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Semi-deciduous

Semi-deciduous refers to a botanical leaf habit in plants, particularly trees and shrubs, that is intermediate between fully deciduous and evergreen, where foliage is retained for most of the year but partially or briefly shed in response to environmental stresses such as seasonal drought or cold temperatures.^[1] This shedding typically involves older leaves dropping as new growth emerges, allowing the plant to maintain some photosynthetic capacity year-round without the complete leafless dormancy of deciduous species or the persistent canopy of evergreens.^[2] In ecological contexts, semi-deciduous plants are prominent in tropical and subtropical dry forests, where they adapt to pronounced wet-dry seasonal cycles by partially defoliating during the dry period, often with rapid leaf flushing upon the return of rains.^[3] Such species exhibit intermediate physiological traits, including higher wood density and slower leaf turnover compared to fully deciduous plants, enabling efficient resource use in water-limited environments.^[3] Semi-deciduous vegetation formations, as classified in standard ecological systems, feature upper canopy trees that are primarily drought-deciduous alongside evergreen understory elements, supporting biodiversity in regions like the neotropics and parts of Africa.^[4] These plants play key roles in ecosystem dynamics, contributing to nutrient cycling through periodic leaf litter and providing habitat continuity despite partial defoliation.^[4] Examples include species like Astronium graveolens in tropical dry forests, which may vary in expression of semi-deciduous behavior across sites depending on local climate.^[3] In temperate zones, the habit appears in response to mild winters, as seen in certain shrubs that drop leaves only under stress.^[1] Overall, the semi-deciduous strategy balances water conservation with sustained growth, influencing forest structure and resilience to climate variability.

Definition and Characteristics

Definition

Semi-deciduous plants, also known as semi-evergreen in some contexts, represent an intermediate leaf habit in botany, characterized by the partial shedding of leaves, particularly older foliage, concurrent with the emergence of new growth, while maintaining some leaves year-round.^[5] This contrasts with fully deciduous species, which lose all leaves during unfavorable seasons, and evergreen species, which retain their foliage continuously without significant seasonal loss.^[6] The semi-deciduous strategy allows plants to balance resource conservation and photosynthetic activity in variable environments. The primary triggers for leaf shedding in semi-deciduous plants are environmental stresses such as seasonal dryness, which limits water availability and prompts partial defoliation to reduce transpiration, leading to a shorter but not entirely dormant growing period.^[7] In some cases, mild cold temperatures in subtropical or transitional climates can also initiate this response, though it remains less pronounced than in fully deciduous habits, avoiding complete leaflessness.^[2] Unlike evergreens, which exhibit lower water use efficiency through persistent leaves, semi-deciduous plants achieve a compromise by shedding only a portion of their canopy, often resulting in visible growth rings that reflect this adaptive phenology.^[8] This leaf retention pattern underscores the semi-deciduous habit's role as a transitional adaptation, enabling sustained but moderated physiological function amid fluctuating conditions.^[5]

Physiological Traits

Semi-deciduous plants exhibit a partial leaf abscission process, similar to that in many plants, where hormonal signals promote the shedding of older, senescent leaves while some foliage is retained. This mechanism involves the formation of an abscission zone at the petiole base, allowing selective detachment without a complete leafless state. New buds often form simultaneously with this shedding, enabling rapid replacement of lost leaves as environmental conditions improve.^[9] Leaf structure in semi-deciduous plants features adaptations like intermediate leaf mass per area (LMA), which is higher than in fully deciduous species but lower than in evergreens, contributing to reduced water loss during periods of partial dormancy. These traits support a balance between resource conservation and functional longevity.^[10]^[6] The growth cycle of semi-deciduous plants includes a shorter dormancy period, typically lasting 1-4 weeks in response to seasonal stress, compared to the months-long dormancy of fully deciduous species. This brief quiescence allows for quicker recovery and resumption of growth upon the return of favorable conditions, such as increased moisture, without entering a prolonged inactive phase.^[9] Photosynthetic continuity is maintained through the retention of a portion of foliage during off-seasons, enabling partial carbon fixation even under suboptimal conditions. This intermediate strategy results in higher maximum photosynthetic rates per leaf area than in evergreens but lower than in deciduous plants, supporting sustained but reduced metabolic activity without a leafless interval.^[6]

Comparisons to Other Leaf Habits

With Deciduous Plants

Fully deciduous plants shed 100% of their leaves in a highly synchronized event, typically during autumn in temperate regions, allowing for complete physiological dormancy over winter. In contrast, semi-deciduous plants exhibit partial leaf drop, losing only a portion of their foliage asynchronously, often retaining some leaves or dead foliage into the unfavorable season.^[11]^[12] This difference in extent enables semi-deciduous species to avoid total shutdown while still reducing exposure to stress.^[13] The seasonal response to environmental cues also diverges between the two habits. Deciduous leaf abscission is primarily triggered by decreasing temperatures and shortening photoperiods in temperate zones, promoting dormancy to conserve energy during cold periods.^[12] Semi-deciduous plants, more common in subtropical or seasonally dry milder climates, respond mainly to water scarcity, shedding leaves irregularly as soil moisture declines to prevent desiccation without entering full dormancy.^[11]^[12] In terms of energy allocation, deciduous plants undergo a total metabolic halt during leaf-off periods, channeling resources into a substantial spring bud burst for rapid canopy redevelopment and high photosynthetic output in the growing season.^[12] Semi-deciduous plants, by retaining some foliage, sustain low-level metabolism and limited photosynthesis year-round, allowing continuous but modest resource acquisition and reducing the energetic cost of full regrowth.^[11]^[13] Evolutionarily, the deciduous habit optimizes for severe cold avoidance in temperate environments, where complete leaf loss minimizes frost damage and mechanical stress from snow or ice loads.^[14] Semi-deciduous strategies represent a trade-off, balancing water conservation during dry spells with opportunities for ongoing growth in less extreme, variable climates, thereby enhancing adaptability in transitional habitats.^[12]^[14]

With Evergreen Plants

Semi-deciduous plants differ from evergreen species in their foliage retention patterns, exhibiting partial retention during dry or unfavorable periods, accompanied by periodic shedding, whereas evergreens retain their foliage year-round through slow, gradual turnover of older leaves.^[15] In terms of leaf longevity, evergreen leaves typically persist for 2-5 years, enabling sustained photosynthesis over extended periods, while semi-deciduous leaves have a shorter average lifespan of 5-12 months, with older leaves preferentially shed to conserve resources during stress.^[16]^[17]^[18] Semi-deciduous plants, akin to deciduous species, demonstrate more efficient nutrient retranslocation from senescing leaves, recovering around 65% of nitrogen compared to about 58% in evergreens, which instead depend on the durability of sclerophyllous leaves to minimize nutrient loss over their longer lifespan.^[19]^[20] Regarding stress tolerance, evergreens emphasize drought resistance through thick, tough leaves that reduce water loss and withstand prolonged aridity, whereas semi-deciduous species employ a flexible strategy, shedding portions of foliage to adjust to variable seasonal conditions like intermittent droughts.^[21]^[14]

Ecological Significance

Environmental Adaptations

Semi-deciduous plants exhibit partial defoliation as a key strategy for water conservation, particularly during dry seasons, by reducing the transpiring leaf surface area and thereby minimizing water loss through transpiration. This gradual shedding of older leaves while retaining younger ones helps maintain hydraulic balance and prevents severe dehydration, allowing the plant to conserve water more effectively than fully evergreen species that retain all foliage under stress. Studies on tropical species indicate that such partial leaf loss can reduce overall transpiration, depending on the intensity of drought, enabling survival in environments with seasonal water limitations.^[22]^[23] This leaf habit also confers climate versatility, enabling semi-deciduous plants to thrive in transitional zones such as subtropical regions characterized by irregular rainfall patterns and mild winters that do not induce full dormancy. In these areas, where dry periods last 3-4 months but are interspersed with moist intervals, partial defoliation allows plants to adjust dynamically to fluctuating water availability without the complete cessation of photosynthesis seen in fully deciduous species. For instance, in neotropical understory herbs, this facultative semi-deciduous response supports adaptation to seasonal moist forests with variable precipitation, balancing growth during wet phases and conservation during dry ones.^[23]^[24]^[25] In terms of interactions with soil and light, semi-deciduous plants enhance nutrient cycling in nutrient-poor soils by resorbing essential nutrients, such as nitrogen and phosphorus, from senescing leaves prior to abscission, thereby recycling them internally for new growth and reducing reliance on external inputs. This resorption efficiency supports persistence in low-fertility environments common to transitional habitats. Additionally, their partial shade tolerance facilitates understory roles, where they exploit dappled light conditions by maintaining photosynthetic activity with reduced leaf area.^[26]^[23]^[27]

Global Distribution

Semi-deciduous plants, characterized by partial leaf shedding during dry periods, are predominantly distributed in tropical and subtropical regions worldwide, where they form key components of seasonal dry forests. These ecosystems are most prevalent in Central and South America, encompassing areas such as Mexico, the Yucatán Peninsula, and eastern Brazil, where they cover extensive tracts interspersed with savannas and woodlands. In Africa, semi-deciduous vegetation thrives in West and Central regions, including the Guineo-Congolian zone and Miombo woodlands, extending from Senegal to Tanzania. Southeast Asia and parts of South Asia, such as India and Thailand, also host significant stands, particularly in monsoon-influenced lowlands and deciduous dipterocarp forests.^[28]^[29]^[30] Habitat preferences for semi-deciduous plants align with transitional environments like savanna edges, open woodlands, and zones affected by seasonal monsoons, typically receiving 500-1500 mm of annual rainfall. These plants peak in prevalence within climates featuring 3-6 month dry seasons, where precipitation drops below 50 mm monthly, allowing for adaptations like partial defoliation to conserve water during drought. Such conditions are common in the Aw/As Köppen climate zones but are less frequent in strictly temperate latitudes or hyper-arid deserts, where full evergreen or complete deciduous strategies dominate instead.^[28]^[30]^[31] Human activities have significantly impacted semi-deciduous habitats, with significant deforestation resulting in 48-68% loss of original cover globally, much of it since 1950, driven primarily by agricultural expansion, logging, and urbanization. In the Americas, for instance, over 60% of original tropical dry forest cover—much of it semi-deciduous—has been cleared in some subregions, exacerbating fragmentation and biodiversity decline. Similar pressures in African and Asian semi-deciduous areas have reduced intact forest patches, though protected areas now safeguard about 30% of remaining extent globally. As of 2025, tropical dry forests, including semi-deciduous formations, are disappearing faster than rainforests due to intensified fires, climate variability, and land-use changes, particularly in Africa where logging and invasive species threaten Miombo woodlands. Conservation efforts emphasize improved monitoring, restoration initiatives, and targeted protection to enhance resilience.^[28]^[29]^[30]^[32]^[33]^[31]

Examples and Applications

Key Plant Species

Prominent examples of semi-deciduous plants include species adapted to varying environmental stresses, such as drought, seasonal monsoons, or mild winters, where they shed a portion of their leaves rather than all foliage. These plants demonstrate partial leaf loss as a strategy for resource conservation while maintaining some photosynthetic capacity. Mesquite trees (Prosopis spp.), desert-adapted species native to arid regions, exhibit semi-deciduous behavior by partially dropping leaves during severe drought conditions to conserve water, while retaining enough foliage to survive and regrow rapidly with moisture return; they are common in the Southwest United States and Mexico.^[34]^[35] In tropical Asia, Magnolia champaca is a semi-deciduous tree that sheds older leaves prior to the onset of new growth, often coinciding with monsoon transitions, allowing it to balance water use in humid yet seasonally variable climates; it is valued for its fragrant flowers and durable timber used in construction and furniture.^[36]^[37] Neotropical legumes of the Inga genus, such as Inga edulis, support their role in agroforestry systems where they fix atmospheric nitrogen to enrich soils; these trees are widespread in Central and South American rainforests and plantations.^[38]^[39] The Southeast Asian ylang-ylang tree (Cananga odorata) displays brief leaf drop following flowering events or under mild drought stress, minimizing water loss while sustaining its evergreen habit overall; it serves as a key source of essential oils extracted from its aromatic flowers for perfumes and aromatherapy.^[40] Regional variations highlight semi-deciduous traits in temperate settings, such as New Zealand's Plagianthus species (e.g., Plagianthus regius), which partially retain leaves through mild winters but shed more in colder conditions, adapting to the archipelago's variable coastal climates.^[41]^[42]

Semi-Deciduous Forests

Semi-deciduous forests represent a distinct vegetation type consisting of mixed woodlands where 50-100% of the canopy trees are deciduous or semi-deciduous, resulting in partial seasonal defoliation and bare areas during dry periods. These ecosystems feature greater than 30% canopy cover, occur primarily below 1200 m altitude, and are dominated by broadleaf species comprising over 75% of the canopy. This mixed leaf habit distinguishes them from fully evergreen or completely deciduous formations, enabling a balance between seasonal adaptation and continuous productivity.^[43] The structure of semi-deciduous forests is typically multi-layered, including emergent trees reaching 20-40 m in height that protrude above the main canopy, a dense understory of shrubs and smaller trees, and a herbaceous ground layer that remains active year-round. Trees in these forests often exhibit greater height for a given diameter compared to those in adjacent evergreen types, contributing to denser stands and higher biomass accumulation. This architectural complexity fosters elevated biodiversity relative to purely deciduous forests, as the retention of leaves by a portion of the canopy supports prolonged habitat availability, nutrient cycling, and faunal interactions throughout the year.^[44] Prominent regions for semi-deciduous forests include transitional zones in the Brazilian Amazon, where they form seasonal woodlands between evergreen rainforests and drier deciduous areas; miombo woodlands across central and southern Africa, characterized by semi-deciduous Brachystegia-dominated canopies; and sclerophyllous mixtures in the Iberian Peninsula of Spain and Portugal, blending evergreen oaks with semi-deciduous elements in Mediterranean climates. These areas exemplify the global patchy distribution of such forests in seasonal tropical and subtropical environments.^[45]^[31]^[46] Conservation challenges in semi-deciduous forests are acute, primarily driven by agricultural conversion, logging, and uncontrolled fires, which fragment habitats and reduce resilience. In Latin America, these ecosystems have faced substantial losses since 2000; for instance, the Brazilian Atlantic Forest, which includes semi-deciduous areas, lost approximately 278,000 hectares between 2000 and 2008, exacerbating threats to endemic biodiversity. As of 2025, the Atlantic Forest continues to lose thousands of hectares of mature forest annually despite legal protections. Similar pressures in African miombo woodlands have led to widespread degradation from invasive species and resource extraction.^[47]^[31]^[48] Biodiversity in semi-deciduous forests is notably rich, often hosting 100-450 tree species per hectare depending on regional variation and disturbance levels, surpassing that of fully deciduous counterparts due to the diverse microhabitats created by mixed phenology. Endemic species such as the tonka bean tree Dipteryx odorata in Amazonian semi-deciduous zones exemplify this diversity, contributing to complex food webs and ecological services like seed dispersal and soil stabilization.^[47]^[49]

References

[1]
[PDF] USA-NPN Botany Primer - National Phenology Network
of a growing season; semi–deciduous—semi–persistent and generally staying on the plant and dropping from the plant when under stress; or succulent—thicker.
[2]
WEC 181/UW206: Tropical Hardwood Hammocks in Florida - Ask IFAS
These forests are dominated by tropical trees and shrubs, most of which are evergreen or semi-deciduous, which means they hold leaves during most or all of the ...
[3]
[PDF] generalities in plant function across 97 tropical dry forest tree species
Semi-deciduous encompasses species that partially shed their crown or synchronously drop their entire canopy with a rapid flushing during the dry season. Plant ...
[4]
[PDF] Vegetation Subcommittee FGDC-STD-005 National Vegetation ...
Semi-deciduous Vegetation - Associations (usually tropical and subtropical) in which most of the upper canopy trees are drought-deciduous and many of the ...
[5]
Beyond leaf habit: generalities in plant function across 97 tropical ...
Jun 25, 2021 · Evergreen species include the species that retain a full canopy during the entire year. Semi-deciduous encompasses species that partially shed ...Introduction · Species Selection And Plant... · Plant Traits
[6]
Leaf habit differentiation explains trait tradeoffs across savanna ...
(2021) also observed an intermediate leaf habit called semi-deciduous, which partly sheds leaves without an obvious leafless stage during the dry season in ...Leaf Habit Differentiation... · 2. Materials And Methods · 2.1. Study Site And Plant...
[7]
Semi-Deciduous - an overview | ScienceDirect Topics
Semi-deciduous refers to trees that shed some of their leaves during the dry season when water resources are limited, resulting in a shorter growing season ...
[8]
https://pmc.ncbi.nlm.nih.gov/articles/PMC11112140/
[9]
Abscission in plants: from mechanism to applications - PMC - NIH
Aug 9, 2024 · Plant hormones play a crucial role in regulating plant abscission and their crosstalk has long been a subject of research interest. A widely ...
[10]
Leaf adaptations of evergreen and deciduous trees of semi‐arid and ...
Feb 22, 2013 · Drought avoidance has been associated with deciduous species, which are able to drop their leaves when soil water potentials decline during dry ...
[11]
https://nph.onlinelibrary.wiley.com/doi/10.1111/nph.17187
[12]
Leaf turgor loss point shapes local and regional distributions of ...
Jan 15, 2021 · Drought-deciduous tree species lose their leaves at the driest time ... semi-deciduous and facultatively deciduous species, as well as ...
[13]
Leaf economics of deciduous and evergreen plants - PubMed Central
Jun 5, 2025 · With a high Aarea and Narea, CO2 diffusion and leaf anatomy become limiting factors. Because CO2 diffusion is very low in the liquid phase ...Missing: semi- | Show results with:semi-
[14]
Chapter 4 Leaf phenology and radiation extinction | The medfate ...
The difference between semi-deciduous and deciduous plants is that the former retain dead leaves in the plant until next season unfolding period, so that ...
[15]
[PDF] Adaptive Significance of Evergreen vs. Deciduous Leaves
Seasonal drought favors deciduous leaves, infertile soils favor long-lived evergreens. Evergreens dominate tropical rainforests, while deciduous trees are in ...
[16]
All About Semi-Evergreen and Semi-Deciduous Shrubs and Trees
Jan 14, 2014 · Plants may be labeled as semi-deciduous if their periods of dormancy are dependent on certain weather conditions—they will shed leaves in ...Missing: ecology | Show results with:ecology
[17]
Types of Trees: Deciduous, Evergreen, and Semi-Deciduous - Quizlet
Oct 17, 2024 · Semi-Deciduous Plants: These plants lose their older leaves as new growth begins, often in response to environmental changes, but do not lose ...
[18]
Photoprotection of evergreen and drought-deciduous tree leaves to ...
Dec 11, 2013 · The leaf lifespan in the evergreen trees (Hopea and Syzygium) was ∼2 years. The leaf lifespan in the semi-deciduous tree (Irvingia) was 1 year, ...
[19]
Leaf life span differs from retention time of biomass and nutrients in ...
Aug 5, 2003 · They thus include evergreen species with a mean leaf life span over 3 years (Pinus spp. and T. baccata), evergreen species with a leaf life ...
[20]
Leaf habit drives leaf nutrient resorption globally alongside ... - BG
Sep 26, 2024 · Overall, we find higher NRE in deciduous plants (65.2 % ± 12.4 %, n=400) than in evergreen plants (57.9 % ± 11.4 %, n=551), likely associated ...
[21]
Leaf life span and nutrient resorption as determinants of plant ...
This is not surprising since mature evergreen leaves have lower nutrient con- centrations than deciduous leaves (Aerts, 1996), but the proportion of nutrients ...
[22]
The Mediterranean evergreen Quercus ilex and the semi-deciduous ...
Plants may exhibit some degree of acclimation after experiencing drought, but physiological adjustments to consecutive cycles of drought and re-watering ( ...<|control11|><|separator|>
[23]
Lessons from a tropical deciduous shrub species: leaf fall can play a ...
Nov 11, 2022 · In this study, we investigated the effect of irrigation and early defoliation on the triggering of leaf budding of the deciduous species ...
[24]
[PDF] Flexible Drought Deciduousness in a Neotropical Understory Herb
Jul 21, 2022 · ... water conservation traits and deep roots to survive in seasonal ... villosissimus' semi-deciduous drought response has not previously ...
[25]
Primer Tropical savannas and dry forests - ScienceDirect.com
May 7, 2018 · Of course, there are formations that are transitional between tropical rain forests and tropical dry biomes, which are often semi-deciduous ...Missing: irregular | Show results with:irregular
[26]
Deciduousness in tropical trees and its potential as indicator of ...
Aug 9, 2025 · In tropical forests, deciduousness is an outcome of integrated effect of drought, tree characteristics and soil moisture conditions and thus ...
[27]
Resource Remobilization Efficiency Varies With Plant Growth Form ...
Effective resource resorption from senescing leaves is an important strategy for internal nutrient recycling (Killingbeck, 1996), which reduces plants' ...<|control11|><|separator|>
[28]
Flexible drought deciduousness in a neotropical understory herb
Jul 21, 2022 · ... water conservation traits and deep roots to survive in seasonal ... Further, the suite of traits that function together in the semi-deciduous ...
[29]
[PDF] Regional Forest Types - Tropical Dry Forests - Clemson OPEN
The highest deforestation rates occur in Africa, even though population densities in dry forests there are only one-fifth of those in Asia. This trend ...
[30]
A global overview of the conservation status of tropical dry forests
Feb 6, 2006 · To analyse the conservation status of tropical dry forests at the global scale, by combining a newly developed global distribution map with spatial data ...<|control11|><|separator|>
[31]
(PDF) Global-Overview-of-Tropical-Dry-Forests - ResearchGate
More than half of the forests are distributed in the Americas, with other portions in Africa, Eurasia, Australia, and Southeast Asia.
[32]
Africa's semi-deciduous forests are under threat - Cirad
Sep 7, 2023 · A semi-deciduous forest is characterized by the partial shedding of leaves in the dry season. In West Africa, these forests are situated in a ...
[33]
Species: Prosopis glandulosa - Forest Service
Sprouting from lateral roots is common [81]. These adaptations allow honey mesquite to retain most leaves in all but the most severe droughts.
[34]
[PDF] Mesquite and Palo Verde Trees for the Urban Landscape
Mesquites are deciduous, and foliage is shed in response to cold temperatures and prolonged drought conditions. The different species of mesquite can be ...
[35]
[PDF] Champ monograph (michelia champaca) - Panda.org
Crown Michelia champaca is a medium or large evergreen or semi-deciduous tree with a crown conical to cylindrical. The old leaves abscised before the arrival ...
[36]
[PDF] Magnolia champaca, Champak - IUCN Red List
Aug 31, 2012 · Magnolia champaca is an evergreen or semi-deciduous, medium-tall sized emergent canopy tree up to. 50 m tall and 200 cm in diameter. It is ...
[37]
How to Plant, Grow, and Care for Ice Cream Bean - Epic Gardening
Apr 4, 2025 · In cooler weather, the tree will drop its leaves and become semi-deciduous. ... Inga trees can grow upwards of 60 feet, so you may need to let the ...
[38]
SYNCHRONIZATION OF SEXUAL REPRODUCTION OF TREES ...
Inga spp., Cecropia, spp., Carua pennata,. Swartzia panamensis, Carapa ... The dropping of leaves during the dry season can certainly contribute to the.
[39]
Ylang Ylang tree - Garden Phenology
New leaf flushes can occur after flowering cycles or in response to rainfall. Leaf drop is minimal but may increase during drought or stress.Missing: deciduous | Show results with:deciduous
[40]
Plagianthus - Trees and Shrubs Online
A genus of 2 deciduous to semi-evergreen species endemic to New Zealand, one a divaricating shrub, the other a normally heteroblastic small to medium tree.
[41]
Plagianthus regius - Future Forests
Out of stockThe toothed leaves are triangular, and pale green, and often persist on the tree in mild winters; unusually, the leaves are small when the tree is young, but ...
[42]
Definitions - Forest - Convention on Biological Diversity
Nov 30, 2006 · Deciduous/semi-deciduous broadleaf forest. Natural forests with > 30% canopy cover, below 1200m altitude in which between 50-100% of the canopy ...
[43]
Taller trees, denser stands and greater biomass in semi-deciduous ...
Aug 15, 2016 · For a given diameter, trees tended to be taller in the semi-deciduous forest than in the evergreen forest, as already reported between moist and ...
[44]
Semideciduous Forest - Cristalino Lodge
The semideciduous forest has partial foliage loss in dry months, transitions between deciduous and evergreen forests, is not as tall, has dense ground level, ...Missing: structure | Show results with:structure
[45]
Iberian Sclerophyllous and Semi-Deciduous Forests | One Earth
The flagship species of the Iberian Sclerophyllous and Semi-Deciduous Forests ecoregion is the spanish imperial eagle. Image credit: Wildscreen Exchange.<|control11|><|separator|>
[46]
Brazil's Atlantic Forest (Mata Atlântica) - The Tropical Rainforest
Semi-deciduous forests: inland forests where trees drop their leaves in the dry season. Atlantic Dry Forests: one of the furthest inland, this forest is a ...Geography Of The Mata... · Biodiversity Profile Of The... · Protected Areas In The Mata...
[47]
Dipteryx odorata (tonka bean) | CABI Compendium
odorata is a large sized tree characteristic and exclusive to Amazonian pluvial forests (preferentially in non-flooded forests, less common in lowland forests).