Copper pheasant
The Copper pheasant (Syrmaticus soemmerringii) is a medium-sized bird in the family Phasianidae, endemic to Japan and renowned for the males' distinctive reddish-chestnut head, neck, and breast plumage, along with a long, cinnamon-colored tail featuring black bars that can measure up to 98 cm in length.[1] Males typically reach 87.5–136 cm in total length and weigh 943–1,348 g, while females are smaller at 51–54 cm (with tails of 14–19 cm) and 745–1,000 g, displaying grey-brown upperparts with a darker breast band and a shorter, blunter tail.[1] This species exhibits sexual dimorphism and geographical variation across five subspecies, such as S. s. scintillans on northern Honshu and S. s. intermedius on Shikoku.[1] Native to the forested hills and mountains of Honshu, Shikoku, and Kyushu islands, the copper pheasant has been introduced to Sado and Oki Islands, with an extent of occurrence spanning approximately 510,000 km².[2] It inhabits a range of woodland types below 1,800 m elevation, including broadleaf, deciduous, coniferous, and mixed forests, as well as plantations and areas near villages, favoring dense undergrowth for foraging and cover while roosting in trees at night.[2][1] The bird's diet consists primarily of invertebrates, seeds, and plant matter, and it is known for its elusive behavior, often remaining hidden in thick vegetation.[1] Historically abundant, the copper pheasant faced severe declines from excessive hunting, with estimates of 500,000–1,000,000 birds harvested annually between 1925 and 1975, alongside habitat loss due to deforestation.[2] Predation by native species like raccoon dogs, foxes, and raptors, as well as disturbance from herbivores and omnivores, pose ongoing risks, though protective measures such as the 1976 ban on hunting females have aided recovery.[2][1] Current population estimates range from 50,000 to 300,000 mature individuals, with an unknown trend, leading to its classification as Least Concern on the IUCN Red List as of 2025.[2] Conservation efforts emphasize habitat management, monitoring, and sustainable practices to ensure the species' persistence in its island homeland.[2]Systematics
Taxonomy
The copper pheasant was first described in 1830 by Dutch zoologist Coenraad Jacob Temminck as Phasianus soemmerringii in his Nouveau recueil de planches coloriées d'oiseaux, honoring the German anatomist Samuel Thomas von Sömmerring for his contributions to natural history.[3] The species is classified in the family Phasianidae within the order Galliformes, with the currently accepted binomial name Syrmaticus soemmerringii.[1] Following its initial placement in the genus Phasianus, the copper pheasant was moved to the monotypic genus Graphephasianus in 1852 by Heinrich Gustav Reichenbach, based on its distinctive long-tailed morphology.[4] In 1918, American ornithologist William Beebe reclassified it into the genus Syrmaticus in his comprehensive Monograph of the Pheasants, expanding the genus from its prior single species (S. reevesii) to include the copper pheasant and others sharing traits such as elongated central tail feathers and a prominent crest, despite some plumage differences among males.[5] This placement has been upheld in modern taxonomy, though historical disputes persisted, including brief considerations of retaining it in Phasianus due to reported hybridization with the green pheasant (Phasianus versicolor), which produces infertile offspring.[1] Phylogenetically, the copper pheasant represents the basal divergence within the genus Syrmaticus, followed by Reeves's pheasant (S. reevesii), with the remaining species including Elliot's pheasant (S. ellioti) and the Mikado pheasant (S. mikado) forming a sister clade, as confirmed by analyses of the mitochondrial cytochrome b gene.[1][6] These genetic studies support the monophyly of Syrmaticus and highlight its distinction from Phasianus based on molecular divergences.Subspecies
The Copper pheasant (Syrmaticus soemmerringii) is recognized as comprising five subspecies, distinguished primarily by variations in male plumage coloration, tail length, and feather fringes, with differences arising from geographic isolation across the Japanese islands.[1] These subspecies exhibit clinal variation, with plumage generally becoming darker toward the south, though some authorities suggest that certain races, such as intermedius and subrufus, may be synonymous with scintillans due to overlapping traits, and further studies in contact zones are recommended to clarify boundaries.[1] Currently, the taxonomy remains stable without evidence from molecular data supporting additional subdivisions.[1] The following table summarizes the recognized subspecies, their distributions, and key morphological distinctions:| Subspecies | Distribution | Distinguishing Features |
|---|---|---|
| S. s. scintillans | Northern and central Honshū | Males paler overall with shorter tails; proposed northern (septentrionalis) and southwestern (inabaensis) variants included herein.[1] |
| S. s. intermedius | Southwestern Honshū and Shikoku | Males with longer, redder tails; richer red tones and golden fringes on scapulars and mantle.[1] |
| S. s. subrufus | Southeastern and southwestern Honshū, southwestern Shikoku | Males darker red with minimal white in upperparts and golden-orange fringes.[1] |
| S. s. soemmerringii | Northern and central Kyūshū | Darkest race overall, with golden-carmine fringes; dark red-brown plumage and black tail bars (nominate form).[1] |
| S. s. ijimae | Southeastern Kyūshū | Similar to soemmerringii but with broad white fringes on back and rump feathers; darker plumage.[1] |