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Redwing

The redwing (Turdus iliacus) is a small bird in the thrush family Turdidae, native to and the western Palearctic, measuring 20–24 cm in length with a wingspan of 33–35 cm. It is characterized by its brown upperparts, pale underparts with dark streaks on the breast, a prominent creamy-white (eyebrow stripe), and distinctive orange-red patches on the flanks and underwing coverts, which are most visible in flight. Slightly smaller than the related (Turdus philomelos), the redwing is the smallest regularly observed in the , where it is primarily a winter visitor rather than a breeder. Breeding across northern Europe including Scandinavia and Iceland, and northern Russia in open woodlands, coniferous forests, and scrubby areas, the redwing constructs a cup-shaped nest of grass, , and twigs lined with or feathers, typically low in trees or shrubs, usually less than 4 m above ground. Females lay 4–6 pale blue-green eggs speckled with brown, incubating them for 12–14 days while males provide food; fledglings leave the nest after about 13 days. The diet shifts seasonally: during breeding, it consists mainly of such as , snails, and foraged on the ground; in winter, it relies heavily on berries from hedgerows, orchards, and gardens, with large flocks forming when northern food sources are scarce. Its song is a series of simple, descending liquid notes delivered from a high , varying by across regions, while calls include a sharp "seep" or buzzy "dsssi" for alarm. The redwing is highly migratory, with northern populations traveling south and west to winter in , including the UK, , and Iberia, often in mixed flocks with fieldfares (Turdus pilaris); vagrants occasionally reach , particularly Newfoundland. Populations are stable overall, classified as Least Concern by the IUCN due to its wide range and adaptability, though it faces threats from habitat loss, affecting berry crops, and hunting in some southern European countries. In the UK, it is on the Amber List of concern, reflecting moderate declines in some areas but strong winter passage numbers.

Taxonomy and systematics

Etymology and naming

The common name "redwing" derives from the bird's distinctive reddish-orange patch on the underwing coverts, which becomes prominently visible during flight. The scientific name Turdus iliacus originates from Latin, with Turdus meaning "thrush," reflecting its membership in the thrush group, and iliacus derived from ilia, referring to the flanks or hips, in allusion to the species' reddish coloration on the flanks. This binomial was formally established by in the 10th edition of Systema Naturae in 1758, although subsequent publications by Linnaeus in 1766 and 1767 introduced some nomenclatural confusion regarding synonyms like Turdus musicus, leading various authors to variably attribute the description to either 1758 or 1766; the 1758 date is now accepted as the original authority. No major taxonomic revisions to the species' naming have occurred since, with the most recent comprehensive review in 2024 confirming its stability. The redwing is classified within the family Turdidae, the thrushes, and the genus Turdus, where it shares close phylogenetic relations with other Palearctic species such as the (Turdus philomelos), with which it exhibits morphological and behavioral similarities.

Subspecies and classification

The redwing (Turdus iliacus) is classified within the thrush family Turdidae and the genus Turdus, with two recognized based on geographic variation and morphological differences. The nominate subspecies, T. i. iliacus, breeds across northern Europe from Scandinavia eastward to central Siberia and winters in western and southern Europe, North Africa, and the Near East; it is characterized by paler, warmer brown upperparts, narrower and more distinct dark streaking on the whitish underparts, a vague breast band, a narrower pale eyeline, a larger pale patch on the rear flanks, and paler legs, with an average size of about 21 cm in length. In contrast, T. i. coburni breeds in and the and winters mainly in ; it is slightly larger (about 3% longer on average, reaching 22 cm), with darker, colder brown upperparts, broader and darker streaking on the underparts that often forms solid dark patches and a more defined breast band, a broader dark eyeline, a reduced pale flank patch, and darker legs. Phylogenetic analyses confirm the of the Turdus, placing T. iliacus within a Palearctic as the sister species to the Eurasian (T. merula), with strong support from ultraconserved element (UCE) genomic data; this relationship highlights the 's complex biogeographic history involving multiple dispersals between the Old and New Worlds. A 2020 phylogenomic study estimated the divergence of the T. iliacusT. merula lineage from other Palearctic thrushes around 5–7 million years ago, reflecting ancient diversification within the . Hybridization between T. iliacus and T. merula has been documented in , though such events are rare and typically occur in overlapping wintering or breeding ranges.

Physical characteristics

Plumage and coloration

The redwing (Turdus iliacus) exhibits a characterized by brownish-olive upperparts, including the crown, back, and rump, which provide effective in its habitats. A prominent pale extends from the bill over the eye, contrasting with the darker ear coverts and giving the bird a distinctive facial pattern. The underparts are cream-colored with bold dark streaks, particularly dense on the breast and sides, creating a mottled appearance. One of the most striking features is the reddish-orange coloration of the axillaries and underwing coverts, which becomes vividly apparent during flight or when the wings are spread, distinguishing the redwing from similar thrushes. The flanks also show rusty-red patches, enhancing this diagnostic trait. is minimal, with males and females displaying nearly identical patterns and colors, though males may average slightly brighter tones in breeding condition. Juveniles possess plumage similar to adults but with notable differences: the upperparts appear scaly due to buffy fringes on the feathers, and the underparts feature spots rather than streaks, especially on the breast where the spots are largest and more rounded. These young birds undergo a post-juvenile moult, typically completing it by late summer, to acquire adult-like streaked underparts and smoother upperparts. Seasonal variations primarily involve feather wear, with fresher in autumn appearing more vibrant and olive-toned, while winter birds show faded, abraded edges without significant color shifts. Subtle differences exist among , such as the Icelandic T. i. coburni having redder flanks and more saturated underparts compared to the paler nominate T. i. iliacus.

Size, measurements, and morphology

The redwing (Turdus iliacus) measures 20–24 cm in length, with a of 33–35 cm and a body mass ranging from 50–75 g, averaging approximately 62 g. These dimensions position the redwing as one of the smaller species within the Turdus, notably more compact than the larger (T. merula) at around 24 cm in length and heavier congeners like the (T. philomelos), which shares a similar length but exceeds 70 g on average. The bill is slender and straight to slightly decurved, with a dark brown to blackish upper and a yellow base on the lower , facilitating precise manipulation of food items. The legs and feet are pinkish to grayish pink, featuring robust tarsi and toes that support typical of thrushes. Wing morphology includes long, pointed primaries that enhance aerodynamic efficiency for long-distance , with a wing measuring 11.8–13.6 cm. This structure, combined with the bird's overall compact build, aids in sustained flight across its Palearctic breeding grounds to wintering sites in .

Vocalizations

Song characteristics

The song of the redwing (Turdus iliacus) is a short phrase of simple, monotonous structure consisting of introductory clear notes followed by a twittering , typically delivered from an elevated such as a tree . It lasts 2–3 seconds and is often described as sweet descending notes with twittery burbling, such as "trúi-trúi-trúi trip-trr-bziriri-rrit." This distinguishes it from the more complex songs of related thrushes. Most males sing one song type, though some use two distinct variants, enabling subtle differences in phrasing and that contribute to individual recognition. At least 38 regional dialects are known from southern , with song patterns varying across populations due to cultural transmission and local ecological conditions, as documented in long-term studies of breeding sites. These dialects maintain within groups but show gradual changes over time, reflecting ongoing evolution in . The serves dual functions in territorial defense and mate attraction, with singing rates highest in early upon territory establishment and pair formation. Playback experiments confirm that males respond aggressively to conspecific , particularly the twittering , underscoring its in intra-sexual . The is dominated by whistles in the introductory part and rapid trills in the concluding , which enhance detectability in forested . Song amplitude and structure also signal male arousal levels during interactions. Environmental factors influence song variation, with habitat acoustics shaping phrase degradation and transmission. Unlike calls, which are shorter and more abrupt, the song's elaborate form is reserved for contexts. A subsong, consisting of low twittering, is produced by flocks during or winter, and by juveniles starting at 17–18 days old.

Calls and other sounds

The redwing (Turdus iliacus) employs a range of short, situational vocalizations for communication, contrasting with its longer, more melodic used primarily in territorial . The alarm call is a sharp, rattling series such as "chet-chet-chet" or "trrt-trrt-trrt", typically produced when the is disturbed or perceives a near its nest or area. These calls alert nearby individuals and may accompany defensive flights toward intruders, as observed during nesting disturbances. Contact calls are soft and repetitive "seep" notes, often given by individuals in flocks to maintain group cohesion while searching for food on the ground. During flight, particularly in nocturnal , the redwing emits high-pitched, drawn-out calls like "tseer", "tseee", or "srieh", which facilitate coordination in loose flocks overhead. A softer nasal variant, described as "kuk", may also occur in flight or contexts. In mobbing scenarios against predators, the alarm calls intensify and are combined with approach behaviors to harass the threat, though physical contact is rare.

Distribution and habitat

Geographic range

The redwing (Turdus iliacus) breeds across a broad expanse of the northern Palearctic, from Iceland and the Faroe Islands in the west through Scandinavia, the British Isles (with a small population in northern Scotland), the Baltic region, and northern Russia to western Siberia in the east, generally reaching up to 70°N latitude, extending eastward to the Altai region and lower Kolyma River in Siberia. This distribution encompasses subarctic and boreal zones, with the nominate subspecies T. i. iliacus predominant across most of the mainland range and T. i. coburni restricted to Iceland and the Faroe Islands. The ' breeding range has undergone historical expansion following post-glacial recolonization after the Pleistocene , allowing northward spread into newly available habitats as ice sheets retreated. In recent decades, slight range extensions have occurred in . Wintering distribution shifts southward to milder regions, primarily in southern and western (including the , , , and the ), northwest (such as and ), and the (extending to the coasts, , and northern ). Vagrant individuals occasionally appear outside the typical ranges, with records in —particularly along the east coast from Newfoundland to the , the west coast including , and multiple records in , including sightings in Seward (2011) and Utqiagvik (2022), as of 2025—where they may briefly overlap with the resident (Turdus migratorius). Rarer vagrants have been documented in , including and .

Habitat preferences and requirements

The redwing (Turdus iliacus) primarily breeds in open woodlands, forest edges, , and habitats featuring scattered trees, often within forests dominated by , , , and interspersed with open country in lowlands and low hills. These environments provide a mix of dense cover for nesting and open areas for foraging, supporting the species' requirements for insect-rich and berry-producing shrubs. The shows particular adaptations to conditions, thriving in cooler, northern latitudes where coniferous and trees offer structural diversity. In winter, redwings shift to more varied low-elevation habitats including open woodlands, orchards, scrubby thickets, farmland, and gardens, where berry-bearing bushes and grassy areas are abundant; during harsh weather, they congregate in coastal regions and dense hedgerows for roosting. These sites allow communal roosting in protective cover, such as thick scrub or hedgerows, to mitigate cold and predation risks. Nest sites are typically placed in trees or shrubs 2–15 m above the , though occasionally in low shrubs or on the under dense vegetative cover to shield eggs and fledglings from predators and weather. The species occupies elevations from to 700 m, with rare occurrences up to 2,100 m, reflecting its affinity for accessible and terrains. Habitat fragmentation, driven by clear-cutting and , reduces suitable sites by disrupting forest mosaics and density, leading to a 61% decline in redwing densities in affected southern forests in between 1993 and 2022. Studies from the highlight climate-driven shifts, including warming temperatures and altered , that are pushing the ' latitudinal center northward while fragmenting remaining habitats and increasing vulnerability to .

Behavior

Breeding and reproduction

The redwing breeds primarily in northern and eastern Europe, including Scandinavia and Iceland, with the season typically spanning April to June in these ranges, allowing for 1–2 broods per year. Nest construction is undertaken almost exclusively by the female, who builds a compact, cup-shaped structure from interwoven grass, moss, twigs, and lichen, often bound with mud and lined with fine grass or leaves; the nest is usually positioned low to the ground in dense vegetation, shrubs, or against tree trunks, at heights of 0.1–2 m. The clutch comprises 4–6 eggs, which are pale greenish-blue with fine brown or black spots and blotches concentrated toward the larger end; egg-laying occurs over 1–4 days, with the female incubating the clutch alone for 10–14 days until hatching. The altricial nestlings, covered in sparse down, remain in the nest for 12–15 days before fledging; both parents feed the young during this period and continue providing care for approximately 2 additional weeks post-fledging, after which the female may initiate a second brood while the male tends the first. Redwings are socially monogamous, with pair bonds forming through male song displays that attract females and help establish breeding territories; the male's singing rate peaks during mate attraction and declines after egg-laying begins.

and

The redwing (Turdus iliacus) exhibits an omnivorous that shifts seasonally to meet nutritional demands. During the breeding season in spring and summer, predominate, with forming a major component alongside , larvae, snails, and slugs. In autumn and winter, the transitions to include a high proportion of matter, particularly berries and fruits such as (Sorbus aucuparia) and (Crataegus monogyna), which provide essential energy for and survival in colder conditions. Nestlings receive a dominated by as a high-protein subset of breeding-season foraging. Foraging occurs primarily on the ground and in low vegetation, employing techniques such as bill-probing into moist soil to extract buried like , from foliage, and surface-pecking for seeds or fallen fruits, often in loose flocks of 10–200 individuals. The ' slender, slightly decurved is adapted for probing and extracting prey from substrates, facilitating access to hidden , while acute vision, including sensitivity to reflectance, enhances detection of visually conspicuous fruits like bilberries (), as demonstrated in laboratory choice experiments where redwings preferentially selected UV-reflecting berries. Redwings frequently forage in mixed-species flocks with other thrushes such as fieldfares (Turdus pilaris) and blackbirds (Turdus merula), resulting in for shared and resources, particularly in resource-limited winter habitats. In agricultural landscapes, their consumption of berries and fruits in orchards and hedgerows can lead to localized impacts on soft fruit crops, prompting considerations in agro-ecosystems to balance with crop protection.

Migration and movements

The redwing (Turdus iliacus) exhibits partial migration, with northern populations from , , and undertaking long-distance journeys southward, typically covering 2,000–4,000 km to wintering grounds in northwest and , while southern European populations remain largely sedentary or make only short movements. This latitudinal variation in migratory allows northern birds to escape harsh winters, whereas milder climates enable resident populations in areas like and to persist year-round. Post- departure from northern breeding grounds begins as early as late July in eastern , extending through October across western populations, with arrivals at wintering sites peaking from September to November. migration occurs primarily from March to April, aligning with improving conditions at breeding areas and often involving nocturnal flights to cover distances efficiently. During these journeys, redwings frequently form large flocks numbering in the thousands, sometimes reaching up to 10,000 individuals, which provides safety from predators and aids in locating food resources en route. In years of poor production in northern regions, irruptive s intensify, driving flocks farther south or westward beyond typical routes to exploit alternative food sources. Redwings navigate using a multifaceted system of cues, including celestial orientation from the setting sun and stars, geomagnetic fields for directional sensing, and visual landmarks such as coastlines and rivers for fine-scale adjustments. Stopover sites, crucial for refueling, are predominantly wetlands and swampy grounds where birds exploit and berry resources to build fat reserves for continued flight. Recent analyses of ringing recoveries from the , including data from the European-African migration network, reveal route variations influenced by weather, with birds from occasionally shifting to more westerly paths over the North Atlantic during favorable tailwinds or avoiding stormy fronts by detouring inland.

Ecology

Predators and natural threats

The redwing faces predation from various avian species, particularly that target adults and fledglings during foraging or migration. Sparrowhawks (Accipiter nisus) are a primary threat, ambushing redwings in edges and gardens with high-speed pursuits, often preying on thrushes including this species. Merlins (Falco columbarius), small falcons, also hunt redwings, especially during autumn migration when flocks are vulnerable, as observed in pursuits where the thrush evades capture through agile flight. , such as short-eared owls (Asio flammeus), occasionally capture adult redwings, exploiting low-light conditions to surprise them near roosts. Mammalian predators pose significant risks to nests, which are typically built low in shrubs or trees. Domestic and feral cats (Felis catus) raid nests for eggs and chicks, particularly in suburban areas where redwings breed or overwinter. Red foxes (Vulpes vulpes) opportunistically prey on ground-level or low nests, digging out or reaching in to consume contents, contributing to high failure rates in open habitats. Weasels (Mustela nivalis) are agile nest predators, squeezing into tight spaces to devour eggs and nestlings, especially in rural breeding grounds. Nest predation rates can reach up to 55% in monitored populations, as recorded in studies where hooded crows ( cornix) were the most frequent predator, though other corvids and mammals contribute regionally. Redwings mitigate these risks through nest , constructing cups from grass, moss, and lichens that blend with surrounding foliage, reducing visibility to visual hunters. Adults also employ alarm calls—a sharp "seep-seep"—to warn of approaching predators, prompting flock evasion or . Beyond predation, abiotic factors like threaten redwing survival. Harsh winters with prolonged and lead to by limiting access to berries and , causing heavy mortality in overwintering flocks. Storms during migration, including autumn gales across the , increase disorientation and exhaustion, with collisions and drowning reported in adverse conditions. Cold, wet summers further reduce breeding success by impairing foraging and chick development.

Parasites, diseases, and interactions

Redwings (Turdus iliacus) are susceptible to several common ecto- and endoparasites, including feather lice (Phthiraptera), , and intestinal helminths such as . Feather lice, such as species in the genera Brueelia and Menacanthus, have been recorded on redwings in , with infestations typically low but contributing to damage and irritation during breeding seasons. , particularly , frequently infest redwings, with thrushes showing high mean infestation intensities in forested habitats; these ticks can transmit pathogens like , exacerbating health risks during migration. Intestinal worms, including like those in the genus Anomotaenia, parasitize the of redwings, potentially leading to nutrient and reduced body condition, as documented in populations. Viral diseases also pose significant threats to redwing health. Avian pox, caused by avipoxviruses, affects European songbirds including thrushes, manifesting as cutaneous lesions on unfeathered areas; severe cases reduce , particularly in juveniles, by impairing vision and foraging. (WNV) exposure has been detected in migratory European passerines, with seroprevalences detected in Turdus species in ; in naïve populations, WNV can cause mortality rates exceeding 50% through neurological damage and . Additionally, antibodies occur at 37% prevalence in Swedish redwings, indicating widespread exposure that may contribute to subclinical infections and reduced fitness. Redwings engage in mutualistic interactions with plants through frugivory, acting as key seed dispersers for berry-producing species. By consuming fruits of rowan () and crowberry () in , redwings deposit viable seeds away from parent plants via endozoochory, enhancing plant recruitment in open habitats with dispersal distances averaging 100-500 meters. This symbiosis benefits plants by promoting and colonization of new areas, while providing redwings with essential energy during pre-migratory fattening. Interspecific competition occurs with congeners like the Eurasian blackbird (Turdus merula), particularly for berry and invertebrate resources in wintering grounds. Redwings exhibit niche segregation by at lower strata and in more open areas compared to blackbirds, reducing overlap; however, in resource-scarce winters, introduced or expanding thrush populations can displace redwings from optimal feeding sites, impacting overwinter survival. Recent research highlights the redwing's gut 's role in health, with compositional shifts during long-distance flights reducing bacterial diversity but enriching taxa like that bolster immune function and resistance. Studies from 2020-2025 on Palearctic migrants show microbiome remodeling influences energy allocation and disease susceptibility, with linked to higher haemosporidian rates in fatigued individuals.

Conservation status

The Redwing (Turdus iliacus) is currently classified as Least Concern on the , following its reassessment in 2025 from Near Threatened, reflecting a global population of approximately 40–70 million mature individuals. In , which hosts the majority of the population, estimates range from 16–28 million mature individuals, based on 8–14 million pairs. Monitoring by indicates an overall stable to increasing trend across , with a 12% rise in numbers from 2012 to 2022. However, long-term data reveal declines in specific regions, particularly agricultural landscapes, where populations have decreased by up to 25–30% since 1980 due to habitat intensification and loss of suitable foraging areas. Key threats include intensified , which reduces prey availability critical for adults and nestlings, as well as illegal during and winter in Mediterranean countries (e.g., an estimated 50,000–300,000 individuals trapped annually in ). exacerbates these pressures by altering berry production cycles, leading to unpredictable winter food supplies; warmer temperatures and shifting have been associated with reduced rowanberry and yields in , forcing greater distances or starvation risks during harsh winters. Over-grazing in habitats further diminishes ground cover for nesting and . Ongoing monitoring highlights potential range shifts linked to climate warming, with northward expansions in distribution and increased records beyond traditional migration routes, such as more frequent sightings in southern and eastern . These shifts may reflect adaptive responses to changing temperatures but also signal if resources fail to synchronize with altered timings. National surveys up to 2025, including those from the British Trust for Ornithology and equivalent programs in , continue to track these dynamics, emphasizing the need for integrated threat assessments.

Conservation measures and efforts

The Redwing (Turdus iliacus) benefits from protection under the EU Birds Directive (Directive 2009/147/EC), which mandates member states to implement measures for the conservation of wild bird populations, including safeguarding and regulation of hunting where applicable. Sustainable hunting is permitted in eight Mediterranean EU countries under Annex II(b) of the directive, with monitoring required to ensure it does not jeopardize . Additionally, the species is covered by the on the Conservation of European Wildlife and Natural Habitats, which promotes international cooperation for migratory species protection across . In the , 486 sites have been designated at least partly for the Redwing, primarily in Mediterranean countries where it occurs during and winter, providing critical stopover and habitats. National parks and protected areas in breeding ranges, such as those in and the , further support populations by preserving forested and woodland environments essential for nesting. In the , the Redwing is protected under the , prohibiting intentional disturbance or harm. Agricultural practices play a key role in conservation, particularly through hedgerow preservation, which maintains berry sources like and vital for winter flocks. Agri-environment schemes, including initiatives, enhance invertebrate prey availability in grasslands, supporting breeding success in fragmented rural landscapes. Habitat restoration projects focus on reconnecting hedgerows and restoring lowland neutral grasslands to mitigate fragmentation, thereby improving connectivity for migratory movements. Research and monitoring efforts rely heavily on citizen science programs, such as the British Trust for Ornithology's (BTO) Garden BirdWatch and Constant Effort Sites scheme, which track annual population fluctuations and breeding productivity. Globally, eBird facilitates real-time data collection from observers, enabling analysis of patterns and winter distributions to inform targeted interventions. These initiatives have contributed to recent assessments, including the IUCN's 2025 reassessment classifying the Redwing as Least Concern following evidence of stable or recovering trends in key regions.

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