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Middle Stone Age

The Middle Stone Age (MSA) is an archaeological period in African prehistory, spanning approximately 300,000 to 30,000 years ago, during which early Homo sapiens populations produced more sophisticated stone tools and exhibited the initial evidence of modern human behaviors, bridging the Earlier Stone Age and the Later Stone Age.^[1] This era is primarily documented across sub-Saharan Africa, with key assemblages reflecting technological advancements, resource exploitation strategies, and cultural innovations tied to the emergence of anatomically modern humans around 300,000 years ago.^[2] Central to the MSA are refined lithic technologies, including prepared-core methods like the Levallois technique, which enabled the production of predetermined flakes and points for hafting onto spears or other composite tools, marking a shift from the larger, less specialized Acheulean handaxes of earlier periods.^[3] Other innovations encompassed heat treatment of silcrete to enhance flaking quality, the use of bone and antler for tools such as awls and points, and diverse unifacial tools like scrapers and denticulates for processing hides, wood, and marine resources.^[4] These developments suggest improved planning, skill, and adaptability, often linked to environmental pressures such as climate fluctuations during Marine Isotope Stages 8–4.^[5] The MSA is particularly renowned for providing the earliest archaeological evidence of behavioral modernity in Homo sapiens, including symbolic practices like ochre processing—possibly for body decoration or ritual—and the creation of perforated shell beads for personal adornment, indicating social signaling and aesthetic awareness as early as 75,000 years ago.^[6] Additional markers include long-distance exchange of raw materials, such as obsidian transported over hundreds of kilometers, and systematic exploitation of coastal shellfish, reflecting expanded foraging ranges and cognitive flexibility.^[1] While regional variability exists— with distinct industries like the Still Bay (characterized by bifacial points) and Howiesons Poort (featuring backed geometric tools)—the MSA overall underscores a gradual buildup of cultural complexity rather than a singular "human revolution."^[3] Prominent MSA sites illuminate these traits, including Blombos Cave in South Africa's Western Cape, where layers from 100,000–70,000 years ago yielded engraved ochre pieces, shell beads, and ochre-processing kits, offering direct proof of abstract thinking and artistic expression.^[6] Similarly, Pinnacle Point Site 5-6 on the southern Cape coast preserves evidence of heat-treated tools and intensive shellfish gathering from 164,000 years ago, highlighting early maritime adaptations during arid phases.^[7] Other significant locales, such as Klasies River Mouth and Sibudu Cave, reveal hafted bone tools and possible projectile technologies, further demonstrating the diverse and innovative lifeways of MSA populations across the continent.^[8]

Definition and Chronology

Temporal and Spatial Extent

The Middle Stone Age (MSA) is broadly dated from approximately 300,000 to 30,000–50,000 years ago, representing a key period in African prehistory associated with the emergence of early modern human technologies and behaviors.^[5] This timeframe is anchored by archaeological evidence from sites such as Jebel Irhoud in Morocco, where Levallois prepared-core reduction techniques— a hallmark of MSA lithic innovation—appear around 315,000 years ago, marking the onset of the period. Regional variations exist, with some South African assemblages, like those at Florisbad, indicating an earlier start near 280,000 years ago, potentially reflecting localized technological transitions from the preceding Acheulean. The end of the MSA is variably defined by the gradual or abrupt shift to Later Stone Age (LSA) industries, characterized by the increasing dominance of microliths and backed tools, often dated between 50,000 and 30,000 years ago across different regions.^[1] Geographically, the MSA is primarily distributed across sub-Saharan Africa, with dense concentrations of sites in eastern and southern regions, including the East African Rift Valley (e.g., Kenya, Tanzania, Ethiopia) and southern Cape landscapes (e.g., South Africa). It extends northward into North Africa, encompassing the Maghreb and Sahara margins, where industries like the Aterian (dated ~145,000–20,000 years ago) show MSA affinities with Levallois elements and hafted tools. Limited evidence appears on the Eurasian fringes, such as in the Levant, but these assemblages are distinguished from the contemporaneous Eurasian Middle Paleolithic by their African technological signatures and lack of integration with Neanderthal-associated industries. This chronological span overlaps significantly with the emergence of anatomically modern Homo sapiens in Africa around 300,000 years ago, as evidenced by fossils from Jebel Irhoud co-occurring with early MSA artifacts, suggesting a close association between hominin evolution and the period's technological developments. Such temporal and spatial parameters provide the foundational framework for understanding MSA variability, influenced by environmental fluctuations like glacial-interglacial cycles that affected site preservation and human adaptations across the continent.^[9]

Terminology and Debates

The term "Middle Stone Age" (MSA) was coined by archaeologist Astley John Hilary Goodwin in 1928 to describe stone tool assemblages from southern Africa that exhibited technological advancements beyond the Earlier Stone Age but lacked the microlithic focus of the Later Stone Age.^[10] Goodwin's framework, introduced in his paper "An Introduction to the Middle Stone Age in South Africa," emphasized flake-based tools and prepared core techniques as hallmarks of this period, initially applied to regional sites before broader continental adoption.^[11] Scholars debate whether the MSA represents a unified "package" of technological and behavioral traits or a regionally variable phenomenon shaped by local environmental and cultural dynamics. While early conceptualizations treated the MSA as a coherent entity spanning much of Africa, recent analyses highlight significant variability, particularly in eastern and central regions, where assemblages often deviate from a "generic" MSA lacking defining characteristics.^[10] In 2024, critiques, such as those by Shea, argued against its use as a formal taxonomic category, noting its absence in approximately 50% of eastern African sites and advocating for more hypothesis-driven approaches to account for African diversity rather than imposing a homogenized label.^[10] The MSA is distinct from the Eurasian Middle Paleolithic, which encompasses Neanderthal-associated industries across Europe and Asia, whereas the MSA is uniquely African and tied to the emergence of Homo sapiens with region-specific behavioral innovations like symbolic material use.^[12] This terminological separation underscores African-centric developments, avoiding direct equivalence despite superficial lithic similarities such as Levallois techniques.^[12] Contemporary research trends favor subdividing the MSA into phases—early, middle, and late— to better capture technological refinements and responses to environmental fluctuations, rather than a monolithic period.^[12] These divisions reflect shifts in tool production and resource exploitation, informed by improved chronologies and regional syntheses that highlight intra-MSA evolution.^[13]

Technological Developments

Transition from Acheulean

The transition from the Acheulean to the Middle Stone Age (MSA) represents a gradual technological shift, characterized by the replacement of large bifacial handaxes and cleavers with more standardized flake-based tools, occurring primarily between approximately 500,000 and 250,000 years ago across Africa.^[11] This change was not abrupt but involved hybrid assemblages that blended Acheulean elements with emerging MSA technologies, as evidenced at sites like Kathu Pan in South Africa, where Fauresmith industry layers dated to 511,000–435,000 years ago contain both large cutting tools and early prepared-core flakes.^[14] Similarly, the Kapthurin Formation in Kenya preserves interstratified Acheulean and MSA artifacts, illustrating overlapping traditions during this period.^[15] A key marker of this transition was the introduction of prepared-core reduction strategies, notably Levallois and discoidal methods, which enabled the production of predetermined flake shapes for points and blades, contrasting with the less controlled flaking of Acheulean bifaces.^[16] Levallois technology, involving hierarchical core preparation to detach flakes of specific morphology, first appears in late Acheulean contexts around 500,000 years ago in East Africa, while discoidal reduction—centripetal flaking around a disc-shaped core—facilitated higher yields of usable blanks and became prominent in transitional assemblages.^[11] These innovations marked a departure from the hand-held, heavy-duty tools of the Acheulean toward lighter, potentially haftable implements suited to diverse functions.^[14] Several factors likely drove this technological evolution, including adaptations to raw material efficiency and enhancements in hominin cognition. Prepared-core techniques like Levallois maximized flake output from limited nodules, promoting economical use of scarce high-quality stone in varied environments, as seen in the finer-grained silcrete tools of transitional sites.^[17] Cognitively, the shift required advanced planning, foresight, and motor skill refinement, correlating with the emergence of archaic Homo sapiens populations capable of more complex problem-solving by around 300,000 years ago.^[14] Regionally, the transition exhibited temporal variation, with earlier evidence in East Africa—such as the Olorgesailie basin, where MSA points and small tools replace Acheulean handaxes after 499,000 years ago—compared to a later onset in South Africa, where full MSA assemblages postdate 300,000 years ago at sites like Florisbad.^[11] This pattern suggests asynchronous developments influenced by local ecological pressures and population dynamics, though the Fauresmith in southern Africa represents an early bridge around 400,000 years ago.^[17]

Lithic Technologies

The lithic technologies of the Middle Stone Age represent a significant advancement in prepared core reduction, emphasizing efficiency and predetermination in blank production over the simpler methods of the preceding Acheulean. Central to these toolkits is the Levallois technique, which dominates MSA assemblages across Africa and involves hierarchical core preparation to detach flakes of predetermined morphology, such as triangular or oval shapes with flat ventral surfaces and convergent dorsal edges.^[16] This method originated in Africa by around 300,000 years ago, predating its appearance in Eurasia, and reflects enhanced planning in knapping sequences.^[18] Levallois variants include the preferential mode, where core preparation focuses on isolating a single high-quality flake from a targeted striking platform, and the recurrent mode, which permits multiple flake removals from the same prepared surface before exhaustion, allowing sustained production of standardized blanks.^[19] Complementing Levallois are discoidal reduction strategies, involving opportunistic flaking from disc-shaped cores via centripetal or bidirectional patterns to yield irregular but versatile flakes suitable for retouching into scrapers or points.^[20] Blade and laminar technologies also emerge prominently, producing elongated blanks at least twice as long as wide through prismatic core reduction, with systematic blade production documented as early as approximately 300,000 years ago in South African sites, as confirmed by recent analyses of core morphology and scar patterns.^[3] Regional variations highlight technological diversity within the MSA, particularly in southern Africa. The Still Bay industry, dated to about 77,000–72,000 years ago, features finely bifacial foliate points crafted via pressure flaking and marginal retouch, often on silcrete or quartzite, as seen in assemblages from Blombos Cave and Hollow Rock Shelter.^[21] Similarly, the Howiesons Poort techno-complex, around 65,000–59,000 years ago, is marked by small blades and bladelets, including backed segments and unifacial points, produced from laminar cores and indicative of specialized reduction for composite tools.^[22] These variations underscore adaptations to local raw materials and subsistence needs, with northern and eastern African MSA sites showing more generalized Levallois and discoidal emphases.^[12] Technological complexity in MSA lithics is evidenced by increased flake standardization, with metrics such as reduced coefficients of variation in length-to-width ratios (often below 20% in Levallois products) demonstrating greater control over blank morphology compared to earlier periods.^[23] Hafting practices further illustrate sophistication, with microscopic residues of plant resins, beeswax, and red ochre on tool edges from sites like Sibudu Cave indicating adhesive attachment to handles as early as 70,000 years ago, enabling multifunctional composite implements.^[24]

Non-Lithic Artifacts and Materials

The use of ochre as a non-lithic material in the Middle Stone Age marks a significant expansion in technological repertoires, with evidence of processing for pigment production and grinding dating to approximately 100,000 years ago at Pinnacle Point Cave 13B in South Africa. Over 500 ochre pieces, primarily hematite, have been recovered from layers spanning 164,000 to 92,000 years ago at this site, many exhibiting striations, scoring, and faceting consistent with deliberate grinding to produce powder, likely for body decoration, hide treatment, or ritual purposes. Recent findings also indicate ochre was deliberately modified into lithic retouchers, showing use-wear patterns and intentional shaping for stone tool production in MSA assemblages.^[25] Such modifications indicate systematic exploitation and manipulation of iron-rich minerals, distinguishing MSA practices from earlier, sporadic occurrences in the Acheulean.^[26] Bone tools represent another key category of non-lithic artifacts in the MSA, emerging around 90,000 years ago in North Africa's Aterian techno-complex, as demonstrated by a finely pointed bone implement from Dar es-Soltan 1 cave in Morocco, shaped through scraping and smoothing for possible use as a projectile or awl.^[27] In southern Africa, bone points and awls from Sibudu Cave's Howiesons Poort layers (~65,000 years ago) show advanced features, including hafting traces and intentional heat treatment via slow heating in ash to harden the material and improve workability.^[28] These tools, often made from long bone fragments of large mammals, reflect specialized manufacturing and integration with lithic components, though their primary fabrication relied on organic media.^[28] Shell beads and associated marine material processing further illustrate MSA innovations in non-lithic technologies, with the earliest examples dated to ~75,000 years ago at Blombos Cave, South Africa, where over 40 Nassarius kraussianus shells were perforated, polished, and strung as ornaments.^[29] These beads, sourced from coastal environments up to 20 km inland, bear use-wear from suspension and ochre staining, pointing to symbolic use and deliberate exploitation of marine shells for both decorative and possibly adhesive purposes.^[29] Concurrent evidence from nearby coastal sites underscores adaptations involving shell collection and processing, expanding material culture beyond terrestrial resources.^[29] Composite tools incorporating adhesives from plant resins appear in the MSA record at Sibudu Cave, South Africa, with findings from layers dated to ~70,000 years ago revealing stone segments hafted using mixtures of plant gum and red ochre as binding agents.^[24] Experimental replications confirm that these adhesives required heating and mixing to achieve durability, enabling the attachment of blades to wooden hafts for enhanced functionality in hunting or processing tasks.^[24] Residue analysis on tools from these contexts identifies acacia-like gums combined with mineral additives, highlighting sophisticated knowledge of organic chemistry and multi-component tool assembly.^[24]

Hominin Biology and Migration

Associated Hominin Species

The Middle Stone Age (MSA) is primarily associated with anatomically modern Homo sapiens, whose earliest fossil evidence appears in African contexts dating to approximately 315,000 years ago. Fossils from Jebel Irhoud in Morocco, including a partial cranium, face, and other skeletal elements, exhibit a mosaic of modern human traits—such as a globular braincase—and more archaic features, like a prognathic face and elongated cranial vault, marking an early stage in the evolutionary emergence of H. sapiens. These remains, found alongside MSA lithic artifacts, indicate that modern human morphology was developing across Africa during the initial phases of the MSA.^[30] Key fossil distributions further link H. sapiens to the MSA, particularly in eastern Africa. The Omo Kibish site in Ethiopia yielded the Omo I partial skeleton, dated to approximately 233,000 years ago, which displays fully modern human cranial and postcranial morphology, representing one of the earliest well-dated examples of anatomically modern humans. Similarly, the Herto Bouri site in Ethiopia produced three crania, including an adult male and female, dated to approximately 160,000 years ago; these fossils show advanced H. sapiens features, such as a high forehead and rounded occipital, while retaining some robusticity, and are associated with MSA tools. These eastern African specimens underscore the widespread presence of early H. sapiens populations during the middle phases of the MSA. While H. sapiens dominates MSA fossil records, possible contributions from archaic hominin forms have been proposed for early MSA contexts, based on anatomical overlaps and chronological proximity. In Africa, fossils attributed to Homo heidelbergensis—such as those from Bodo (Ethiopia, ~600,000 years ago) and Elandsfontein (South Africa, ~1.0–0.6 million years ago)—exhibit traits transitional to modern humans, suggesting potential ancestral roles or regional persistence. Likewise, Homo naledi, known from the Rising Star Cave in South Africa and dated to 236,000–335,000 years ago, represents a small-brained, primitive hominin contemporaneous with early MSA sites, raising questions about its ecological overlap or indirect influence on H. sapiens populations in southern Africa, though no direct artifact associations exist for H. naledi.^[31]^[32] Debates persist regarding the exclusivity of H. sapiens in the MSA, particularly concerning species boundaries and interbreeding with archaic forms. Some researchers argue that early MSA hominins reflect a polytypic H. sapiens clade incorporating H. heidelbergensis-like variation, while others propose distinct lineages with gene flow. Genetic evidence from modern African populations reveals archaic admixture—up to 2–19% from unidentified "ghost" lineages—potentially tracing to MSA-era interbreeding events with non-sapiens hominins in Africa, complicating taxonomic attributions and highlighting a reticulate evolutionary history.

Cranial and Brain Evolution

During the Middle Stone Age (MSA), hominin endocranial volumes exhibited a gradual increase, reflecting ongoing neuroanatomical development in early Homo sapiens. Fossils from early MSA sites, such as Jebel Irhoud in Morocco dated to approximately 315,000 years ago, yield endocast-based estimates of 1,375 cm³ for Irhoud 1 and 1,467 cm³ for Irhoud 2, aligning with the lower end of modern human ranges but surpassing many archaic forms.^[33] By the middle MSA, around 233,000 years ago, specimens like Omo II from Ethiopia show volumes reaching 1,491 cm³, approaching the modern average of ~1,450 cm³ and indicating a modest expansion of roughly 100–200 cm³ over the period.^[33] These measurements, derived from virtual endocasts of fossil crania, underscore a trend toward larger brain capacities without dramatic leaps, consistent with the broader trajectory of hominin encephalization.^[34] A key feature of MSA cranial evolution was the internal rearrangement of brain lobes, particularly the relative expansion of parietal and temporal regions, which enlarged areas for sensory integration. Endocasts from early MSA individuals, such as those at Jebel Irhoud, display elongated brain shapes with less pronounced parietal bulging compared to later forms, suggesting progressive reorganization toward greater globularity.^[35] This shift involved disproportionate growth in the parietal lobe, including the precuneus, and temporal association zones, as evidenced by comparisons of fossil endocasts with modern human brains using geometric morphometrics.^[33] The temporoparietal junction and posterior superior temporal sulcus also underwent expansion and rearrangement during this era, enhancing cortical connectivity in regions linked to multimodal sensory processing.^[36] Fossils like the Broken Hill 1 (Kabwe) cranium from Zambia, dated to ~300,000 years ago and associated with early MSA contexts in Africa, exemplify transitional cranial robusticity. This specimen features a robust brow ridge, broad face, and thick vault alongside an endocranial volume of ~1,280 cm³, bridging archaic Homo heidelbergensis-like traits with emerging H. sapiens morphology.^[37] Such intermediates highlight morphological variation among MSA hominins, with robusticity decreasing over time as brain expansion and reorganization progressed.^[31] Estimating these changes relies on advanced methodologies, including virtual reconstructions of incomplete crania via computed tomography (CT) scans to produce digital endocasts. These endocasts enable quantification of brain volume and shape through landmark-based geometric morphometrics, allowing estimation of missing regions by mirroring bilateral symmetries or referencing complete conspecific skulls.^[38] For instance, 3D warping and thin-plate spline interpolation facilitate the reconstruction of distorted fossils, providing insights into lobe proportions and regional expansions without physical alteration of specimens.^[39] This approach has been pivotal in analyzing MSA fossils, revealing subtle reorganizations that distinguish early from late period brains.^[35]

Genetic and Population Evidence

Genomic analyses of modern African populations reveal deep divergent lineages within Homo sapiens that trace back to approximately 200,000 years ago, supporting the emergence of anatomically modern humans in Africa during the Middle Stone Age (MSA). These lineages, identified through whole-genome sequencing, exhibit the highest levels of genetic diversity among human populations globally, reflecting long-term isolation and regional adaptations without significant input from non-sapiens archaic hominins. For instance, mitochondrial DNA and autosomal markers in southern and eastern African groups, such as the San and Hadza, show basal branches that predate non-African lineages by tens of thousands of years, indicating structured metapopulations across the continent. Evidence for low levels of archaic admixture in African sapiens genomes further underscores these deep indigenous lineages. Unlike non-African populations, which carry 1-2% Neanderthal DNA from dispersals out of Africa, sub-Saharan African genomes generally show minimal to no Neanderthal or Denisovan ancestry, with archaic contributions estimated at less than 0.5% on average. However, statistical methods like S* have detected signals of "ghost" archaic introgression—likely from an unknown hominin species related to Homo heidelbergensis or earlier forms—in up to 8% of West African genomes, such as in Yoruba and Mende populations, suggesting rare, localized gene flow events predating the MSA. This admixture is low overall and does not substantially alter the predominantly sapiens character of African genetic diversity.^[40] Population genomic models indicate severe bottlenecks in early hominin ancestors between approximately 930,000 and 813,000 years ago, reducing the breeding population to around 1,280 individuals for about 117,000 years, followed by gradual recoveries and expansions. These expansions, inferred from linkage disequilibrium and site frequency spectra in modern genomes, align temporally with MSA onset around 300,000 years ago, coinciding with technological innovations like Levallois flaking and hafted tools in eastern and southern Africa. Demographic simulations suggest that post-bottleneck growth in effective population size from ~10,000 to over 100,000 individuals by 200,000-100,000 years ago facilitated the spread of sapiens across diverse African environments, enhancing genetic diversity.^[41] Phylogeographic reconstructions using coalescent-based models and ancient DNA proxies point to multiple out-of-Africa dispersals during the late MSA, with a primary pulse between 70,000 and 50,000 years ago originating from eastern African source populations. These models, calibrated with mutation rates from modern and archaic genomes, depict a southern coastal route along the Indian Ocean, carrying lineages that founded non-African populations while leaving traces of back-migration in African genomes. The timing correlates with MSA cultural complexes like the Howiesons Poort in South Africa, suggesting demographic pulses driven by climatic amelioration enabled rapid colonization.^[42] Recent genomic studies, including reanalyses of Ethiopian ancient DNA, highlight regional diversity and ghost lineages within Africa. The ~4,500-year-old Mota genome from Ethiopia, one of the earliest successfully sequenced from the continent, reveals a basal East African lineage with no Eurasian admixture, underscoring pre-MSA isolation and subsequent regional structuring. A 2025 study suggests modern humans result from admixture between two ancient African populations, while 2024 genomic evidence indicates multiple early dispersals out of Africa, including interbreeding with Neanderthals as early as 250,000 years ago, aligning with late MSA migrations.^[43]^[44] These findings emphasize Africa's role as a mosaic of sapiens diversity rather than a single-origin cradle.

Evidence for Behavioral Modernity

Cultural Complexes and Industries

The Middle Stone Age (MSA) in Africa is characterized by distinct cultural complexes or industries that reflect regional technological traditions and adaptations, serving as key markers of variability across the continent. These complexes, often defined by specific tool assemblages and production techniques, emerged within the broader MSA timeframe of approximately 300,000 to 30,000 years ago, highlighting shifts from earlier Acheulean technologies to more diverse and specialized lithic strategies.^[45] Prominent examples include the Still Bay and Howiesons Poort in southern Africa, the Sangoan and Lupemban in central and eastern regions, and the Aterian in the north, each associated with particular environmental contexts and hominin behaviors.^[46] The Still Bay complex, primarily known from southern Africa, dates to around 72,000–71,000 years ago and is distinguished by finely crafted, pressure-flaked points and backed artifacts, indicating advanced knapping skills adapted to open landscapes during Marine Isotope Stage 4. Following closely, the Howiesons Poort industry, also centered in southern Africa, spans approximately 65,000–59,000 years ago and features backed microlithic tools and segmental forms, suggesting innovations in hafting and composite tool use suited to diverse habitats.^[47] In central and eastern Africa, the Sangoan complex represents an early MSA woodland adaptation, dated to roughly 250,000–130,000 years ago, with heavy-duty core-axes and picks for processing wood and other resources in forested environments.^[48] The Lupemban, evolving from or overlapping with the Sangoan in the Congo Basin and surrounding plateaus, is placed in the upper Pleistocene around 200,000–40,000 years ago and includes large foliate points and core tools, reflecting specialized exploitation of tropical woodlands.^[49] Further north, the Aterian industry, widespread across North Africa from Morocco to Sudan, extends from about 150,000 to 40,000 years ago and is identified by tanged or pedunculated tools, likely hafted for projectile use in arid and semi-arid settings.^[50] Debates surrounding these complexes center on whether they represent continuous regional developments within a "generic" MSA substrate or episodic replacements driven by population movements, climate shifts, or cultural diffusion. Recent analyses, including 2024 syntheses, challenge notions of pan-African uniformity, emphasizing instead high regional heterogeneity and questioning the extent to which specific industries like the Still Bay or Aterian overlay a persistent, undifferentiated MSA tradition across the continent. For instance, while some evidence suggests continuity in core reduction methods over millennia, others highlight abrupt shifts in tool forms that may indicate replacement by innovative groups, with limited chronological overlap between complexes in different regions.^[45] These discussions underscore the need for refined dating and comparative studies to clarify sequencing and interconnections, revealing the MSA as a mosaic of localized traditions rather than a monolithic progression.^[45]

Symbolic and Abstract Thinking

The Middle Stone Age (MSA) in Africa provides some of the earliest archaeological evidence for symbolic and abstract thinking among hominins, manifested through deliberate modifications of natural materials that suggest intentional behaviors, including both non-utilitarian (e.g., personal adornment and pattern creation) and practical applications. These proxies for symbolism, including engraved ochre pieces and perforated shells, indicate cognitive capacities for abstract representation and possibly social signaling, predating similar behaviors in other regions by tens of thousands of years. Such artifacts, primarily from South African sites but with earlier examples from North Africa, highlight a tradition of symbolic expression tied to the Still Bay and Howiesons Poort cultural phases around 100,000 to 60,000 years ago.^[51]^[52] At Blombos Cave, engraved ochre fragments dating to approximately 100,000–75,000 years ago represent early instances of abstract patterning, with incisions forming crosshatched, parallel, and geometric designs that exceed simple functional scoring. These 13 documented pieces, recovered from secure MSA layers, show consistent engraving techniques using fine stone points, suggesting a purposeful aesthetic or symbolic intent rather than practical use alone.^[53] Complementing these are ochre processing kits from the same site, including abalone shells containing ground ochre mixtures with charcoal and quartzite, dated to about 100,000 years ago, which imply preparation of pigments potentially for body decoration or ritual applications. Recent 2025 analysis of seven ochre pieces from Still Bay and pre-Still Bay layers (~90,000–70,000 years ago) reveals their modification into specialized retouchers for lithic pressure flaking and refinement, demonstrating multifunctionality in ochre use that combined symbolic and technological purposes.^[25] The presence of such grinding tools and stored compounds underscores advanced manipulation of materials for diverse purposes, though their exact contexts remain interpretive based on ethnographic analogies. Personal adornment is evidenced by perforated Nassarius kraussianus shells from Blombos Cave, dated to around 75,000 years ago, which bear uniform drill holes and wear patterns consistent with suspension as beads; earlier examples (~142,000 years ago) from Bizmoune Cave in Morocco's Aterian context represent the current earliest known in Africa. Over 40 such shells from Blombos, sourced from marine environments 20 kilometers away, indicate deliberate collection, modification, and likely stringing for necklaces or other ornaments, signaling social identity or status. This practice demonstrates abstract thinking in transforming natural objects into culturally meaningful items.^[54] Further evidence of patterning emerges from Diepkloof Rock Shelter, where fragments of engraved ostrich eggshells, dated to approximately 60,000 years ago, feature repetitive geometric motifs such as parallel lines, chevrons, and lattices, possibly used as decorative containers. Hundreds of these fragments from Howiesons Poort layers reveal a sustained engraving tradition spanning generations, with motifs standardized enough to suggest shared symbolic conventions across a regional network. These artifacts collectively illustrate the MSA's role in pioneering abstract expression, laying foundations for later symbolic complexity.^[55]

Planning, Innovation, and Cognition

One hallmark of advanced planning in the Middle Stone Age (MSA) is the deliberate heat treatment of silcrete, a silica-rich stone, to improve its flaking properties for tool production. At Pinnacle Point Site 5-6 in South Africa, archaeological evidence shows systematic heat treatment occurring as early as 164,000 years ago, with predominant use around 72,000 years ago.^[56] This process required hominins to heat the stone to approximately 400°C in controlled fires, then cool it slowly to alter its microstructure, demanding foresight to anticipate future tool-making needs and precise control over fire temperatures without damaging the material.^[56] Such pyrotechnological expertise indicates elevated cognitive abilities, including delayed gratification and abstract planning, as the treatment's benefits—sharper edges and more predictable fractures—were not immediately apparent.^[56] Logistical planning is further evidenced by long-distance transport of raw materials, reflecting organized mobility and resource management. In eastern African MSA sites, obsidian and other lithic materials were moved over distances exceeding 300 km, as seen at sites like Mumba Rockshelter in Tanzania (up to 305 km from sources) and Porc-Epic in Ethiopia (139 km).^[8] At Tsodilo Hills in Botswana, silcrete was transported over 220 km despite local alternatives, suggesting deliberate provisioning strategies where groups anticipated needs during extended foraging or seasonal movements.^[57] These patterns imply coordinated group efforts, possibly involving exchange networks or scheduled expeditions, which required mental mapping of landscapes and prediction of future shortages.^[8] Innovation rates during the MSA show episodic bursts, particularly in the Howiesons Poort techno-complex (~65,000–59,000 years ago), where technological diversification enhanced hunting efficiency. This period marks a rupture in southern African MSA traditions, with rapid adoption of novel production techniques and multifunctional implements forming diversified kits for varied prey.^[58] Use-wear analysis reveals these innovations supported complex hafting and projectile systems, indicating iterative experimentation and adaptation to ecological pressures. Such bursts suggest accelerated cultural transmission and cumulative knowledge, contrasting with slower earlier MSA phases. Cognitive models from the 2020s infer language capabilities through the syntactic complexity of MSA tool-making sequences. Hierarchical planning in knapping, involving nested actions like preparing cores before striking flakes, mirrors syntactic structures in language, as shown by overlapping neural activations in Broca's area during experimental replication.^[59] Predictive processing frameworks posit that MSA hominins used proto-syntactic rules to sequence multi-step operations, facilitating communication of technical knowledge via gesture or speech.^[59] This complexity, evident in Levallois and blade technologies, underscores abstract thinking and recursive cognition essential for innovation.^[59]

Environmental and Subsistence Context

Paleoclimatic Influences

The Middle Stone Age (MSA) in Africa, spanning approximately 300,000 to 30,000 years ago, coincided with significant paleoclimatic variability driven by global Marine Isotope Stages (MIS). During MIS 6 (approximately 191,000–130,000 years ago), pronounced arid phases prevailed across much of eastern and southern Africa, characterized by reduced precipitation and expanded desert conditions.^[13] These dry intervals are associated with a contraction in MSA occupations and technological stasis, as populations likely retreated to refugia with limited innovation in tool technologies.^[60] Such aridity may have contributed to a genetic bottleneck in early modern human populations, constraining cultural developments.^[60] In contrast, MIS 5 (approximately 130,000–71,000 years ago) featured wetter intervals with increased monsoon intensity and expanded habitable zones, facilitating population expansions and bursts of technological innovation.^[61] These humid phases, particularly during MIS 5e (the last interglacial peak around 125,000–110,000 years ago), supported denser vegetation and more reliable water sources, correlating with the emergence of advanced MSA industries like Levallois techniques and hafted tools in regions such as the southern Cape.^[61] Abrupt climatic shifts within MIS 5, including rapid wetting events, appear to have driven adaptive responses that enhanced human dispersal and cultural complexity.^[61] Recent modeling studies as of 2025 further indicate that climate seasonality and predictability during the MSA influenced behavioral diversity, particularly between eastern and northwestern African populations.^[5] Paleoclimatic reconstructions for the MSA rely on multiple proxy records from East and South Africa, revealing regional variability. Speleothems from sites like Pinnacle Point on the southern Cape coast document fluctuating precipitation and temperature between 90,000 and 53,000 years ago, with δ¹⁸O values indicating wetter conditions during interstadials and drier spells linked to global cooling.^[62] Pollen records from inland sites such as Wonderkrater in Limpopo Province show shifts from grassland-dominated landscapes during arid phases to more wooded environments in wetter periods, reflecting changes in effective moisture availability.^[63] In East Africa, lake level proxies from Lake Malawi indicate deep, overflowing conditions during much of MIS 5, contrasting with severe lowstands exceeding 400 meters below modern levels around the MIS 6–5 transition, underscoring the role of orbital forcing in hydroclimate dynamics.^[64] Global events further influenced MSA paleoclimates, notably the Toba supereruption approximately 74,000 years ago during early MIS 4. While the eruption caused widespread cooling and ash deposition, proxy evidence from East African lakes like Malawi shows no significant volcanic winter or ecosystem collapse, with minimal disruption to vegetation and human populations.^[65] In southern Africa, archaeological records from coastal sites indicate continued human occupation and resource use through this period, suggesting resilience to any localized aridification rather than a major population impact. These findings highlight that African MSA groups adapted to transient climatic perturbations without evidence of widespread extinction or migration bottlenecks tied to Toba.

Resource Exploitation Strategies

During the Middle Stone Age, hominins in Africa exhibited a notable shift toward more diverse hunting strategies, incorporating the use of hafted projectile points to target a broader range of prey, including small game such as antelope and smaller mammals, around 70,000 years ago. This innovation is evidenced by stone points from sites like Blombos Cave in South Africa, where experimental analyses confirm their suitability as spear tips for thrusting or throwing, enabling efficient hunting of agile, smaller animals that were less accessible to earlier thrusting spears. Such adaptations reflect increased technological sophistication in subsistence, allowing for exploitation of varied ecological niches beyond large herbivores.^[66] Parallel to these hunting advancements, there was intensified processing of plant resources, as indicated by the presence of grinding stones bearing starch residues from grasses and tubers, suggesting systematic preparation of carbohydrate-rich foods. At sites in the Niassa Rift of Mozambique, dated between approximately 105,000 and 42,000 years ago, microscopic analysis of these tools revealed starch grains from wild sorghum and other Poaceae species, pointing to deliberate grinding for flour or porridge production to supplement faunal diets. This practice underscores a broadening of resource base, enhancing nutritional resilience in fluctuating environments.^[67] Coastal adaptations during the MSA are exemplified by the exploitation of marine resources, including shellfish collection that resulted in substantial middens at sites like Klasies River in South Africa around 115,000 years ago. Uranium-thorium dating of shell layers confirms intensive gathering of limpets and mussels during Marine Isotope Stage 5, with accumulations exceeding 10 meters in thickness, indicating repeated visits and a reliable protein source amid terrestrial variability. These middens highlight early behavioral flexibility in accessing intertidal zones, potentially supporting population growth in southern African coastal regions.^[68]

Key Sites and Discoveries

Iconic African Sites

Blombos Cave, located on the southern coast of South Africa, represents a cornerstone of Middle Stone Age research due to its rich assemblage of artifacts demonstrating early symbolic behavior and advanced lithic technology. Excavations have uncovered engraved pieces of red ochre from layers dated to approximately 100,000 to 70,000 years ago, including abstract geometric patterns incised with fine lines using pointed stone tools, suggesting intentional symbolic expression rather than utilitarian function. These engravings, found in Still Bay layers around 77,000 years ago, are among the earliest evidence of abstract art in Africa, with over 13 pieces documented, some featuring crosshatched designs up to 3.5 cm long. Additionally, the site yielded Still Bay points—bifacial foliate stone tools—crafted from silcrete and other fine-grained materials, numbering over 370 specimens, which indicate specialized pressure flaking techniques and possible use as spear tips for hunting. The ochre pieces also show signs of grinding and mixing, hinting at pigment use in body decoration or ritual, while the points reflect technological innovation linked to marine resource exploitation in the region's coastal environment. Jebel Irhoud in Morocco stands as the earliest known site associated with anatomically modern Homo sapiens, pushing back the timeline of our species' origins. Discovered remains include a partial cranium (Irhoud 10), mandible (Irhoud 11), teeth, and limb bones from multiple individuals, exhibiting a modern facial structure with a flat face and small chin, combined with an elongated braincase more akin to archaic hominins.^[69] Thermoluminescence dating of heated flint artifacts places these fossils at 315,000 ± 34,000 years ago, making Jebel Irhoud the richest and most securely dated early Homo sapiens site in Africa.^[69] The associated lithic assemblage features Levallois core technology, characterized by prepared flakes and points for scrapers and blades, typical of Middle Stone Age industries and indicating planned knapping strategies for tool production.^[69] This mosaic morphology and tool kit support a pan-African model for Homo sapiens evolution, with gradual morphological changes occurring across the continent over at least 400,000 years rather than a singular recent origin.^[69] Klasies River Mouth, situated along South Africa's southern coast, provides critical evidence for the emergence of modern human morphology and adaptive strategies in coastal settings. The site has yielded over 50 human fossil fragments, primarily from adults, including cranial and postcranial elements showing fully modern Homo sapiens traits such as a high forehead and globular braincase, dated to between 120,000 and 60,000 years ago, with key layers in Cave 1 around 115,000 to 90,000 years ago.90054-Y) These remains, recovered from hearths and debris, indicate repeated occupation by behaviorally modern populations during Marine Isotope Stage 5.90054-Y) Notably, the faunal assemblage documents intensive marine resource exploitation, including shellfish middens with limpets and mussels, seal bones, and fish remains, representing one of the earliest sustained uses of aquatic foods to supplement terrestrial hunting.^[70] Bone tools and grinding stones further suggest processing of marine and plant resources, underscoring the site's role in understanding how coastal adaptations supported population resilience during environmental fluctuations.^[70] Border Cave in South Africa's KwaZulu-Natal province offers a continuous stratigraphic sequence spanning much of the Middle Stone Age, with key findings illuminating early ritual practices. The site preserves layers from approximately 200,000 to 50,000 years ago, including Howiesons Poort industries marked by backed tools and heat-treated silcrete. A significant discovery is the intentional burial of a 4- to 6-month-old infant (BC3) in a shallow pit within a Howiesons Poort layer dated to around 74,000 years ago, accompanied by a perforated Conus shell bead, suggesting deliberate interment and possible ornamentation as an early form of symbolic burial. The infant's remains, including skull fragments and long bones, were placed in a flexed position with red ochre traces nearby, indicating ritualistic elements in death practices among Middle Stone Age groups. Additional artifacts, such as bone points and digging sticks, point to diverse subsistence strategies involving plant processing and small-game hunting, contributing to models of technological continuity in the region.

Recent and Emerging Findings

In 2025, archaeological surveys in the Río Campo region of Equatorial Guinea uncovered widespread evidence of Middle Stone Age (MSA) occupation in the Atlantic rainforests of West Central Africa, with lithic assemblages including Levallois flakes and cores dated to the late Pleistocene, approximately 40,000 years ago, demonstrating early human adaptation to dense tropical forest environments previously thought inhospitable for such technologies.^[71] These findings, derived from 11 field campaigns since 2014, identified over 50 Paleolithic sites with recurring technological patterns, such as prepared core reduction methods, suggesting sustained MSA presence in a region long underrepresented in the archaeological record and challenging assumptions about human avoidance of humid equatorial zones during Marine Isotope Stage 3.^[72] The artifacts, primarily quartzite tools, indicate resource exploitation strategies tailored to rainforest settings, including woodworking and plant processing, thereby expanding the known geographic and ecological scope of MSA populations.^[73] At Olduvai Gorge in Tanzania, a 2025 lithic technological analysis of the newly documented Dorothy Garrod Site (DGS), located at the junction of the main gorge and Side Gorge, revealed evidence of early blade production within an MSA context dating to approximately 300,000 years ago, predating previously recognized blade technologies in East Africa.^[74] The assemblage, comprising over 1,000 artifacts dominated by chert and quartzite, exhibits prismatic blade knapping and Levallois methods, indicating advanced planning and raw material optimization during the early MSA, potentially linked to environmental shifts in the Eastern Rift Valley.^[75] This discovery refines understandings of technological continuity from the Acheulean-MSA transition, highlighting Olduvai's role in tracing the emergence of elongated flake production across African landscapes.^[76] A 2025 excavation at the Longtan site in Yunnan Province, southwest China, yielded stone tools dated to around 55,000 years ago that closely parallel Middle Stone Age styles from Africa and Europe, such as Quina scrapers and Levallois points typically associated with Neanderthals, sparking debates on convergent evolution versus intercontinental dispersal of lithic traditions.^[77] The tools, made from local quartz and sandstone, show flake-scar patterns and retouch techniques indistinguishable from European Middle Paleolithic industries, suggesting that unidentified hominin groups in East Asia independently developed or adopted similar reduction strategies during the late Middle Pleistocene, without direct evidence of migration.^[78] This finding prompts reevaluation of global MSA equivalents, emphasizing parallel technological innovations in response to comparable ecological pressures rather than a singular African origin for such advancements.^[79] Reappraisals of Moroccan sites in 2025, including Rhafas Cave and other North African locales, have refined the chronology of the MSA to Late Stone Age (LSA) transition through new optically stimulated luminescence (OSL) and radiocarbon dates for Aterian industries, placing their emergence at approximately 150,000 years ago and persistence until 20,000 years ago, later than previously estimated.^[80] These updates, based on integrated stratigraphic and use-wear analyses of quartzite tools, reveal gradual shifts from tanged points and Levallois cores to microlithic forms, indicating behavioral continuity rather than abrupt replacement during Marine Isotope Stages 5-2.^[81] The revised timeline underscores the Aterian's role as a bridge between MSA hunting adaptations and LSA mobility patterns in the Maghreb, influenced by climatic fluctuations and resource availability.^[82]

References

[1]
Continuity of the Middle Stone Age into the Holocene - Nature
Jan 11, 2021 · The African Middle Stone Age (MSA) is a cultural phase characterized by features such as a focus on prepared core lithic technology, hafting, ...
[2]
Homo sapiens | The Smithsonian Institution's Human Origins Program
Jan 3, 2024 · Homo sapiens, the species of all living humans, evolved in Africa about 300,000 years ago. They have large brains, lighter skeletons, and a ...<|separator|>
[3]
Technological Trends in the Middle Stone Age of South Africa ...
Its unifying technological characteristic is the almost exclusive use of a blade and bladelet production system, but subtle changes in types of backed artefacts ...Skip main navigation · Abstract · Early Middle Stone Age · MIS 5 Middle Stone Age
[4]
Scientists Discover Evidence of Early Human Innovation, Pushing ...
Mar 15, 2018 · The first evidence of human life in the Olorgesailie Basin comes from about 1.2 million years ago. For hundreds of the thousands of years, ...
[5]
Climate seasonality and predictability during the middle stone age ...
Apr 4, 2025 · The Middle Stone Age (MSA) represents the earliest behavioural signature of our species, Homo sapiens, across Africa from ca. 300,000 to 30,000 ...Introduction · Results · Methods
[6]
Blombos Cave, Southern Cape, South Africa: Preliminary Report on ...
The Later- and Middle Stone Age levels at Blombos Cave (BBC) were excavated over four field seasons between 1992 and 1999. Here we report on the results.
[7]
Middle and late Pleistocene Middle Stone Age lithic technology from ...
Excavations at a complex of caves and open air sites at Pinnacle Point, Mossel Bay, Southern Africa have uncovered rich stratified assemblages of Middle Stone ...
[8]
Variability in the Middle Stone Age of Eastern Africa
All of the early fossils of H. sapiens from eastern Africa are associated with Middle Stone Age (MSA) artifacts. These include those from the Kibish Formation ( ...Missing: timeline | Show results with:timeline
[9]
A spatiotemporally explicit paleoenvironmental framework for the ...
Mar 7, 2022 · A spatiotemporally explicit paleoenvironmental framework for the Middle Stone Age of eastern Africa. Lucy Timbrell,; Matt Grove, ...
[10]
Full article: The generic Middle Stone Age: fact or fiction?
Mar 20, 2024 · The Middle Stone Age (MSA) constitutes the archaeological record and material culture of the earliest populations of Homo sapiens on the African ...
[11]
Chronology of the Acheulean to Middle Stone Age transition in ...
A. J. H. Goodwin, An introduction to the Middle Stone Age in South Africa. S. Afr. J. Sci. 25, 410–418 (1928).
[12]
The African Middle Stone Age
### Summary of Middle Stone Age Content
[13]
The structure of the Middle Stone Age of eastern Africa - ScienceDirect
Sep 1, 2018 · The Middle Stone Age (MSA) of eastern Africa has a long history of research and is accompanied by a rich fossil record, which, combined with ...Missing: timeline | Show results with:timeline
[14]
A Chronological Perspective on the Acheulian and Its Transition to ...
A number of researchers have hypothesised that the Acheulian first occurred simultaneously in southern and eastern Africa at around 1.7-1.6 Ma. A chronological ...
[15]
From Acheulean to Middle Stone Age in the Kapthurin Formation ...
The Kapthurin Formation shows MSA technology before 285,000 years ago, with interstratified Acheulean and MSA artifacts, and the Levallois tradition beginning ...Missing: evidence | Show results with:evidence
[16]
[PDF] Levallois Lithic Technology from the Kapthurin Formation, Kenya
Aug 19, 2006 · MSA Levallois technology differs from that of the Acheulian in (1) the reduction in flake size, (2) increased variation in flake shape ...Missing: discoidal | Show results with:discoidal
[17]
New radiometric ages for the Fauresmith industry from Kathu Pan ...
New radiometric ages for the Fauresmith industry from Kathu Pan, southern Africa: Implications for the Earlier to Middle Stone Age transition · Abstract.
[18]
Early Levallois technology and the Lower to Middle Paleolithic ...
Sep 26, 2014 · The technique appeared in the archeological record across Eurasia 200 to 300 thousand years ago (ka) and appeared earlier in Africa. Adler et al ...
[19]
Why Levallois? A Morphometric Comparison of Experimental ...
Jan 23, 2012 · Middle Palaeolithic stone artefacts referred to as 'Levallois' have caused considerable debate regarding issues of technological ...
[20]
Middle Stone Age technology from MIS 6 and MIS 5 at ...
Oct 15, 2023 · Klipfonteinrand 1 (KFR1) is a foundational but poorly documented Middle Stone Age (MSA) site located in the south west of South Africa.Missing: key | Show results with:key
[21]
The Still Bay points of Blombos Cave (South Africa) - ScienceDirect
The late Middle Stone Age of South Africa between 77 and 35 ka contains three technocomplexes known as the Still Bay, the Howiesons Poort and the post-Howiesons ...
[22]
Blade technology and tool forms in the Middle Stone Age of South ...
One such episode is the Howiesons Poort (HP). The appearance of a range of small geometric forms, apparently used as insets in multi-component tools, has been ...
[23]
Variable Perspectives on “Standardization in the Stone Age”
Nov 24, 2023 · They explore basic flake morphological standardization in each method by comparing 728 flakes produced using seven lithic reduction strategies ( ...
[24]
Implications for complex cognition from the hafting of tools with ... - NIH
Jun 16, 2009 · Abstract. Compound adhesives made from red ochre mixed with plant gum were used in the Middle Stone Age (MSA), South Africa.
[25]
The pigments from Pinnacle Point Cave 13B, Western Cape, South ...
This article is part of 'The Middle Stone Age at Pinnacle Point Site 13B, a Coastal Cave near Mossel Bay (Western Cape Province, South Africa)' Special Issue.
[26]
The Emergence of Habitual Ochre Use in Africa and its Significance ...
Dec 14, 2022 · At archaeological sites attributed to the Middle Stone Age (MSA) of Africa, ochre is typically found as red, yellow or brown-colored lumps or ...
[27]
90000 year-old specialised bone technology in the Aterian Middle ...
With a date of approximately 90 ka, the tool from Dar es-Soltan 1 represents the oldest specialised bone tool yet found in a well-dated context that is clearly ...Missing: ya heat treatment
[28]
Middle Stone Age bone tools from the Howiesons Poort layers ...
This paper describes three bone tools: two points and the end of a polished spatula-shaped piece, from unequivocal HP layers at Sibudu Cave (with ages greater ...
[29]
Nassarius kraussianus shell beads from Blombos Cave: evidence ...
Since 1991, excavations at Blombos Cave have yielded a well-preserved sample of faunal and cultural material in Middle Stone Age (MSA) levels.
[30]
New fossils from Jebel Irhoud, Morocco and the pan-African origin of ...
Jun 8, 2017 · New human fossils from Jebel Irhoud (Morocco) document the earliest evolutionary stage of Homo sapiens and display modern conditions of the ...
[31]
Middle and later Pleistocene hominins in Africa and Southwest Asia
200,000 years ago, individuals more similar to recent humans are present in the African record. These fossils are associated with Middle Stone Age lithic ...
[32]
Homo naledi and Pleistocene hominin evolution in subequatorial ...
May 9, 2017 · Homo naledi, dated to 236-335 ka, was found in the Rising Star cave system, showing a diverse hominin lineage in subequatorial Africa, with ...
[33]
The evolution of modern human brain shape - PMC - PubMed Central
Jan 24, 2018 · The endocranial volumes of early H. sapiens individuals (Jebel Irhoud 1, 1375 ± 6 ml; Jebel Irhoud 2, 1467 ± 6 ml; Omo 2, 1491 ± 4 ml) fall ...
[34]
Endocranial volumes and human evolution - F1000Research
May 30, 2023 · Enlarging brains have been held up as the classic (if not the only) example of a consistent long-term trend in human evolution.
[35]
The evolution of modern human brain shape | Science Advances
Jan 24, 2018 · Incomplete and distorted fossils were reconstructed using established methods ... Bookstein, Principles for the virtual reconstruction of hominin ...Missing: methodologies MSA
[36]
The evolution of the temporoparietal junction and posterior superior ...
Feb 7, 2019 · In this review, we discuss how the expansion and rearrangement of the temporoparietal junction and posterior superior temporal sulcus (TPJ-pSTS) ...
[37]
https://www.nature.com/articles/s41586-020-2165-4
[38]
Principles for the virtual reconstruction of hominin crania - PubMed
We outline a general methodological framework by which missing information about biological specimens can be estimated using geometric morphometric methods.Missing: methodologies MSA
[39]
(PDF) Virtual Reconstruction of Modern and Fossil Hominoid Crania
PDF | Most hominin cranial fossils are incomplete and require reconstruction prior to subsequent analyses. Missing data can be estimated by geometric.Missing: methodologies MSA
[40]
Recovering signals of ghost archaic introgression in African ...
Feb 12, 2020 · Our results reveal the substantial contribution of archaic ancestry in shaping the gene pool of present-day West African populations.
[41]
Genomic inference of a severe human bottleneck during ... - Science
Aug 31, 2023 · Results showed that human ancestors went through a severe population bottleneck with about 1280 breeding individuals between around 930,000 and ...
[42]
Out-of-Africa, the peopling of continents and islands - PubMed Central
AMH left Africa via a single southern exit about 70 000 years ago and rapidly spread around the Indian Ocean towards the Antipodes.
[43]
https://www.sciencedaily.com/releases/2025/03/250318141412.htm
[44]
The Still Bay and Howiesons Poort at Sibudu and Blombos
We present the results of a systematic technological and typological analysis of the Still Bay assemblages from Sibudu and Blombos.
[45]
Single-grain OSL chronologies for the Still Bay and Howieson's ...
In 2008, we proposed a chronology for the SB and HP, based on a systematic dating study of nine sites spanning a variety of climatic and ecological zones in ...<|separator|>
[46]
A West African Middle Stone Age site dated to the beginning of MIS 5
... Still Bay and the Howiesons Poort (for synthesis see e.g., Wurz, 2013; Porraz et al., 2013). Furthermore, on a larger scale, models of early H. sapiens ...
[47]
Riddles wrapped inside an enigma. Lupemban MSA technology as ...
Mar 7, 2022 · The early MSA Lupemban industry has been considered a circumstantially strong candidate for the initial settlement of the African rainforest ...
[48]
The Aterian and its place in the North African Middle Stone Age
In the north of Africa, these technological changes are most often identified with the highly distinctive tanged tool assemblages of the middle and later MSA.
[49]
Fire As an Engineering Tool of Early Modern Humans - Science
Aug 14, 2009 · The controlled use of fire was a breakthrough adaptation in human evolution. It first provided heat and light and later allowed the physical properties of ...
[50]
Provenancing of silcrete raw materials indicates long-distance ...
Provenancing of silcrete raw materials indicates long-distance transport to Tsodilo Hills, Botswana, during the Middle Stone Age · Results of geochemical ...
[51]
Functional insights into the innovative Early Howiesons Poort ...
The Howiesons Poort ('HP') is characterized by a set of technological innovations that mark a rupture in the Southern African Middle Stone Age.
[52]
Stone tools, predictive processing and the evolution of language - Pain
Apr 25, 2022 · Recent work by Stout and colleagues indicates that the neural correlates of language and Early Stone Age toolmaking overlap significantly.
[53]
Technological Trends in the Middle Stone Age of South Africa ...
This historical review of the technological diversity in the Middle Stone Age of South Africa emphasizes that the roots of some innovations may lie in the ...Missing: phases stasis
[54]
Development of Middle Stone Age innovation linked to rapid climate ...
May 21, 2013 · Here we show that, contrary to some previous studies, the occurrence of innovation was tightly linked to abrupt climate change.
[55]
A high resolution and continuous isotopic speleothem record of ...
Here we present the first such record for climate and environmental change from 90 000 to 53 000 years ago from the southern Cape coast.
[56]
Multiproxy record of late Quaternary climate change and Middle ...
Sep 1, 2014 · Here we provide a multiproxy record of climate change and human occupation at Wonderkrater, a spring and peat mound site situated in the ...
[57]
Continuous 1.3-million-year record of East African hydroclimate, and ...
Results show evidence for 24 lake level drops of more than 200 m during the Late Quaternary, including 15 lowstands when water levels were more than 400 m lower ...
[58]
Toba supereruption: Age and impact on East African ecosystems
Aug 13, 2013 · Toba supereruption: Age and impact on East African ecosystems. ... Two recent synopses (and the primary references cited therein) highlight ...
[59]
An Experimental Investigation of the Functional Hypothesis ... - NIH
Aug 27, 2014 · Middle Stone Age points from the Gademotta Formation, Ethiopia date ... 70,000-year-old Middle Stone Age levels at Blombos Cave, South ...
[60]
Middle Stone Age starch acquisition in the Niassa Rift, Mozambique
Jan 20, 2017 · Keywords. AfricaMiddle Stone AgeMozambiqueNiassaStarch grains ... A functional analysis of grinding stones from an early holocene site at ...
[61]
U-Th dating, taphonomy, and taxonomy of shell middens at Klasies ...
The three ages, 110,060 ± 1,100, 109,800 ± 970, and 106,000 ± 2,100 years, place the BOS layer as the base of the SASL sub-member at over 110 ka, making the ...
[62]
Frontiers | A strontium (87Sr/86Sr) isoscape of Southern Ethiopia
We used the Middle Stone Age (MSA) site of Gotera as a case study and conducted 87Sr/86Sr isotope analysis of faunal tooth enamel. The results show that our ...
[63]
https://www.sciencedirect.com/science/article/abs/pii/S0277379114002467
[64]
Page Not Found - University of Chicago Press
**Summary:**
[65]
Middle Stone Age (MSA) in the Atlantic rainforests of Central Africa ...
Feb 1, 2025 · This study addresses these issues by analyzing recently recovered Pleistocene prehistoric evidence from the Republic of Equatorial Guinea, a ...
[66]
(PDF) Widespread evidence of Middle Stone Age (MSA) presence in ...
Aug 4, 2025 · Widespread evidence of Middle Stone Age (MSA) presence in Equatorial Guinea (West-Central Atlantic Africa). May 2025; Quaternary International ...
[67]
40,000-year-old stone tools uncover early human adaption to ...
Jan 15, 2025 · Morales, J. I. (2025). Middle Stone Age (MSA) in the Atlantic rainforests of Central Africa. The case of Río Campo region in Equatorial Guinea.
[68]
Exploring the Middle Stone Age lithic technology at DGS, Olduvai ...
Jan 18, 2025 · This paper presents the lithic technology analysis of Dorothy Garrod Site (DGS), a newly-documented MSA site located at the junction of the main gorge and the ...
[69]
(PDF) Exploring the Middle Stone Age lithic technology at DGS ...
Aug 6, 2025 · This paper presents the lithic technology analysis of Dorothy Garrod Site (DGS), a newly-documented MSA site located at the junction of the main ...
[70]
Lithic technology from level 22 at Nasera (Tanzania): The Kisele ...
This study presents a techno-typological analysis of level 22 at the site of Nasera (northern Tanzania), shedding new light on the Kisele industry (Middle ...
[71]
Stone Tools Discovered in China Resemble Neanderthal ...
Apr 4, 2025 · Stone Tools Discovered in China Resemble Neanderthal Technology Used in Europe, Creating a Middle Stone Age Mystery. Archaeologists ...Missing: paralleling convergent
[72]
Stone tools discovery suggests Neanderthals migrated as far ... - CNN
Apr 1, 2025 · Stone tools discovered in southwest China could challenge what's known about human origins during the Stone Age. The tools, found at Longtan ...Missing: paralleling convergent evolution
[73]
50,000-Year-Old Tools Found in China Could Point to Unknown ...
Apr 1, 2025 · Researchers found European-style Quina tools in China, dating back 55,000 years, challenging the view that East Asia's Middle Paleolithic ...Missing: paralleling convergent
[74]
https://link.springer.com/article/10.1007/s12520-024-02137-8
[75]
A reappraisal of the Middle to Later Stone Age prehistory of Morocco
Here we outline some of the critical findings concerning the Middle (MSA) to Later (LSA) Stone Age periods. We describe shifts in the understanding of the ...
[76]
Insights from use-wear analysis of Rhafas Cave, Northeast Morocco
This study presents the results of the use-wear analysis on the quartzite assemblage from the Aterian sequence of Rhafas cave.