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Common sandpiper

The Common sandpiper (Actitis hypoleucos) is a small, migratory wading bird belonging to the family Scolopacidae, characterized by its slender body, measuring approximately 19–21 cm in length with a wingspan of 32–35 cm, featuring a rich brown upperparts, white underparts, and a distinctive white wing bar visible in flight.^[1]^[2] It is closely related to the North American spotted sandpiper (Actitis macularia), sharing similar teetering gait and bobbing tail behavior while foraging along water edges, but distinguished by its lack of spotting in non-breeding plumage and a slightly longer, more wedge-shaped tail.^[2] This species breeds in scattered pairs during May to June across northern and central Europe, Siberia, and parts of Asia, favoring habitats near fast-flowing rivers, upland streams, lakeshores, and coastal areas with gravelly or stony substrates.^[1] It undertakes long-distance migration, often at night overland on a broad front, wintering in a wide range from sub-Saharan Africa and the Indian subcontinent to Southeast Asia, Australasia, and even southern Pacific islands, where it occupies a variety of wetland environments including pools, marshes, estuaries, and rice fields up to 4,000 meters elevation.^[1] The global population is estimated at 3.2–5.2 million mature individuals, with a stable trend, and it is classified as Least Concern by the IUCN due to its extensive range and lack of severe threats, though localized disturbances from recreation and habitat alteration pose minor risks in some breeding areas.^[1] Behaviorally, the common sandpiper is primarily solitary outside the breeding season, forming small flocks only during migration, and forages diurnally by running rapidly along shorelines to peck at aquatic insects, crustaceans, and small fish.^[1]^[2] Its flight is distinctive, involving alternating bursts of rapid wingbeats and gliding on stiff wings, often low over water, accompanied by a high-pitched, piping call of "twee-wee-wee."^[2] Although rare in the Western Hemisphere, with vagrant records primarily in western Alaska during migration, it is a common sight in the Old World, playing a key ecological role in controlling invertebrate populations in wetland ecosystems.^[1]^[2]

Taxonomy

Etymology

The common name "sandpiper" refers to the bird's characteristic behavior of foraging along sandy shores while producing a piping vocalization, with the term first appearing in English in the 1670s as a compound of "sand" and "piper."^[3] The descriptor "common" highlights its broad distribution and relative abundance across Eurasia, Africa, and parts of Australasia.^[4] The binomial scientific name Actitis hypoleucos originates from Ancient Greek roots: the genus Actitis derives from aktitēs, meaning "coast-dweller," which itself stems from aktē for "coast," alluding to the species' affinity for shoreline habitats.^[5] The specific epithet hypoleucos combines hypo-, meaning "under" or "beneath," with leukos, meaning "white," describing the bird's pale underbelly.^[5] The species was first formally described by Swedish naturalist Carl Linnaeus in 1758 under the name Tringa hypoleucos in the tenth edition of Systema Naturae.^[6] In 1811, German zoologist Johann Karl Wilhelm Illiger established the genus Actitis to include A. hypoleucos and its North American congener, the spotted sandpiper (A. macularius), distinguishing them from the broader Tringa genus based on morphological traits like tail shape and wing patterns.^[7] This reclassification marked a key shift in sandpiper nomenclature, separating the Actitis species from other tringine waders.^[7]

Systematics

The common sandpiper (Actitis hypoleucos) belongs to the family Scolopacidae, encompassing sandpipers and their allies, and is placed within the order Charadriiformes.^[8] The genus Actitis contains only two species: the common sandpiper (A. hypoleucos) and the spotted sandpiper (A. macularius), with the common sandpiper recognized as monotypic and lacking any subspecies.^[8] Molecular phylogenetic analyses confirm that A. hypoleucos and A. macularius are sister species, having diverged approximately 2 million years ago.^[9] Hybridization between the common sandpiper and the green sandpiper (Tringa ochropus) is rare but has been documented.^[6] This classification is upheld by major authorities, including the IOC World Bird List (version 12.1, 2022) and subsequent updates through 2025.^[10]

Description

Morphology

The common sandpiper (Actitis hypoleucos) is a small shorebird with a body length of 19–21 cm, a wingspan of 32–35 cm, and a weight of 40–60 g, with an average of approximately 55 g.^[11]^[12]^[13] It possesses a slim, streamlined build suited to its agile foraging habits, featuring relatively long wings and a long tail relative to its body size. The bill is short and straight, typically measuring 19–24 mm in length, while the legs are short and colored dark yellowish to greenish.^[4]^[14]^[13] Sexual dimorphism is minimal, though females are slightly larger than males, particularly in wing length, with thresholds around 111–117 mm serving as reliable indicators for sex determination based on linear measurements.^[4] Juveniles closely resemble non-breeding adults in overall structure but exhibit distinct features including a scaled appearance on the upperparts due to buff tips and bars on the feathers, along with more heavily barred wing coverts.^[11]^[15] Morphological adaptations include short legs that facilitate the species' characteristic bobbing motion while wading in shallow waters, and a flexible, slender bill designed for probing into mud and sand to extract invertebrate prey.^[16]^[14] The common sandpiper is similar in overall size to its North American sister species, the spotted sandpiper (Actitis macularius).^[4]

Plumage and variation

The common sandpiper (Actitis hypoleucos) exhibits distinct seasonal plumages that aid in camouflage and identification. In breeding plumage, adults display greyish-brown upperparts with fine dark streaks and chevrons on the mantle and scapulars, contrasting with unmarked white underparts and a white supercilium extending into a supraloral line.^[4] The breast sides may show subtle dark streaking, while the legs are greenish-yellow, and a prominent white wing bar is visible in flight, formed by the inner webs of the secondaries.^[17] Unlike its North American congener, the spotted sandpiper (A. macularia), the common sandpiper lacks spotting on the breast in any plumage.^[4] During the non-breeding season, the plumage becomes duller overall, with grey-brown upperparts showing reduced streaking and less distinct chevrons on the mantle.^[17] The breast streaking fades significantly or disappears, resulting in smaller, less prominent patches on the sides, while the white underparts remain unmarked and the white wing bar persists in flight.^[4] This subdued coloration provides better adaptation to winter habitats in tropical and subtropical regions.^[17] The species undergoes two main molts annually. Adults complete a full post-breeding (pre-basic) molt from September to February, primarily in winter quarters, replacing all body feathers, flight feathers, and tail to attain non-breeding plumage.^[17] This is followed by a partial pre-breeding (pre-alternate) molt from February to May, involving the body feathers, tertials, and some wing coverts to acquire breeding plumage before northward migration.^[17] Juveniles perform a complete or near-complete post-juvenile molt from August to December, transitioning from their initial plumage to one resembling non-breeding adults.^[17] There is no sexual dimorphism in plumage, though females are slightly larger than males overall.^[4] Juvenile plumage closely resembles non-breeding adults but features buff fringes and speckles on the upperpart feathers, along with more heavily barred greater wing coverts and a crisper, tidier appearance due to fresher feathers.^[17]^[4] As adults age, feather wear becomes evident on the upperparts during the breeding season, accentuating the dark streaks and chevrons.^[17]

Range and habitat

Breeding range

The common sandpiper (Actitis hypoleucos) breeds across a vast expanse of the Palearctic region, spanning Eurasia from western Europe—including Iceland, the United Kingdom, and Ireland—eastward through Scandinavia, central and eastern Europe, and Russia to Japan, extending northward to approximately 70°N in suitable northern latitudes.^[18]^[1] This breeding distribution encompasses diverse temperate and subarctic environments, with the species favoring freshwater systems over coastal areas during this period.^[1] Within this range, the bird selects habitats characterized by fast-flowing rivers, clear lake shores, and tundra regions featuring gravel or shingle banks, where it can access open, sparsely vegetated ground for nesting.^[1] Nests are typically constructed on the ground near water edges, often on shingle, moss, or pebbly substrates, deliberately avoiding dense vegetation to minimize predation risks and facilitate rapid access to foraging sites.^[1]^[12] These preferences extend to sheltered inland wetlands, small ponds, and riverine meadows, providing the open terrain essential for territorial displays and chick rearing.^[1] Recent studies indicate declines in some European breeding populations, attributed to habitat changes and disturbances.^[18] Densities are notably higher in Fennoscandia, reflecting optimal conditions in the region's upland rivers and boreal wetlands.^[11] The species occupies elevations from sea level up to 4,000 m in mountainous areas, such as the Alps and Central Asian ranges.^[1]

Winter range

The common sandpiper (Actitis hypoleucos) is a full migrant with no overlap between its breeding and winter ranges, undertaking long-distance journeys from northern temperate and subtropical zones to southern wintering grounds. Western breeding populations primarily overwinter in sub-Saharan Africa, ranging from Senegal and Angola in the west to Kenya and South Africa in the east, where they are widespread and common across diverse wetland systems. Eastern populations migrate to southern Asia, including India, Malaysia, Indonesia, and the Philippines, and extend into Australasia, with key concentrations in northern and western Australia, Papua New Guinea, and occasionally Melanesia.^[1]^[19]^[4] In its winter range, the species favors a broader array of habitats than during breeding, showing a stronger preference for coastal and lowland environments. It commonly occupies mudflats, estuaries, and mangroves along coastlines, as well as inland sites like riverbanks, small pools, ditches, marshes, and tidal creeks; rice fields serve as supplementary foraging areas, particularly in Asia, providing shallow-water edges amid agricultural landscapes. Unlike its more upland breeding preferences, winter habitats emphasize soft substrates near water for accessibility, though it generally avoids large open mudflats in favor of sheltered, vegetated margins.^[1]^[16]^[19] Population densities are notably high in African wintering areas, with birds abundant from West Africa southward, though a few individuals may linger year-round at inland sites in southern Africa. In Asia and Australasia, densities vary but are common in northern Australia and coastal Southeast Asia. Vagrants occasionally appear outside the typical range, including records in the Americas (such as the USA), Fiji, and New Zealand.^[1]^[20]^[19]

Migration

Patterns and routes

The common sandpiper (Actitis hypoleucos) is a long-distance migrant, with breeding populations across Europe and Asia undertaking extensive journeys to wintering grounds in sub-Saharan Africa, southern Asia, and Australasia.^[1] European birds typically follow the African-Eurasian flyway, migrating south-southwesterly across continental Europe toward West Africa, often via routes passing through central France, Iberia, or the Mediterranean region before reaching destinations in countries like Guinea-Bissau or further south to Zimbabwe.^[12]^[21] In contrast, eastern Asian populations utilize the East Asian-Australasian flyway, traveling through key areas such as China, Korea, Japan, Hong Kong, and the Yellow Sea wetlands en route to wintering sites in Southeast Asia, Wallacea, New Guinea, and Australia.^[19]^[11] Important stopover sites along these routes include the wetlands of the Yellow Sea, which serve as critical refueling areas for East Asian birds during migration, and Palau in Micronesia, where hundreds may congregate briefly on the eastern edge of the flyway.^[22] The species employs a combination of navigational strategies, including magnetic cues for orientation and visual landmarks for route correction, as observed in related shorebirds and supported by broader avian migration studies.^[23] During flight, common sandpipers achieve speeds of up to 60 km/h, facilitating efficient travel over both land and sea.^[24] Vagrancy occurs outside typical routes, with records in North America primarily in western Alaska during migration periods, occasional sightings in Hawaii, and rare appearances in northern South America, likely resulting from overshoots or storm displacements.^[2]^[22]^[25]

Timing and phenology

The common sandpiper undertakes long-distance migration between its Eurasian breeding grounds and wintering areas in sub-Saharan Africa, southern Asia, and Australasia, with spring migration typically commencing in February to May from winter quarters. Birds depart overwintering sites in Africa as early as late February, progressing northward and reaching European and Asian breeding grounds between April and June, with peak arrivals in late April to early May in western Europe.^[12]^[21] Autumn migration follows breeding and begins in July to September, as adults initiate post-breeding movements shortly after fledging, often preceding juveniles by 10-15 days. In western populations, median departure from breeding sites occurs around early July, with passage through Europe peaking in August to October; eastern populations, such as those stopping over in Palau, may depart northward as early as May. The full migratory journey for individuals spans 4-6 weeks, involving stopovers, while some birds cover over 10,000 km round-trip along major flyways.^[12]^[26]^[19] Breeding phenology aligns with arrival, with clutch initiation averaging late May (range: early May to early July) across the range, enabling synchronization with peak invertebrate availability. Phenological shifts have been observed, with earlier spring arrivals and breeding starts linked to warmer conditions in preceding winters and springs; however, such changes vary regionally and may not fully compensate for trophic mismatches.^[12]^[27]

Behavior

Locomotion and displays

The common sandpiper exhibits a distinctive terrestrial locomotion characterized by a constant teetering or bobbing motion of the head and tail while walking or running along shorelines and water edges.^[4] This behavior, often described as a tail-flicking or wagtail-like gait, aids in maintaining balance on slippery surfaces and is prominent during foraging, where the bird darts quickly with jerky movements to probe mud or pick prey from rocks.^[16]^[28] Occasionally, individuals wade into shallow water or swim short distances, powered by wings and feet, particularly when pursuing prey or evading threats despite lacking webbed feet.^[4]^[29] In flight, the common sandpiper employs stiff, bowed wings with rapid, shallow wingbeats, producing a characteristic bowing motion that flashes a prominent white wing bar.^[4]^[29] This low, skimming flight path, typically close to the water surface, facilitates efficient travel during foraging or migration, though longer distances involve higher altitudes and stronger beats with intermittent glides on arched wings.^[28]^[16] Courtship displays occur primarily in the breeding season, with males performing aerial circuits over potential territories, involving high-speed chases with the female, glides on arched wings, and occasional wing salutes where one or both wings are thrown outward.^[28]^[4] These visual signals, often accompanied by trilling calls in a "flight song," help establish monogamous pairs and defend nesting areas.^[29] Territorial aggression includes similar wing salutes as warnings before chasing intruders, while near nests, parents employ distraction tactics by flying away to lure predators from chicks or young.^[4]^[16] Outside breeding, the species is typically solitary or forms small flocks, with aggressive chases at boundaries during migration or wintering.^[28]

Vocalizations

The common sandpiper produces a variety of vocalizations that serve key communicative roles, including alarm signaling, contact maintenance, and reproductive behaviors. These sounds are typically high-pitched and whistled, reflecting the bird's small size and habitat preferences near watercourses. Vocalizations are most frequent during the breeding season and migration, when individuals are more interactive. The primary alarm call is a high-pitched, ringing series of notes often rendered as seep-seep-seep or swee-wee-wee, delivered rapidly when the bird is disturbed or threatened, such as by predators or intruders near the nest.^[30] An alternative alert call is a thin, sharp, drawn-out wheet, used to warn nearby individuals of potential danger.^[30] These calls function primarily in territory defense and anti-predator responses, helping to deter threats and coordinate escape behaviors among flock members or family groups.^[31] The contact call, a piping twee-wee-wee or tsee-wee-wee, is commonly given during foraging or in flight, maintaining spatial awareness between individuals without indicating alarm.^[31] This call facilitates social cohesion, particularly in loose wintering flocks or during migration stopovers. For parent-offspring communication, adults use softer variants of these contact calls to guide or reassure chicks, while juveniles emit quieter, less assertive versions to solicit food or attention.^[4] The song consists of a melodious, trilling series of descending whistles, often described as a rapid repetition of three-note phrases like chiu-wi-wit or a continuous piping trill, performed primarily by males but occasionally by females during aerial display flights.^[29] This vocalization plays a central role in mate attraction and territory advertisement, accompanying the bird's characteristic stiff-winged flight displays over breeding grounds.^[11] Songs are more elaborate and frequent in the early breeding season (May–June), becoming subdued post-pairing.^[16] Overall, vocal output intensifies during breeding, with calls becoming louder and more persistent to assert dominance and pair bonds, whereas non-breeding individuals are generally quieter, relying more on visual cues.^[4] Geographic variations in call pitch and tempo may occur across the Palearctic range, though structural similarities persist.^[32]

Ecology

Breeding

The common sandpiper breeds from May to July in its northern ranges, forming monogamous pairs that maintain a nesting territory for the season. Pairs typically arrive at breeding sites in mid-April, with courtship displays involving aerial chases and ground pursuits leading to egg-laying by early May.^[33]^[12]^[34] Nests are constructed as shallow scrapes on the ground, often lined with grass, leaves, or small stones, and situated within 50 meters of water in areas of sparse vegetation such as riverbanks or lake edges. Nests are typically placed in open habitats with low cover near freshwater bodies to provide concealment while allowing quick access to foraging areas. Females lay a clutch of 3 to 5 eggs, most commonly 4, which are pear-shaped, measuring 35–38 mm in length, and feature a creamy or buff background with reddish-brown or dark brown spots.^[35]^[4]^[36] Both parents share incubation duties for 21–22 days, with the male often taking the longer night shift and the female the day; eggs are laid at 1–2 day intervals, resulting in asynchronous hatching over 1–3 days. Chicks are precocial and mobile upon hatching, leaving the nest within hours, though they are initially brooded and fed by both parents. When threatened, the young may cling to a parent's body to be carried to safety, particularly near water.^[12]^[36]^[16] Fledging occurs at 25–30 days, by which time the chicks are independent in foraging but may remain with the male parent longer if the female has already departed. Breeding success varies but averages 0.5–1 fledged young per pair, with hatching rates of 85–89% but lower post-hatching survival due to predation by mammals and birds, flooding, and disturbance.^[16]^[35]^[18]

Foraging and diet

The common sandpiper forages primarily along the edges of water bodies, employing a distinctive teetering or bobbing gait while running or walking rapidly to pursue prey. It captures food by pecking at the surface or probing shallow water and soft soil with its bill, often stunning larger items like small crabs against rocks before consumption. This visual hunting strategy allows it to target mobile prey effectively, with birds typically foraging individually or in pairs to minimize competition and predation risk.^[37]^[4] The diet consists mainly of invertebrates, comprising approximately 70% of intake, including insects such as beetles and Diptera larvae, earthworms, crustaceans like amphipods and crabs, polychaete worms, and mollusks; small fish are occasionally consumed, particularly in wetlands. During the breeding season, the diet shifts toward terrestrial prey, with Diptera making up around 30%, beetles 23%, and earthworms 12% based on fecal analysis. In winter and on migration, it favors more aquatic items, such as amphipods (47%), polychaetes (39%), and juvenile crabs (6%) in estuarine habitats.^[38]^[39]^[40] Feeding occurs throughout the day but peaks at dawn and dusk, with intake rates averaging about 75 mg dry weight per minute in winter, supporting high energy needs for migration. Prey selection is opportunistic yet selective for high-profitability items, such as larger polychaetes and small crustaceans, while avoiding less energetic options. By foraging in low-density groups, common sandpipers reduce interspecific competition with other waders for shared resources along shorelines.^[39]^[4]

Conservation

Population trends

The global population of the common sandpiper (Actitis hypoleucos) is estimated at 3,230,000–5,190,000 mature individuals.^[1] In Europe, the breeding population is approximately 1,030,000–1,490,000 pairs, representing a significant portion of the global total.^[1] Overall, the species' population trend is considered stable, based on assessments from the Waterbird Populations Portal.^[1] However, regional variations exist; in the United Kingdom, numbers have declined by 21% from 1995 to 2023, with a steady decrease since the 1990s that may be linked to climate change effects on wintering grounds.^[12] Populations in Asia appear stable, contributing to the species' overall resilience.^[1] The common sandpiper is monitored through programs such as the British Trust for Ornithology (BTO) Breeding Bird Survey and the African-Eurasian Waterbird Agreement (AEWA), which track breeding densities and migration patterns via ringing and census data.^[12] Ringing recoveries indicate a typical lifespan of around 8 years, with the maximum recorded age exceeding 15 years.^[12] Regionally, the species is classified as vulnerable in some Australian states, such as Victoria, due to localized habitat pressures, despite its global status.^[41] The International Union for Conservation of Nature (IUCN) lists the common sandpiper as Least Concern worldwide as of 2025.^[1]

Threats and measures

The Common sandpiper faces several key threats throughout its range, primarily related to its migratory lifestyle. Habitat loss and degradation along flyways, particularly reclamation of intertidal mudflats in the Yellow Sea region, reduce critical stopover sites for refueling during migration, leading to increased energy expenditure and higher mortality rates among migrants.^[19]^[42] Similarly, hunting pressure in wintering areas of southern Asia and sub-Saharan Africa contributes to adult mortality, with birds occasionally targeted for food or sport.^[19] Climate change exacerbates these risks by altering wintering habitats, such as through drier conditions in Africa and southern Asia that diminish wetland availability and prey resources, potentially increasing overwinter survival challenges.^[12] On breeding grounds in Europe and northern Asia, nest predation by mammals and avian predators accounts for a significant portion of reproductive losses, while periodic flooding in riverine habitats destroys nests and chicks.^[43] Pollution from agricultural and industrial runoff further impacts prey populations, such as aquatic invertebrates, reducing foraging efficiency at key sites.^[44] To address these threats, the species is protected under the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA), which promotes habitat safeguards and limits hunting across its range.^[1] Numerous Ramsar-designated wetlands along migration routes, including stopover sites in the East Asian-Australasian Flyway, provide legal protection against development and support habitat restoration efforts.^[45] The East Asian-Australasian Flyway Partnership coordinates international initiatives to conserve shared habitats and monitor migratory shorebirds, benefiting the Common sandpiper through reduced disturbance and pollution controls. In Europe, ongoing monitoring via national breeding bird surveys tracks population health and informs site-specific protections, such as restrictions on recreational activities near nests.^[12] These measures have contributed to population stabilization in protected wetland areas, with no major global declines reported since 2020, supporting the species' overall Least Concern status.^[1]

Human interactions

Cultural significance

In the Nukumanu language spoken on the Nukumanu Islands of Papua New Guinea, the common sandpiper is commonly known as tiritavoi, while matakakoni serves as a taboo name due to its descriptive meaning—"bird that walks a little and then copulates"—alluding to the species' characteristic tail-bobbing behavior. Local informants emphasized avoiding the taboo term, reflecting cultural norms around explicit references to reproduction in naming wildlife. This linguistic distinction highlights the bird's integration into indigenous knowledge systems, where its movements evoke specific behavioral interpretations.^[46] The common sandpiper features occasionally in European literature as a symbol of swift, migratory life along waterways. In William Wordsworth's 1795 poem "Lines Left upon a Seat in a Yew-Tree," the bird is evoked as the "glancing sandpiper," capturing its agile, reflective presence near water and linking it to themes of transience and natural harmony. Such portrayals underscore its role in Romantic-era art and poetry, where it represents adaptability to coastal and riparian environments. In modern contexts, the common sandpiper appears in birdwatching guides and educational materials as an emblem of long-distance migration, emphasizing its reliance on interconnected wetlands across continents. This portrayal supports broader wetland conservation efforts by illustrating the bird's vulnerability to habitat disruption along flyways.

Vagrancy and records

The Common sandpiper is a regular vagrant to western Alaska, where it occurs primarily during spring migration from mid-May to mid-June, with fewer records in autumn from late July to early September.^[11] The species has been documented over 100 times across North America, with notable concentrations in Alaska and more than 50 sightings in California, typically in late September to early November and occasionally in summer.^[47] Vagrants also reach other regions, including rare occurrences in Hawaii and Iceland.^[1] Further south, isolated records exist in South America, such as in Chile, highlighting transoceanic dispersal events.^[48] The first confirmed North American record dates to 1966, when two specimens were collected on St. George Island in the Pribilof Islands, Alaska, on 21 and 24 May.^[49] Subsequent sightings, including the first Aleutian Islands record on 23 May 1971 at Adak, established the pattern of overshoot from Asian breeding grounds.^[50] Records have increased since the mid-20th century, potentially influenced by shifting weather patterns, including stronger storms and climate-driven changes in migration corridors that facilitate vagrancy.^[51] These vagrant occurrences provide valuable insights into long-distance migration dynamics, as demonstrated by ringing recoveries of European-banded individuals traced to wintering sites in sub-Saharan Africa, confirming non-stop flights of up to several thousand kilometers.^[52] Such data from traced birds have helped map connectivity between Eurasian breeding populations and African non-breeding areas, revealing flexible routes via western Europe and Iberia.^[53] Extralimital breeding attempts are exceedingly rare, though vagrants have occasionally prospected suitable habitats in Iceland, where the species does not breed regularly.^[1]

References

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Undergo partial first pre-alternate moult Jan. to early Apr., resulting in breeding plumage similar to adult breeding but slightly duller; then migrate N. Adult.
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Factors Affecting Survival of Common Sandpiper (Actitis hypoleucos ...
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Common Sandpiper - The Australian Museum
The Common Sandpiper is a small sandpiper with a rather long body and short legs. It is grey-brown above and white below, extending up in a pointed shape.
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A few birds remain in southern Africa throughout the austral winter, particularly at inland sites, but relatively fewer overwinter as compared to predominantly ...<|control11|><|separator|>
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Breeding populations from C & W Europe largely take SW routes, while the UK population migrates mainly N-S. W Africa (especially the Gulf of Guinea) is an ...
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The Common Sandpiper is a possibility because it reaches Micronesia as a regular winter visitor (Baker 1951) and western Polynesia and Alaska as a vagrant ( ...
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Upland Sandpipers traveled long distances during southbound migration and the total length of routes averaged 8,793 km (range = 5,410–10,675 km, n = 6 birds, ...
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The median date of female migration was 6 days earlier than that of males. In the first days of July, males comprised only 22% of common sandpipers caught, ...
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Common sandpiper. –0.38. –0.38. –0.13. –5.9. –0.13. 0.32. 0.22. –1.7. Goldcrest. –0.26 ... There is little information on the effect of climate change on ...
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[31]
Common Sandpiper (Actitis hypoleucos) :: xeno-canto
Bird observations: a Common Sandpiper calls, sitting on the ground, appearing to be agonized by the presence of the recordist. Weather conditions: partly cloudy ...
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Common sandpiper | New Zealand Birds Online
Within the East Asian-Australasian Flyway, the common sandpiper is considered the most widely distributed shorebird species, but the southern limits of its ...
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Common Sandpiper - Sound Approach
Common Sandpipers Actitis hypoleucos calling excitedly as they gain height, turn a wide arc, then head off into the night.
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Not-so-Common Sandpipers - wadertales - WordPress.com
May 4, 2019 · The Breeding Bird Survey** indicates a fall in breeding Common Sandpiper ... Green Sandpiper · Greenshank · Grey Plover · Hudsonian Godwit ...
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Common Sandpiper - Facts, Diet, Habitat & Pictures on Animalia.bio
The Common sandpiper (Actitis hypoleucos) is a small Palearctic wading bird, formally described by the Swedish naturalist Carl Linnaeus in 1758.
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The breeding biology of the Common Sandpiper Actitis hypoleucos ...
The breeding biology of the Common Sandpiper Actitis hypoleucos in the Peak District. P. K. Holland 43 Thornhill, Stockport, Greater Manchester. ,. J. E. ...
[37]
Common Sandpiper Life Cycle: Nest Building To Fledging (and ...
They can be found in a wider range of wetland habitats, including estuaries, mangrove swamps, and even man-made water bodies like sewage farms and irrigation ...What Is A Common Sandpiper... · How Long Does It Take For A... · Eggs
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Common Sandpiper: Foraging behaviour - Bird Ecology Study Group
Dec 17, 2008 · “When it is on the hunt, the Common Sandpiper is more relaxed and controlled, it does not do wild sprints like the Terek, as it walks bobbing ...<|control11|><|separator|>
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Diet, food availability and habitat selection of breeding Common ...
Aug 9, 2025 · The diet of the Common Sandpiper, as studied by faecal analysis, includes a wide variety of prey, the majority of which are of terrestrial ...Missing: foraging | Show results with:foraging
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Diet and prey selection of Common Sandpiper Actitis hypoleucos ...
Aug 6, 2025 · Results: During the winter the diet of this wader species was mainly composed of three marine invertebrates, the amphipod Talitrus saltator ( ...
[41]
(PDF) Fish eating by Common Sandpiper - Academia.edu
The study reveals that fish was consumed by Common Sandpipers, despite previous records excluding fish from their diet. This was observed in a wetland in ...
[42]
Actitis hypoleucos : Common Sandpiper | Atlas of Living Australia
Tringa aurita Latham, 1801 synonym. AFD. Tringa hypoleucos Linnaeus, 1758 synonym. AFD. Common Name, Source. Common Sandpiper preferred Australia Australia. ALA ...<|separator|>
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Conserving unprotected important coastal habitats in the Yellow Sea
In 2008–2018, 43 shorebird species were recorded along this coastline, including 12 globally threatened or 'Near Threatened' species. In terms of number of ...Original Research Article · 2. Material And Methods · 2.3. Satellite Tracking
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Factors Affecting Survival of Common Sandpiper (Actitis hypoleucos ...
Jul 10, 2024 · Our research showed that mammalian and avian predation accounted for 51% of losses and flooding for 49% of losses. The negative impact of floods ...
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[PDF] Threatened Species Assessment Actitis hypoleucos
Jun 9, 2021 · Common Sandpipers nest mainly in cool temperate to low arctic regions of the northern hemisphere, usually near freshwater habitats (especially ...
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https://www.birdsnz.org.nz/wp-content/uploads/2021/12/Notornis_51_2_91.pdf
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[PDF] Notornis June 04.indd - Birds New Zealand
Birds of North Solomons atolls. 101. *Common sandpiper (OJ) [Tiritavoi]. *Grey-tailed tattler (OJ) [Tekapo]. *Turnstone (OJ) [Teikaika or Kanakapo]. On Ontong ...<|control11|><|separator|>
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Common Sandpiper Actitis hypoleucos - eBird
Small bicolored sandpiper which often bobs its tail in a distinctively wagtail-like manner. Plain brown with white underparts; distinguished from bulkier ...Identification · Range Map · Media
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[PDF] Punta Rasa, South America's first vagrant trap? - Neotropical Bird Club
Even less is known concerning vagrancy in the continent, and, due to the lack of baseline data for many areas, vagrants can be mistaken for range extensions.
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(PDF) Birds New to North America and the Pribilof Islands, Alaska
This is the first positive North American record. Common Sandpiper (Actitis hypoleucos).--Two specimensof this sandpiper were taken on St. George Island, one ...
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New Information on Asiatic Birds in the Aleutian Islands, Alaska - jstor
there are five other Alaska records of the species. Actitis hypoleucos. Common Sandpiper. Since the first Aleutian record of the species at Adak on 23 May ...
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When Birds Get Lost, Space Storms May Be to Blame | Audubon
Jun 5, 2023 · New analysis of 60 years of bird banding data shows that vagrancy increases during periods of geomagnetic disturbance.
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Migration patterns and morphometrics of Common Sandpipers ...
In this paper, we present results from an analysis of 62 years' trapping and ringing data of autumn-migrating Common Sandpipers at Ottenby Bird Observatory, ...
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[PDF] Non-breeding areas and timing of migration in relation to weather of ...
It would therefore seem that. 289 common sandpipers have the potential to accomplish their migration from northern Europe to. 290. West Africa in a single ...Missing: banded | Show results with:banded