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Chenodeoxycholic acid

Chenodeoxycholic acid (CDCA), also known as chenodiol, is a primary naturally synthesized in the human liver from through a series of enzymatic steps. It has the C24H40O4 and a molecular weight of 392.572 g/mol, featuring a nucleus with hydroxyl groups at the 3α and 7α positions. As one of the two main primary s—alongside cholic acid—CDCA is secreted into , where it acts as a physiological to emulsify and facilitate the and absorption of dietary fats, sterols, and fat-soluble vitamins in the . It undergoes conjugation with or in the liver before release and is efficiently reabsorbed via the , with about 95% recycled daily. In its biological role, CDCA helps regulate by promoting its excretion from the liver into and modulating flow and lipid secretion. A portion of CDCA is metabolized by gut into the secondary lithocholic acid, which is largely excreted in , contributing to elimination. This also activates nuclear receptors such as the farnesoid X receptor (FXR), influencing genes involved in synthesis, transport, and inflammation. Medically, chenodeoxycholic acid is approved for the dissolution of small, radiolucent cholesterol gallstones in patients with functioning gallbladders who are at high risk for surgery or elect non-surgical options. Treatment typically involves oral doses of 13–16 mg/kg daily in divided doses for up to two years, achieving partial or complete gallstone dissolution in 15–30% of cases, though recurrence rates can reach 50% upon discontinuation. It is also indicated for treating cerebrotendinous xanthomatosis (CTX), a rare autosomal recessive disorder caused by sterol 27-hydroxylase deficiency, where it replaces deficient bile acids, normalizes cholestanol levels, and inhibits overproduction of atypical bile acids by suppressing cholesterol 7α-hydroxylase (CYP7A1); it is approved by regulatory agencies such as the EMA and FDA (as of 2025 for adults in the US). For CTX, dosing starts at 5–15 mg/kg/day in pediatrics and up to 750–1,000 mg/day in adults, with monitoring of plasma cholestanol and liver function. The therapeutic mechanism of CDCA involves decreasing hepatic cholesterol synthesis via inhibition of and reducing biliary saturation to promote dissolution. However, it can cause dose-related due to its cathartic effects and transient elevations in serum aminotransferases in up to 30% of patients, with rare instances of clinically apparent linked to its metabolite lithocholic acid. Largely supplanted by for therapy due to better tolerability, CDCA remains a key treatment for CTX and select deficiencies.

Chemical properties

Molecular structure

Chenodeoxycholic acid (CDCA) is a primary with the molecular formula C24H40O4 and a molecular weight of 392.58 g/. Its systematic IUPAC name is 3α,7α-dihydroxy-5β-cholan-24-oic acid, reflecting its classification as a dihydroxy of the cholanoic acid backbone. The molecule features a characteristic composed of four fused rings: three six-membered rings (A, B, and C) and one five-membered ring (D), forming a curved core derived from . Attached to this nucleus are two hydroxyl groups oriented at the 3α and 7α positions on rings A and B, respectively, which contribute to its , and an eight-carbon at C-17 terminating in a group at C-24. These structural elements endow CDCA with amphipathic properties, where the hydrophobic rings contrast with the hydrophilic hydroxyl and carboxyl moieties. In comparison to cholic acid, another primary , CDCA lacks the additional hydroxyl group at the 12α position on ring C, resulting in only two hydroxyl groups instead of three. This difference influences the molecule's overall hydrophobicity and characteristics. The of CDCA is defined by the 5β configuration of the A/B ring junction, which imparts a cis fusion between rings A and B, along with α-orientation (axial, below the plane) of the hydroxyl groups at C-3 and C-7, and specific chiral centers at C-8 (β), C-9 (α), C-10 (β), C-13 (β), C-14 (α), C-17 (β), and C-20 (R). In three-dimensional representations, this arrangement positions the polar groups on the concave α-face of the steroid nucleus, enhancing its detergent-like behavior in formation and contributing to its biological in aqueous environments.

Physical and chemical characteristics

Chenodeoxycholic acid is a crystalline powder that may absorb under ambient conditions. It exhibits low in , with a reported value of approximately 0.09 mg/mL, rendering it practically insoluble, but it dissolves readily in organic solvents such as (up to 20 mg/mL), , acetone, and glacial acetic acid. increases in alkaline solutions owing to the of the carboxyl group. It is weakly soluble in and but insoluble in petrol ether and . The compound has a of 165–167 °C. The pKa of its group is approximately 4.6. Chenodeoxycholic acid demonstrates good , remaining unchanged for up to 24 months when stored at without special packaging requirements; it is unaffected by exposure to light and resists degradation under acidic, basic, or oxidizing conditions, although thermal decomposition occurs above 130 °C. Commercially, chenodeoxycholic acid is produced through from animal sources, such as porcine by-products, or via semi-synthetic routes involving microbial transformation of precursor bile acids like cholic acid. from is possible but less common for large-scale production due to complexity.

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Biosynthetic pathway

Chenodeoxycholic acid (CDCA) is a primary synthesized in hepatocytes from primarily via the classic (neutral) pathway, which accounts for the majority of bile acid production in the liver. This pathway initiates with the rate-limiting step catalyzed by 7α-hydroxylase (CYP7A1), a microsomal that hydroxylates at the 7α position to form 7α-hydroxycholesterol. Subsequent enzymatic modifications oxidize this intermediate via 3β-hydroxy-Δ⁵-C₂₇-steroid dehydrogenase (HSD3B7) to 7α-hydroxy-4-cholesten-3-one, followed by stereospecific reduction of the Δ⁴ double bond by Δ⁴-3-oxosteroid-5β-reductase (AKR1D1) and oxidation at the 3α position by 3α-hydroxysteroid dehydrogenase (AKR1C4), yielding 3α,7α-dihydroxy-5β-cholestanoic acid (DHCA). The side chain is then oxidized at the 27 position by sterol 27-hydroxylase (CYP27A1), a mitochondrial and peroxisomal , leading to oxidative cleavage and formation of unconjugated CDCA. Unlike the parallel synthesis of cholic acid, CDCA production bypasses 12α-hydroxylation by sterol 12α-hydroxylase (CYP8B1). The biosynthetic pathway is subject to regulation primarily through the farnesoid X receptor (FXR), a activated by s such as CDCA. FXR activation in the liver induces small heterodimer partner (SHP), which represses CYP7A1 transcription by inhibiting liver receptor homolog-1 (LRH-1). Additionally, intestinal FXR activation by reabsorbed bile acids upregulates fibroblast growth factor 19 (FGF19), which travels to the liver and further inhibits CYP7A1 expression via signaling through fibroblast growth factor receptor 4 (FGFR4) and β-klotho, maintaining bile acid homeostasis. In humans, CDCA comprises approximately 40% of the total pool, reflecting its significant contribution alongside cholic acid. A minor alternative (acidic) pathway, initiated extrahepatically by CYP27A1-mediated 27-hydroxylation of to 27-hydroxycholesterol followed by 7α-hydroxylation via oxysterol 7α-hydroxylase (CYP7B1), converges with the classic pathway to produce CDCA but plays a limited role in overall synthesis, particularly in adults.

Physiological functions

Chenodeoxycholic acid (CDCA), a primary synthesized in the liver, serves as a key component of bile salts that facilitate the emulsification and of dietary in the . By forming micelles with , phospholipids, and monoglycerides, CDCA solubilizes these hydrophobic molecules, enabling their efficient uptake by enterocytes and preventing their precipitation in the aqueous environment of the intestinal . This process is essential for the of fats and the of fat-soluble vitamins, contributing to overall nutrient . CDCA participates in the , a highly efficient recycling mechanism that conserves within the body. Approximately 95% of secreted , including CDCA, are reabsorbed in the terminal via the apical sodium-dependent transporter (ASBT, also known as SLC10A2), which actively transports them into enterocytes against a concentration gradient using sodium co-transport. From there, CDCA is shuttled to the and returned to the liver via basolateral transporters like OSTα/OSTβ, allowing it to be recycled 10-12 times daily to support repeated cycles of secretion and intestinal function. This recirculation minimizes the need for , with only about 5% of the pool lost in each day. As a for the farnesoid X receptor (FXR), CDCA exerts regulatory effects on metabolic pathways. Upon binding to FXR in hepatocytes and enterocytes, CDCA induces the expression of small heterodimer partner (SHP), which suppresses transcription of the rate-limiting enzyme in synthesis, cholesterol 7α-hydroxylase (CYP7A1), thereby providing to maintain homeostasis. FXR activation by CDCA also promotes glucose and by enhancing insulin sensitivity, inhibiting , and stimulating oxidation in the liver. Additionally, CDCA-mediated FXR signaling modulates by repressing pro-inflammatory cytokines such as TNF-α and IL-1β in immune cells and endothelial tissues. CDCA activates the G protein-coupled membrane receptor TGR5 (also known as GPBAR1), which mediates diverse physiological responses independent of FXR. In enteroendocrine L cells, TGR5 stimulation by CDCA promotes the secretion of (GLP-1), an hormone that enhances insulin release and suppresses , aiding postprandial glucose control. In and , CDCA-induced TGR5 signaling increases energy expenditure through cAMP-dependent activation of type 2 (DIO2), which converts thyroxine to active , thereby boosting . Furthermore, TGR5 activation by CDCA exerts anti-inflammatory effects in macrophages by inhibiting signaling and reducing the production of pro-inflammatory mediators like IL-6 and MCP-1. Within the bile acid pool, contributes to compositional diversity and influences the overall hydrophobicity index, which affects properties and . Compared to the more hydrophilic cholic acid, CDCA is relatively hydrophobic, enhancing the pool's ability to solubilize while balancing risks associated with excessive hydrophobicity. This property supports by promoting the biliary excretion of free into , where it is solubilized for elimination via the feces, thus preventing . Through these mechanisms, CDCA helps regulate systemic levels and maintains intestinal barrier .

Clinical uses

Gallstone dissolution

Chenodeoxycholic acid, marketed under the brand name Chenodiol, received FDA approval in 1983 for the oral dissolution of small, radiolucent, non-calcified cholesterol gallstones measuring less than 15 mm in diameter, specifically in patients with well-opacifying gallbladders for whom elective surgery would otherwise be considered. This approval was based on clinical evidence demonstrating its ability to nonsurgically treat suitable gallstones by altering bile composition. The therapeutic mechanism involves suppressing hepatic cholesterol synthesis and secretion into bile, which decreases the biliary cholesterol saturation index (SI) to below 1, rendering bile unsaturated and capable of solubilizing cholesterol from gallstones over time. This desaturation promotes gradual dissolution, particularly effective for cholesterol-rich stones that appear radiolucent on imaging. Standard dosing regimen is 13 to 16 mg/kg body weight per day, administered orally in two divided doses (morning and evening), typically for 6 to 24 months depending on stone size and response. Treatment success, defined as complete dissolution, ranges from 40% to 60% for small stones under 10 mm, though rates are lower for larger ones; for instance, the landmark National Cooperative Gallstone Study (NIH-NIDDK, 1980s) reported complete dissolution in 13.9% of patients after 2 years of therapy at 750 mg/day. Appropriate patient selection is critical, favoring those with floating gallstones on oral cholecystography, which indicates a high content and low density conducive to dissolution. is contraindicated in cases of calcified (radiopaque) stones, stones exceeding 15 mm, nonvisualizing gallbladders, or conditions like that increase pigment stone risk. Combination with (UDCA) at equimolar doses has shown improved efficacy, achieving higher dissolution rates (up to 70% in some trials) while minimizing chenodeoxycholic acid-associated side effects like and . This approach leverages the complementary actions of both acids to more effectively desaturate and inhibit stone formation.80004-3/fulltext)

Cerebrotendinous xanthomatosis treatment

Chenodeoxycholic acid (CDCA) is the standard treatment for cerebrotendinous xanthomatosis (CTX), a rare autosomal recessive disorder caused by mutations in the CYP27A1 gene, which encodes sterol 27-hydroxylase and leads to deficient production of chenodeoxycholic acid, resulting in accumulation of cholestanol in plasma and tissues. In CTX, the lack of CDCA disrupts the normal on bile acid synthesis via the farnesoid X receptor (FXR), promoting excessive production of abnormal bile alcohols and cholestanol through pathways, including defective 26-hydroxylation. CDCA therapy restores the deficient bile acid, activates FXR to downregulate cholesterol 7α-hydroxylase (CYP7A1), suppresses the overproduction of toxic bile alcohols, and reduces cholestanol accumulation. CDCA received designation in the United States in 2007 and in the in 2014, with EU approval for CTX treatment in 2017 and US approval as chenodiol (Ctexli) in 2025. Therapeutically, CDCA normalizes bile acid synthesis, reduces plasma and tissue cholestanol levels by 70-90%, and alleviates neurological and extraneurological symptoms in most patients. It halts disease progression, shrinks xanthomas, and improves , , , and cerebellar function, with benefits emerging over months to years of lifelong therapy. The recommended adult dose is 750 mg/day orally (250 mg three times daily), with adjustments for children based on body weight (typically 5-15 mg/kg/day divided into three doses); treatment is lifelong to maintain suppression of abnormal metabolites. Pivotal evidence from a 1984 study of 17 CTX patients showed that CDCA (750 mg/day) reduced mean cholestanol threefold, cleared in 10 patients, resolved in 6 of 7, and improved cerebellar and in most, arresting or reversing progression. Long-term follow-ups confirm sustained efficacy: a 2025 nationwide study of 86 patients reported 100% stabilization or improvement in those starting before age 28, with reduced cholestanol in all and halted progression of pyramidal and cerebellar symptoms, though psychiatric features like were less responsive. Monitoring involves regular cholestanol levels as a of efficacy (target: normalization) and MRI to assess lesions, alongside .

Other established applications

Chenodeoxycholic acid is authorized by the for the treatment of cerebrotendinous xanthomatosis (CTX), an inborn error of primary synthesis due to sterol 27-hydroxylase deficiency. It has also shown in other rare cholestatic disorders characterized by defective production, such as oxysterol 7α-hydroxylase deficiency, a condition presenting with progressive intrahepatic , neurological impairment, and liver dysfunction in infancy. Oral chenodeoxycholic acid therapy has demonstrated in normalizing profiles, resolving , and improving clinical outcomes, including liver levels and growth parameters, when initiated early. Doses typically range from 10 to 15 mg/kg/day, administered orally, with monitoring of to ensure safety. Historical investigations explored chenodeoxycholic acid for based on its potential to induce cholesterol 7α-hydroxylase, the rate-limiting enzyme in bile acid synthesis, thereby promoting conversion to bile acids and reducing levels. However, clinical studies showed limited efficacy in lowering serum LDL , with some reports indicating no change or even paradoxical increases due to reduced LDL clearance, leading to its diminished use in favor of more effective lipid-lowering agents. Due to risks of , including , chenodeoxycholic acid is contraindicated in patients with pre-existing or acute , overlapping with precautions in cholestatic applications where baseline liver assessments are required.

Pharmacology and safety

Mechanism of action

Chenodeoxycholic acid (CDCA) primarily exerts its pharmacological effects through activation of the farnesoid X receptor (FXR), a that serves as a key regulator of . Upon binding to FXR in hepatocytes and enterocytes, CDCA induces the transcriptional upregulation of the small heterodimer partner (SHP), a corepressor that inhibits liver receptor homolog-1 (LRH-1)-mediated activation of the cholesterol 7α-hydroxylase gene (CYP7A1). This repression of CYP7A1, the rate-limiting in the classic biosynthetic pathway, results in decreased hepatic synthesis and prevents bile acid overload. In addition to FXR, CDCA acts as an agonist for the Takeda G-protein-coupled receptor 5 (TGR5, also known as GPBAR-1), a membrane-bound receptor expressed on enteroendocrine L-cells, macrophages, and adipocytes. TGR5 activation by CDCA elevates intracellular (cAMP) levels via G-protein signaling, which in L-cells promotes the secretion of (GLP-1) to enhance insulin sensitivity and glucose homeostasis. In , this cAMP-mediated pathway stimulates type 2 (DIO2) expression, increasing local hormone production and thereby promoting and energy expenditure. CDCA also modulates cholesterol metabolism by inhibiting the enzyme 3-hydroxy-3-methylglutaryl-coenzyme A reductase () and the intestinal cholesterol transporter Niemann-Pick C1-like 1 (NPC1L1), primarily through FXR-dependent mechanisms. FXR activation in the liver represses sterol regulatory element-binding protein-2 (SREBP-2), which downregulates expression and reduces de novo synthesis, while in the intestine, FXR induces 19 (FGF19) secretion to suppress NPC1L1-mediated absorption. Furthermore, CDCA exhibits properties by suppressing the nuclear factor kappa B () pathway in hepatocytes and macrophages; FXR activation antagonizes translocation and transcriptional activity, reducing pro-inflammatory cytokine production such as tumor necrosis factor-α (TNF-α) and interleukin-6 (IL-6), while TGR5 contributes by inhibiting and c-Jun N-terminal kinase (JNK) signaling in immune cells. Compared to (UDCA), CDCA demonstrates greater hydrophobicity, which enhances its potency in altering composition by more effectively reducing biliary saturation and forming less toxic micelles at therapeutic doses. At pharmacological concentrations, which exceed physiological levels, CDCA amplifies FXR and TGR5 signaling pathways without the full engagement of endogenous negative feedback mechanisms, such as those mediated by FGF19, leading to sustained metabolic modulation.

Pharmacokinetics

Chenodeoxycholic acid (CDCA) is administered orally and undergoes efficient absorption primarily in the proximal through a combination of passive diffusion and via the apical sodium-dependent bile acid transporter (ASBT). is nearly complete, ranging from 80% to 99% depending on the dose and site of infusion, with peak concentrations achieved 1 to 2 hours after . Food intake delays but does not significantly alter overall . The systemic bioavailability of CDCA is approximately 70-80%, limited by extensive first-pass extraction in the liver following absorption. Once absorbed, CDCA enters the portal circulation and is efficiently taken up by hepatocytes, contributing to its confinement within the enterohepatic system. CDCA exhibits high plasma protein binding, primarily to albumin at levels exceeding 95%, which facilitates its transport. Its distribution is largely restricted to the enterohepatic circulation, involving the intestine, portal vein, liver, and biliary tract, with minimal entry into systemic circulation. The endogenous pool size of CDCA in healthy individuals is typically 1-2 g, part of the total bile acid pool of 2-4 g, maintained through efficient reabsorption. Metabolism of CDCA occurs primarily in the liver, where it undergoes conjugation with glycine or taurine to form more soluble bile salts, with a typical glycine:taurine ratio of about 3:1 in humans. In the intestine, gut bacteria perform 7-dehydroxylation to form lithocholic acid, a secondary bile acid and potentially toxic metabolite. Elimination of CDCA is predominantly fecal, with 80-90% excreted unchanged or as bacterial derivatives like lithocholic acid, while a small (about 5%) enters the systemic circulation and undergoes minor urinary . Due to extensive enterohepatic recirculation, the biological half-life of CDCA is 3-5 days, reflecting the slow turnover of the pool. In special populations, may vary. Liver impairment reduces hepatic uptake and biliary secretion of CDCA, leading to decreased clearance and potential accumulation in . In pediatrics, particularly for conditions like cerebrotendinous xanthomatosis, dosing is adjusted based on body weight (e.g., 15 mg/kg/day), though specific pharmacokinetic alterations are not well-characterized and require monitoring.

Adverse effects and contraindications

Chenodeoxycholic acid therapy is commonly associated with gastrointestinal adverse effects, primarily occurring in 30-40% of patients due to in the colon, along with , , , cramps, and dyspepsia. These effects are typically dose-related and manageable with dose reduction, though approximately 3% of patients discontinue treatment due to uncontrolled . Hepatic adverse effects include transient elevations in serum aminotransferases in 20-30% of patients, often appearing within the first few months of and resolving with dose adjustment or discontinuation; rare cases of clinically apparent have been reported, particularly in early dissolution trials. A portion of this arises from bacterial conversion of chenodeoxycholic acid to lithocholic acid in the gut, which can accumulate and cause , though levels remain minor in humans and the risk is mitigated by co-administration of , which reduces lithocholic acid formation. Other reported adverse effects include and pruritus, as well as dose-related in some patients, particularly at lower doses. In early clinical trials for dissolution, overall discontinuation rates due to side effects ranged from 18-30%, though long-term data indicate that most hepatic changes are reversible upon cessation of therapy. Contraindications for chenodeoxycholic acid include known hepatobiliary disorders such as biliary obstruction, , or intrahepatic ; non-functioning ; radiopaque or bile pigment s; complications from s requiring surgery; and to acids. It is also contraindicated in (FDA Category X) due to evidence of fetal harm in at high doses, though human data are limited. Monitoring recommendations include monthly liver function tests (aminotransferases, , ) for the first three months, followed by assessments every three months thereafter, with discontinuation advised if aminotransferases exceed three times the upper limit of normal or if rises above twice normal. Periodic imaging, such as cholecystograms or ultrasonograms every 6-9 months, is also suggested to evaluate response and detect complications.

Research developments

Role in cancer

Chenodeoxycholic acid (CDCA), a primary characterized by its hydrophobic properties, promotes () development by inducing DNA damage in colon epithelial s, as evidenced by its ability to trigger unscheduled indicative of repair responses in colon cell lines. Furthermore, CDCA upregulates (COX-2) expression, contributing to inflammatory processes that foster tumorigenesis in the colon. In cells harboring APC mutations, CDCA activates β-catenin signaling, enhancing and leading to increased tumor formation in the intestinal tract of animal models. Epidemiological studies have linked elevated fecal CDCA levels to higher risk, with significantly greater excretion observed in patients with large bowel cancer compared to controls (weighted mean difference of 0.28 mg/g dry feces). In (HCC), CDCA exhibits a synergistic effect with , enhancing its antitumor efficacy through FXR-mediated pathways that promote ; a 2022 preclinical study demonstrated a twofold increase in treatment efficacy in HepG2 cell models and xenografts when CDCA was combined with . This combination inhibits proliferation, migration, and invasion more effectively than alone by modulating the PI3K/AKT/ signaling pathway, underscoring CDCA's potential as an adjunct in HCC . CDCA also holds anticancer potential, particularly in , where it inhibits progression via FXR activation; administration of CDCA as an FXR agonist attenuated pancreatic intraepithelial neoplasia (PanIN) development in Kras-mutant models. A 2023 metabolomics analysis further identified CDCA levels as predictive of reduced pancreatic necrosis, with higher recovery-phase concentrations correlating with less tissue damage in , a precursor to pancreatic ductal . Through modulation, CDCA reduces the production of secondary carcinogens like (DCA), which promotes and , while CDCA itself suppresses tumor growth in certain contexts by altering microbial composition to favor metabolites. This dual role highlights CDCA's context-dependent influence on via interactions. Preclinical evidence supports CDCA's delivery via nanoformulations for targeted HCC therapy, with lipoprotein-mimicking nanoparticles incorporating CDCA showing enhanced tumor accumulation and efficacy in models. Therapeutic trials exploring CDCA derivatives as FXR agonists, such as , are ongoing, including phase II evaluations as adjuncts in HCC to improve outcomes in advanced disease.

Emerging therapeutic applications

Chenodeoxycholic acid (CDCA) has shown promise in preclinical models of acute injury () by activating the farnesoid X receptor (FXR) to exert effects. In lipopolysaccharide-induced mouse models, CDCA treatment significantly reduced pathological damage and lowered levels of pro-inflammatory cytokines such as interleukin-6 (IL-6) and tumor factor-alpha (TNF-α), highlighting its potential to mitigate inflammation through FXR-mediated pathways. In disorders, particularly non-alcoholic (NAFLD), CDCA regulates via multi-omics approaches, demonstrating therapeutic effects on and . A 2025 study using transcriptomics, , and revealed that CDCA supplementation in high-fat diet-induced NAFLD models lowered levels and improved profiles by modulating key pathways in hepatic lipid synthesis and oxidation. For obesity management, nanoparticle-based delivery of CDCA promotes browning and enhances fat burning. A single subcutaneous injection of CDCA-loaded nanoparticles in obese models reduced size and fat mass by inducing mitochondrial function and in , offering a targeted approach to combat without systemic side effects. CDCA has been identified as a potential therapeutic for pancreatic in through bile acid profiling. In cerulein-induced models, CDCA levels decreased during the acute phase but rose in recovery, with supplementation improving outcomes by reducing acinar cell and , suggesting its role in modulating signaling to protect pancreatic tissue. In type 1 diabetes, CDCA enhances the viability of islet cells for encapsulation and transplantation by providing anti-apoptotic protection. Preclinical studies demonstrated that incorporating CDCA into alginate-based microencapsulation matrices preserved primary islet function and reduced apoptosis during in vitro culture and transplantation, potentially improving graft survival and insulin production in diabetic models. Beyond these applications, CDCA holds potential in metabolic syndrome through activation of the Takeda G protein-coupled receptor 5 (TGR5), which regulates energy expenditure and glucose homeostasis. A 2024 review emphasized TGR5 agonism by CDCA as a mechanism to increase thermogenesis and insulin sensitivity, addressing interconnected features of metabolic syndrome such as dyslipidemia and hyperglycemia in preclinical settings.