variation within the two isolated Alpine populations and very high genetic variation between the two Alpine populations was found, despite ...
[39]
Aigas Twinflower Project Update
Apr 6, 2024 · The project aims to secure the Twinflower by planting insurance populations. 72% of cuttings survived, and more flowering plants are seen each ...
[40]
Entering a New World: The Northern Travels of Carl Linnaeus in 1732
Jul 11, 2017 · In his right hand Linnaeus holds his favorite northern flower, linnaea borealis (twinflower). The Dutch botanist Jan Frederik Gronovius named ...
[41]
The Linnaean collection | Natural History Museum
... Hortus Cliffortianus (1737) based on the plants in Clifford's garden. One of the Library's copies of Hortus Cliffortianus includes a pressed specimen of Linnaea ...
[42]
Cabinet 14 - Towards Species Plantarum - University of Otago
Carl Linnaeus, Hortus Cliffortianus. Amsterdam, 1737. Lehre [Germany]: J ... ' One of note was Linnaea borealis, the hardy twinflower which became a floral ...
[43]
[PDF] Dear Martlet - University of Iowa Libraries Publishing
is the emblematic flower of Smaland Province. On the Linn? coat of arms, the cartoon plant flows as a garland from his helmet's ruffed crest. In real life ...
[44]
Twinflower - Gardening at USask - College of Agriculture and ...
May 3, 2021 · Twinflower is very rarely cultivated. It is sometimes finicky, and it has very shallow roots that dry out quickly. It will not tolerate wind or drying out.
[45]
Ethnobotany of Linnaea borealis (Linnaeaceae) in Norway - 2006
Jul 7, 2006 · In folk medicine, Linnaea was also used to treat other kinds of rash, eczema, hives, measles, ringworms, scabies, water blisters, rheumatism ...Missing: remedies | Show results with:remedies
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Linnaea borealis - Twinflower - Wild About Flowers
In stockSome birds may forage for insects among its foliage, and while not a preferred browse, young shoots may occasionally be eaten. Thriving in cool, moist soils ...
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Ethnobotany of Linnaea borealis (Linnaeaceae) in Norway
Aug 7, 2025 · In Norwegian folk tradition, Linnaea borealis has a long-standing reputation as a cure for shingles (Herpes zoster).