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Pugmark

A pugmark is the footprint or paw print left by an animal, particularly large wild animals such as tigers (Panthera tigris) and other felids, serving as a primary sign in wildlife tracking to detect presence, movement, and individual identity. The term derives from the Hindi word "pug," meaning foot. In , particularly in , pugmarks have been central to population monitoring since the development of the pugmark census method in 1966 by forester S.R. Choudhury. Male pugmarks are typically larger and wider, while female pugmarks are more elongated and narrower, with overall size also indicating age categories like cubs, sub-adults, or adults; fresh prints, free of dust or debris, signal recent activity and help trackers predict direction and timing of movement. Despite its widespread use for decades across India's tiger reserves, the pugmark census faced significant limitations, including incomplete coverage, inconsistent print quality, and challenges in accurate by untrained personnel, leading to unreliable population estimates as critiqued in scientific analyses. Consequently, it has largely been supplanted by more precise methods like camera trapping and genetic analysis, though pugmarks remain a valuable supplementary tool for assessing occurrence and relative abundance, as used in recent es as of 2025.

Definition and Terminology

Definition

A pugmark is the paw print or footprint left by a wild , particularly carnivores such as tigers (Panthera tigris) and leopards (Panthera pardus), formed when the animal walks on soft substrates like , , or . These impressions serve as a primary tool for species identification, individual tracking, and monitoring animal presence in ecological studies. In wildlife contexts, pugmarks differ from general footprints by their specific application in conservation and forensic analysis, where they are examined to infer details about the animal's movement, habitat use, and population dynamics rather than mere casual observation. Unlike hoofmarks of herbivores, pugmarks feature a central pad with toes, enabling differentiation across species. Pugmark characteristics vary by but commonly include a rounded or oval central pad surrounded by four toes, with retractable claws that typically do not leave visible marks in felines due to sheathing during normal locomotion. For instance, pugmarks exhibit a tri-lobed pad with toes arranged in a semi-circle, while pugmarks are proportionally smaller with similar toe placement but distinct pad proportions relative to overall size. Pugmarks have been historically employed in population censuses to estimate numbers and .

Etymology and Usage

The term "pugmark" derives from "pug," an English borrowing from the Hindi word pag (पग), meaning "foot," "paw," or "step," combined with "mark" to denote the impression left by an animal's foot. This compound originated in 19th-century Anglo-Indian hunting terminology during British colonial rule in India, where English speakers adapted local words to describe animal tracks encountered in the field. The standalone term "pug" for an animal's footprint was first documented in English around 1860–65, reflecting its integration into colonial and tracking practices. By the early , "pugmark" as a specific compound had entered wider usage, with its earliest recorded appearance in 1922, and it has since become standardized in to refer precisely to paw prints suitable for identification. In regional contexts, particularly in Hindi-speaking areas of , pag directly signifies a , and "pugmark" retains this connotation in English-influenced discourse. While occasionally used interchangeably with broader terms like "" or "spoor" in informal narratives, "pugmark" distinctly emphasizes the detailed, measurable impression of a , distinguishing it for analytical purposes in .

Historical Development

Origins in India

Pugmark tracking emerged as a systematic, non-invasive method for estimating populations in during the mid-20th century, driven by concerns over the ' sharp decline due to habitat loss, , and human-wildlife conflict. In 1966, Indian forester S.R. Choudhury developed the pugmark census technique while working in Odisha's Similipal forests, aiming to identify individual tigers through the unique patterns of their footprints without direct disturbance to the animals. This approach involved collecting plaster casts or tracings of pugmarks during intensive field surveys, allowing for differentiation based on size, shape, and ridge patterns, which served as a practical alternative to more invasive counting methods prevalent at the time. The technique gained national prominence with its first widespread application in the 1972 All-India Tiger Census, organized by the Indian government in response to alarming reports of dwindling tiger numbers. The census involved systematic surveys across potential tiger habitats throughout , mobilizing thousands of forest personnel to systematically record pugmarks over a short intensive period in key areas such as , Kanha, and Ranthambore, resulting in an estimate of 1,827 tigers nationwide. This figure highlighted the urgency of conservation action, as it represented a significant drop from historical estimates of around 40,000 tigers in the early , underscoring the ecological facing the species. Building on the 1972 census data, the Indian government launched on April 1, 1973, under , establishing a network of protected reserves to safeguard tigers and their habitats. Pugmark tracking was integrated into the project's framework as the primary tool for establishing baseline population estimates and monitoring trends, enabling regular assessments to evaluate effectiveness without relying on lethal control or direct sightings. This initiative marked pugmark methods as a cornerstone of India's early strategy, setting the stage for sustained efforts to reverse tiger population declines.

Evolution and Adoption

During the 1980s and 1990s, the pugmark technique underwent significant refinements to improve its reliability for tiger population estimation. H.S. contributed to early enhancements in 1979 by emphasizing standardized tracings of pugmarks in field surveys, as detailed in the Indian Forester special issue on wildlife census methods. In the 1990s, Dr. L.A.K. Singh advanced identification protocols, enabling differentiation of tiger sexes and ages from hind pugmarks, as published in the Indian Forester in 1993. The () played a central role in , recommending the left hind pugmark as the uniform reference for measurements and analysis in their monitoring manuals. These efforts culminated in formal guidelines issued by Singh in 1999 through , which specified procedures for creating plaster casts—using a 350ml water-to-Plaster of Paris ratio and documenting site conditions—and tracings with tools like the Tiger Tracer on standardized forms to ensure consistency across surveys. The refined pugmark method saw widespread adoption in India's efforts, becoming the primary tool for annual censuses in tiger reserves from the 1970s through 2006. By the 1990s, it was extended to monitor , with Singh's protocols adapted for distinguishing leopard pugmarks from those of tiger cubs and other sympatric species. Applications also emerged for tracking in select reserves, where impressions supplemented dung-based surveys to map movements and individual identification. This expansion supported integrated carnivore and herbivore monitoring under , covering over 17 states by the early 2000s. Following critiques of its limitations, the pugmark method entered a transition phase for , with phased-out reliance starting post-2010 in favor of camera trapping for more accurate, non-invasive data collection. The 2010 All Estimation marked a shift, incorporating camera traps as the core methodology under and NTCA protocols, reducing pugmark use to supplementary sign detection. As of 2025, while no longer primary for national censuses, pugmark tracking persists in forensic applications—such as identifying poached animals or conflict individuals via plaster casts—and in low-tech areas with limited infrastructure, like remote reserves in and , where it aids real-time movement tracking alongside GPS mapping. This retention leverages its simplicity for ground-level patrols, as evidenced in ongoing NTCA advisories.

Collection and Analysis Methods

Field Collection Techniques

Field collection of pugmarks involves non-invasive methods to capture footprints in their natural habitats, primarily focusing on the paws for consistency in tracking. Surveyors select sites where fresh prints are likely to appear, such as along riverbanks, dry stream beds, animal trails, firelines, roads, and near holes, where soft substrates like moist or fine earth preserve clear impressions. Environmental conditions are critical; optimal prints form in substrates with adequate to hold detail without , typically at depths of 0.5 to 1.0 cm, and prints should be fresh—ideally less than one day old—to ensure accuracy. Common collection techniques include paper tracings, plaster of casts, and , each suited to different field conditions. For tracings, a specialized tool like the Tracer—a plate with a hinged frame—is placed over the hind pugmark, and its outline is drawn with a sketch pen on standardized forms, using pins to mark key points without touching the print. Plaster casts are prepared for impressions at least 2-3 mm deep, suitable for clear but shallower prints, by encircling the print with an aluminum strip, mixing plaster of with water, pouring it gently into the track, and allowing it to set for preservation of three-dimensional features. has gained prominence as a modern alternative, involving high-resolution images taken from a fixed-height stand (e.g., 50 cm) with a scale reference like a coin or placed adjacent to the print for , minimizing human error compared to manual methods. Best practices emphasize timing and documentation to enhance data reliability. Collections are ideally conducted during the (December to March) when is firm yet impressionable, reducing and print degradation, though early dry periods ( to ) serve as a secondary option. Surveyors record comprehensive for each sample, including GPS coordinates for precise , and time of , track estimate, of , substrate type (e.g., dusty or sandy riverbank), and stride or step measurements taken without disturbing the prints. Only undistorted hind pugmarks from a normal are selected, with multiple replicates (at least 5-10 per individual set) collected to account for variability. These techniques ensure pugmarks are gathered ethically and efficiently for later analysis.

Identification and Measurement

Pugmarks, once collected from suitable substrates such as soft soil or mud, are analyzed to extract biological attributes including species, sex, and potentially individual identity. This process typically begins with tracing or photographing the impressions using standardized tools like plaster casts or digital cameras to preserve details, followed by precise measurements with calipers or image analysis software such as Sigma Scan Pro or JMP. Key measurements focus on pad dimensions, toe configurations, and ancillary features; measurements typically include total length (from toe tip to heel) and width (maximum perpendicular), as well as pad dimensions excluding toes. For instance, in tigers, adult male hind pugmarks are approximately 12% larger in length and width than females, with total widths typically exceeding 11 cm for males and under 11 cm for females. Toe spacing, such as the distance between toes 2 and 3 (typically 4-6 cm in felids), and claw mark depth (up to 1 cm in non-retractable claw species) provide additional quantitative data, with software enabling sub-millimeter precision on digitized images. Species differentiation relies on morphological criteria like toe count and pad shape. Felids, including tigers, exhibit four prominent toes with rounded, teardrop-shaped pads and typically no visible claw marks due to retractable claws, contrasting with canids that show five toes (including dewclaw impressions) and oval pads with prominent, non-retractable claw marks. Pad shape further aids distinction: felid prints are more compact and rounded, while canid prints appear elongated with a triangular leading toe arrangement. These features allow field experts to classify tracks preliminarily before detailed analysis. Sex determination in species like tigers uses size ratios and shape indices from hind paw measurements, as forepaws show greater overlap. Adult male tiger pugmarks are approximately 12-15% larger in length and width than females, with male pads often exceeding 14 cm in length and 10 cm in width for hind feet, while females measure under 13 cm in length and 9 cm in width; males also display more circular outlines fitting a square, versus the rectangular female form. Discriminant function analysis on variables like total length, width, and pad area achieves 87-100% accuracy in classification when applied to verified samples. Individualization of tigers from pugmarks involves assessing unique morphological traits like ridge patterns, scars, and subtle variations in toe alignment or pad contours, though accuracy is limited without multiple impressions. Techniques such as stepwise discriminant function analysis on 11-128 morphometric variables (e.g., toe areas, angles between toes, and heel-to-toe distances) yield 96-100% identification success with 10 or more pugmarks per individual, outperforming single-print assessments. These methods, including the Footprint Identification Technique (FIT), digitize prints for clustering and validation but falter in variable substrates or with fewer than five prints, achieving only 70-80% reliability in such cases. In forensic contexts, pugmark analysis serves as evidence in investigations by linking impressions to specific individuals or confirming species presence at crime scenes, supporting legal convictions through morphometric matching.

Applications in Wildlife Conservation

Tiger Population Monitoring

The pugmark method plays a central role in tiger population monitoring in by enabling the and enumeration of individuals through their distinctive footprints. In the process, tiger reserves are divided into smaller administrative units such as beats, sections, ranges, and divisions to facilitate systematic coverage. Teams of forest personnel, often numbering in the thousands, conduct intensive searches for fresh pugmarks along roads, trails, riverbanks, and other likely tiger paths during the from December to March, typically over a 7- to 14-day period. Plaster-of-Paris casts or paper tracings of the left hind paw are collected for later analysis, with measurements of pugmark length and breadth recorded to aid . The inaugural systematic all-India tiger census using this pugmark approach was conducted in 1972, establishing a baseline estimate of 1,827 tigers across the country and serving as a for subsequent monitoring efforts, which became standardized on a four-year cycle starting in 2006. Unique pugmarks are distinguished based on morphological variations, including size, shape, stride length, and ridge patterns, allowing trackers to attribute prints to specific individuals while eliminating duplicates through cross-comparisons across blocks and reserves. techniques, such as measuring key dimensions and noting impression details, are essential for this differentiation. Population estimates under the pugmark method are calculated by tallying the total number of unique pugmark sets, each representing an individual , and adjusting for assumed family units—such as one associated with 2–3 females—and incomplete coverage of ranges to extrapolate the overall . This approach assumes territorial behavior and consistent pugmark deposition within a tiger's range, though it relies heavily on the expertise of analysts to classify prints by sex and individuality. Notable case studies illustrate the method's vulnerabilities, particularly its susceptibility to overestimation. The 2001–2002 all-India census reported a national population of 3,642, but subsequent audits revealed significant inflation due to flawed identification and potential fabrication of prints. In the , pugmark data claimed 16–18 tigers in 2002 and 25–28 in 2003, yet investigations in 2005 confirmed the local population had been entirely extirpated by , with officials having fabricated pugmarks to mask the decline. These incidents prompted a reevaluation of the pugmark technique's reliability. Following these revelations, censuses transitioned to protocols post-2010, integrating pugmark tracking with camera , genetic analysis, and modeling for enhanced accuracy. The 2010 , the first to employ this combined approach nationwide, estimated 1,706 s, providing a more robust foundation that addressed pugmark's limitations in detecting population changes. As of the 2022 , pugmark tracking continues as a supplementary tool in protocols, contributing to the estimated national population of 3,682 s.

Use for Other Species

Pugmarks, or paw prints, have been adapted for tracking (Panthera pardus) in addition to , particularly in overlapping habitats within protected areas. Leopard pugmarks are notably smaller, typically measuring 8–10 cm in length, allowing for differentiation from tiger prints during field surveys. This method has been employed since the 1990s in reserves like Similipal Tiger Reserve to map leopard densities and monitor populations in tiger-leopard shared landscapes, aiding in habitat management and conflict mitigation. Beyond felids, footprint tracking extends to elephants (Elephas maximus and Loxodonta africana) and ungulates, where grouped footprints facilitate herd size estimation and movement assessment. In Asian contexts, such as India's Sarguja region, elephant pugmark supports identification, efforts, and migratory route tracking to inform conservation strategies. In African savannas, teams use elephant patterns to estimate group compositions during patrols, enabling rapid response to threats in protected areas like those managed by . For ungulates, including species like (Syncerus caffer) and Asian deer, clustered hoof prints from track surveys help gauge densities and support operations across continents by revealing corridors and poacher intrusion points. Recent advancements highlight emerging applications of pugmarks in behavioral studies and forensics. Track sequences from pugmarks enable analysis of movement patterns, such as territorial ranging and foraging behaviors, in multi-species monitoring protocols as detailed in 2025 research. In wildlife forensics, pugmark impressions serve for species confirmation in poaching investigations, providing non-invasive evidence to verify animal presence and aid .

Advantages and Limitations

Benefits as a Tracking Tool

Pugmark tracking stands out for its cost-effectiveness, relying on low-tech methods that require minimal equipment such as basic and plaster casts, making it particularly suitable for remote areas lacking or advanced . This approach enables wildlife monitoring in resource-limited settings, where more sophisticated technologies like GPS collars may be impractical due to high costs and logistical challenges. Additionally, its non-invasive nature ensures that animals remain undisturbed during data collection, preserving natural behaviors and ecological dynamics without the need for capture or handling. Beyond accessibility, pugmarks provide valuable ecological insights, such as differentiation through variations in and marks, and on movement patterns via stride lengths, contributing to a deeper understanding of . These attributes make pugmark tracking an effective tool for long-term monitoring in sensitive habitats. As a supplementary , pugmark tracking enhances the utility of camera traps by addressing limitations in dense , where visibility and placement challenges reduce photographic capture rates; it offers corroborative evidence of presence and activity in such terrains. Recent 2025 studies underscore its ongoing supplementary role in wildlife monitoring, including integration with digital tools like and GIS for improved accuracy in assessing presence and movements.

Criticisms and Challenges

One major criticism of the pugmark method is its limited accuracy in distinguishing individual tigers, which frequently results in double-counting due to overlapping prints from the same animal and variations in footprint impressions caused by conditions or . This subjectivity in by field personnel has been shown to produce unreliable estimates, with the method's core assumptions—such as complete detection and unique track signatures—often failing, leading to errors in enumeration. Environmental limitations exacerbate these accuracy issues, as pugmarks degrade rapidly under rain or wind, become indistinct on hard or rocky substrates, and are rarely found in dense or large reserves where habitat coverage is incomplete, potentially missing a significant portion of activity areas. For instance, in regions like the or Northeastern , unsuitable terrain renders track detection nearly impossible despite known presence. Scientific critiques from 2003 to 2010, including those by Karanth et al., emphasized the method's inherent biases toward overestimation, as untested assumptions about track coverage and individual segregation masked population declines, such as undetected local extinctions in reserves like and . These analyses revealed that pugmark-based censuses lacked statistical rigor and failed to correlate with ecological realities like prey availability or range overlaps. Consequently, the pugmark method has been largely supplanted by genetic and DNA-based approaches since 2006, which use fecal or hair samples for individual and offer greater precision in abundance estimation without relying on transient tracks. These modern techniques address the pugmark method's flaws by enabling non-invasive, verifiable identification even in challenging environments.

References

  1. [1]
    Five Tips To Help You Spot A Tiger In The Wild! - Wildlife SOS
    Aug 21, 2025 · Usually a bigger and wider pugmark represents that of a male tiger, whereas a more elongated and narrower pugmark represents a female tiger. The ...Missing: definition | Show results with:definition
  2. [2]
    Frequently Asked Questions - Tiger - WWF India
    The most commonly used technique in the past was 'Pugmark Census Technique'. In this method the imprints of the pugmark of the tiger were recorded and used as a ...Missing: definition tracking
  3. [3]
    Why the 'Pugmark Census' Used to Monitor Tiger Populations Failed
    The 'pugmark census' was invented in 1966 by Indian forester SR Choudhury. In this method, during a 1–2-week period, thousands of personnel would simultaneously ...Missing: tracking | Show results with:tracking
  4. [4]
    [PDF] Characterization of Pugmark for Animal Species Identification for ...
    Jun 12, 2020 · No claw marks were present and the specific feature was that dew claw marks were present behind. Dog pugmark heel pad shape was up-side down ...
  5. [5]
    [PDF] Guidelines1or estimating wild tiger populations using the Pugmark ...
    The assessment of tiger numbers through the identification of footprints, for which this volume provides the guidelines, is a traditional method which has.
  6. [6]
    pug - Wiktionary, the free dictionary
    Etymology 3. The noun is borrowed from Hindi पग (pag, “step; (less common) foot; leg”), from Old Hindi पगु (pagu), from Sanskrit पद्ग (padga), from ...
  7. [7]
    PUGMARK - Definition & Meaning - Reverso English Dictionary
    pugmark definition: mark left by an animal's paw. Check meanings, examples, usage tips, pronunciation, domains, related words.Missing: etymology | Show results with:etymology
  8. [8]
    PUGMARK Definition & Meaning - Merriam-Webster
    The meaning of PUGMARK is footprint; specifically : the footprint of a wild mammal (such as tiger or leopard). How to use pugmark in a sentence.
  9. [9]
    PUG Definition & Meaning - Dictionary.com
    Also called pugmark. a footprint, especially of a game animal. verb (used ... First recorded in 1860–65; from Hindi pag “footprint”. Discover More. Word ...
  10. [10]
    pugmark - Wiktionary, the free dictionary
    Etymology. From pug (“pawprint or footprint of an animal”) +‎ mark. Noun. pugmark (plural pugmarks). English Wikipedia has an article on: pugmark · Wikipedia.
  11. [11]
    17 Words for Dog Breeds - Merriam-Webster
    Oct 13, 2025 · The pug that is used for the footprint of a wild mammal (also called pugmark comes pag, a word in both Hindi and Urdu meaning “foot, step.
  12. [12]
    [PDF] Science deficiency in conservation practice: the monitoring of tiger ...
    The pugmark census approach to monitoring tigers. The 'pugmark census' was invented in 1966 by Indian forester S. R. Choudhury (Choudhury, 1970, 1972). In ...Missing: origins | Show results with:origins
  13. [13]
    Imaginary tigers - Down To Earth
    Mar 15, 2005 · Invented in 1966 by Indian forester S R Choudhury, the method requires thousands of people to fan out across jungles for a 1-2 week period ...
  14. [14]
    [PDF] India's Tiger Census Gets Into Guinness Records
    Aug 22, 2020 · The census estimated a total of 1827 tigers according to the. 'Report of Task Force, 1972, Government of India.' This census figure (1972) is ...
  15. [15]
    Tiger Conservation and 50 Years of Project Tiger in India
    First ever all India tiger census by pugmark method was conducted in 1972 which revealed the figure of 1,827 tigers. The Camera Trap Method: After the debacle ...
  16. [16]
    India's successful Project Tiger could do even better, say experts
    Jul 26, 2023 · The rise in numbers is a large change from 1972, when the first all-India tiger census revealed just 1,827 individuals, down from an estimated ...
  17. [17]
    Project Tiger – India's Iconic Wildlife Conservation Program
    Recognising this urgent crisis, the Government of India launched a pioneering initiative on April 1, 1973, called Project Tiger. Spearheaded by then Prime ...
  18. [18]
    [PDF] A Geographical Study of Tiger Projects in India
    Launched on April 1973 Project Tiger has successfully emerged as one ... by the 1972 All India Tiger Census, which enumerated the Tiger population at ...<|control11|><|separator|>
  19. [19]
    Identification of individual tigers (Panthera tigris) from their pugmarks
    Aug 7, 2025 · Tracings and photographs of hind pugmarks were obtained from 23 pugmark-sets of 19 individually known tigers (17 wild and two captive tigers).
  20. [20]
    [PDF] COMPENDIUM OF GUIDELINES/ ADVISORIES/ GAZETTE ...
    Nov 13, 2000 · Tiger Pugmark and other signs: Regular monitoring of tiger signs (pugmark tracings, plaster casts, etc) should be undertaken in every beat ...
  21. [21]
    [PDF] CENSUS OF TIGER AND LEOPARD - Original Wisdom
    For any Tiger or Leopard area, the routes along which pugmark tracking is to be carried out are identified at first in the month of October-November. This is ...Missing: documented | Show results with:documented
  22. [22]
    L.A.K. Singh's Page : Tiger tracking using pugmarks
    Pugmark tracking technique to enumerate tigers and leopards. Dedicated to Late Saroj Raj Choudhury who first refined the technique of pugmark enumeration.
  23. [23]
    [PDF] Tracking the Wild: A Multidimensional Study on the Scientific ...
    Further advancements in pugmark-based studies began to emerge in the 1990s when the Indian government institutionalized the “Pugmark Method” for tiger ...
  24. [24]
    Why India's tiger census is misleading - The Guardian
    May 20, 2015 · The so-called “pugmark census” unrealistically hoped to identify every wild tiger from its paw-prints. That tiger census approach, which was ...<|separator|>
  25. [25]
    'No more pugmark-tracking for tiger count' - The Times of India
    Jul 8, 2009 · Environment and forest minister Jairam Ramesh told Rajya Sabha that the old system of tiger census through pugmarks was 'faulty' and a new ...
  26. [26]
    https://www.theguardian.com/environment/2015/may/20/why-indias-tiger-census-is-misleading
  27. [27]
    Importance of Digital Pugmark Technique & Pugmarks in Wildlife ...
    The 'pugmark census' was invented in 1966 by Indian forester SR Choudhury. ... track of tiger populations to evaluate the effectiveness of conservation schemes.
  28. [28]
    [PDF] Identification of individual tigers (Panthera tigris) from their pugmarks
    We recommend collecting pugmarks from soil depths ranging between 0.5 and 1.0 cm, and advocate the use of pugmark photographs rather than pugmark tracings to.
  29. [29]
    [PDF] Monitoring Tigers in Ranthambhore National Park using the Digital ...
    The. Committee proposed that three different methods should be used: (1) Plaster cast and pugmark tracing method (used traditionally by the Forest Department), ...
  30. [30]
    https://www.wpsi-india.org/images/ranthambore_census_report.pdf
  31. [31]
    [PDF] Gender discrimination of tigers by using their pugmarks
    Tiger gender can be discriminated using pugmark length and width. Male pugmarks are more circular, while female's are misshapen. DFA and logistic regression ...
  32. [32]
    Canine vs. Feline Tracks - How to tell the difference between them.
    The toe pads of cats are more teardrop-shaped than those of dogs. The outer toes of cats are not triangular in shape. Also note that claw marks don't ...
  33. [33]
    Pugmark guide. | Big Cat Sightings In The UK
    Canine pugmark, with non-typical claw marks, due to deep, wet mud. Honing Your Skills: The Value of Studying Canine Tracks.<|control11|><|separator|>
  34. [34]
    Full article: Distinguishing sex of free-ranging tigers using pugmark ...
    Apr 23, 2014 · Univariate analysis​​ The pugmarks of male adult tigers were 11.8% longer and 12.8% wider than female pugmarks, but these differences were not ...
  35. [35]
    (PDF) Distinguishing sex of free-ranging tigers using pugmark ...
    Apr 23, 2014 · The accuracy of sex determination of male and female tigers using total length and total width was 87.5%.
  36. [36]
    'I know the tiger by his paw': A non-invasive footprint identification ...
    This paper describes a statistically robust, cost-effective and non-invasive footprint identification technique (FIT) to identify individual tigers from ...<|control11|><|separator|>
  37. [37]
    Importance of Pugmarks in Veterinary & Wildlife Forensics
    Pugmarks are important evidence in wildlife crime investigation. They alone explicate a lot of information regarding the animal that includes- the species, age ...
  38. [38]
    Tiger Census—How it was Done | Oryx | Cambridge Core
    Apr 24, 2009 · In April-May this year the Indian Forest Service held the first-ever tiger census. Hitherto all assessments of tiger numbers had been based ...
  39. [39]
    [Commentary] Tiger-tiger burning bright: Historical efforts have ...
    Aug 21, 2019 · Since the inception of Project Tiger in April 1973, approximately 14,441 families have been rehabilitated outside tiger reserves from the core ...
  40. [40]
    India tigers fall to 1,411, half earlier estimate | Reuters
    Feb 13, 2008 · The previous census, carried out in 2001 and 2002, said there were 3,642 tigers. A century ago there were 40,000. "The tiger has suffered ...
  41. [41]
    Concerns with India's Tiger Surveys - Shankar IAS Parliament
    These numbers were based on the 'pugmark census method', which assumed that the pugmarks of every tiger could be found, recognised and tallied. · Over time, ...
  42. [42]
    No tigers left in Sariska: CBI - The Times of India
    May 7, 2005 · There are 3,500 to 3,700 Indian tigers left, according to official estimates which are increasingly in dispute, against 40,000 tigers prior to ...
  43. [43]
    [PDF] Status of tigers, co-predators and prey in India, 2010
    The study reports a countrywide increase of 20% in tiger numbers but a decline of 12.6% in tiger occupancy from connecting habitats. The methodology consisted ...
  44. [44]
    How Scientists Count Tigers in India | World Wildlife Fund
    Jan 27, 2022 · Camera trapping has evolved from earlier methods of counting tigers by using their footprints—or pugmarks—which provided inaccurate population ...
  45. [45]
    1636 Wild Tiger Estimation of Tiger Census-2010 - PIB
    Mar 28, 2011 · The latest Tiger Census in India puts estimated tiger population in the country at 1636. Announcing this here today, Shri Jairam Ramesh said that close to 30% ...
  46. [46]
    Measurements of pugmarks of the Indian leopard at Inani Reserve ...
    Length of two pugmarks of the front legs on the sandy-clay streambeds were 8.6cm and 8.7cm (Table 1). It was more or less circular.
  47. [47]
    Carnivora: Felidae: Panthera pardus) in Cox’s Bazar District of ...
    Jun 26, 2017 · The pugmark length of an adult leopard would range from 7–9.5 cm (Talwar & Usmani 2005). On the other hand, the length of a full grown leopard ...
  48. [48]
    [PDF] INFERENCES ON THE BIOLOGYOF TIGER AND LEOPARDIN ...
    Similipal Tiger Reserve (1989-1992): Evidence of male and female. Leopard collected during pugmark tracking. Figures indicate PML or. Pugmark length in cm.<|control11|><|separator|>
  49. [49]
    (PDF) Let's Count Tigers CENSUS OF TIGER AND LEOPARD
    Pugmark Tracking remains a validated method for enumerating tiger and leopard populations. In 2002, 8946 PIPs were laid over 1773 km, yielding 764 pugmark ...
  50. [50]
    Feeding behaviour and pugmark analysis of elephants in Sarguja ...
    Aug 9, 2025 · The pugmark analysis of the elephants offers basic information to track the wild elephant migratory route, identification and census.
  51. [51]
    African Parks Uses Tracking to Combat Poaching and Protect Animals
    African Parks uses GIS to track animal movements, revealing poacher activity, and helps rangers intercept poachers, disrupting supply chains.
  52. [52]
    An investigation of lightweight AI models to classify African ungulate ...
    Sep 26, 2025 · Tracks are cryptic ground evidence that record the history of movement of animals and can be used for monitoring, conservation, anti-poaching ...
  53. [53]
    A Multidimensional Study on the Scientific, Ecological, and ...
    Aug 7, 2025 · The paper outlines the significance of pugmarks in preventing human-wildlife conflict, aiding anti-poaching operations, and contributing to ...
  54. [54]
    Pugmark techniques in tiger population estimation: a review
    Oct 1, 2025 · The pugmark technique involves collecting and analyzing tiger footprints to identify individual animals and estimate population sizes. Despite ...Missing: origins | Show results with:origins<|control11|><|separator|>
  55. [55]
    Science deficiency in conservation practice: the monitoring of tiger ...
    Feb 28, 2006 · We study this phenomenon in the context of monitoring tiger populations in India, by evaluating the 'pugmark census method' employed by wildlife ...
  56. [56]
    Can the abundance of tigers be assessed from their signs? - Jhala
    Nov 18, 2010 · Tiger pugmark encounter rates explained 84% of the observed variability in tiger densities. After removal of an outlier (Corbett), square root ...Introduction · Materials and methods · Results · Discussion
  57. [57]
    Genotyping faecal samples of Bengal tiger Panthera tigris tigris for ...
    Oct 17, 2006 · We describe here a pilot study using DNA extracted from faecal samples of tigers for the purpose of population estimation.