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Cretaceous–Paleogene extinction event

The Cretaceous–Paleogene (K–Pg) extinction event was a catastrophic mass extinction that occurred approximately 66 million years ago, marking the boundary between the Cretaceous period of the Mesozoic era and the Paleogene period of the Cenozoic era, and resulting in the loss of roughly 76% of Earth's species, including all non-avian dinosaurs.^[1]^[2] This event, one of the "Big Five" mass extinctions in the geologic record, profoundly reshaped global ecosystems by eliminating dominant marine and terrestrial groups such as pterosaurs, mosasaurs, plesiosaurs, ammonites, and many planktonic foraminifera and nannoplankton, while sparing smaller organisms like mammals, birds, crocodilians, and certain plants that later diversified.^[2]^[3] The extinction's rapidity—unfolding over mere years to millennia—distinguishes it from gradual environmental changes, with fossil records showing abrupt discontinuities at the K–Pg boundary worldwide.^[4] The event's primary trigger was the collision of a ~10–15 km diameter asteroid with the Yucatán Peninsula in present-day Mexico, forming the Chicxulub impact crater, a 150–200 km wide structure buried beneath sediments and dated precisely to 66.04 ± 0.05 million years ago through argon-argon dating of impact melt rock.^[4]^[1] This hypervelocity impact released energy equivalent to billions of atomic bombs, vaporizing rock and ejecting debris that heated the atmosphere, ignited global wildfires, and generated tsunamis up to 1 km high, while the resulting dust cloud blocked sunlight for months to years, halting photosynthesis and causing a collapse in food chains.^[4]^[3] Compounding factors included massive volcanic eruptions from the Deccan Traps in India, which released climate-altering gases over millennia bracketing the boundary, though stratigraphic and geochemical evidence indicates the impact as the dominant, synchronous cause of the extinction pulse.^[4] Diagnostic evidence for the impact includes a thin, global layer of iridium-rich clay at the K–Pg boundary—iridium being rare on Earth but abundant in asteroids—along with shocked quartz crystals, nickel-rich spinels, and tektite-like microkrystites formed under extreme pressures and temperatures.^[3]^[4] Recovery was uneven: marine ecosystems at the impact site showed microbial rebound within years, but full biodiversity restoration took millions of years, with surviving lineages like mammals undergoing adaptive radiations that define modern faunas.^[1] This extinction underscores the vulnerability of complex life to extraterrestrial perturbations, informing ongoing research into planetary defense and paleobiological resilience.^[4]

Overview

Event definition

The Cretaceous–Paleogene (K–Pg) extinction event, also referred to as the Cretaceous–Tertiary (K–T) extinction in older literature, represents the fifth major mass extinction in Earth's history and occurred approximately 66 million years ago.^[5]^[6]^[7] This event demarcates the boundary between the Cretaceous period of the Mesozoic era and the Paleogene period of the Cenozoic era, signifying a profound transition in Earth's biota.^[8] The extinction eradicated an estimated 75–80% of all species across marine and terrestrial realms, including the complete demise of all non-avian dinosaurs.^[1]^[9] It stands out for its abrupt and global scope, with fossil records showing a sharp faunal turnover at the boundary layer.^[10] Key geological signatures of the event include a widespread iridium anomaly—indicating extraterrestrial material—and shocked quartz grains in sediments, both concentrated precisely at the K–Pg boundary worldwide.^[11]^[12] The abrupt nature of these changes was first recognized by 19th-century geologists examining stratigraphic discontinuities and faunal absences, such as the sudden disappearance of dinosaurs noted by early paleontologists.^[13] A pivotal advancement came in 1980 with the Alvarez hypothesis, which proposed that a large asteroid impact triggered the catastrophe, supported by the iridium enrichment as evidence of extraterrestrial delivery.^[11]^[14]

Geological and biological significance

The Cretaceous–Paleogene (K–Pg) extinction event profoundly reshaped Earth's biosphere, extinguishing approximately 75% of species and allowing only about 25% to survive, including key lineages ancestral to modern birds (Neornithes) and placental mammals that later diversified into primates and other groups.^[15] This massive biotic turnover ended the dominance of non-avian dinosaurs, which had constrained mammalian evolution for over 150 million years, thereby releasing ecological opportunities for surviving small-bodied mammals to radiate into diverse niches.^[16] The event's biological legacy is evident in the survival of adaptable, often generalist taxa, such as small theropod dinosaurs that evolved into birds and insectivorous mammals, setting the stage for avian and mammalian dominance in the Cenozoic. Geologically, the K–Pg boundary marked a pivotal transition from the warm, greenhouse conditions of the Late Cretaceous to cooler Paleogene climates, driven by the Chicxulub impact's atmospheric soot and associated global cooling, which disrupted photosynthesis and initiated an "impact winter."^[17] This climatic perturbation facilitated a major floral shift, with angiosperms rising to dominance in terrestrial ecosystems; for instance, pollen records from Colombian sites show angiosperm representation increasing from roughly 48% in the Maastrichtian to 84% in the early Paleocene, coinciding with the development of closed-canopy rainforests featuring diverse angiosperm growth forms.^[18] Such changes in vegetation structure enhanced habitat complexity, influencing subsequent biogeochemical cycles and soil formation across continents. In the long term, the K–Pg extinction paved the way for the Cenozoic Era, often termed the "Age of Mammals," as placental orders and crown-group diversification accelerated post-boundary, filling vacated niches and driving innovations in locomotion, diet, and sociality.^[16] Relative to other mass extinctions, the K–Pg was more ecologically selective than the Permian–Triassic event, which eradicated over 80% of species and disrupted nearly all functional groups in marine ecosystems, whereas the K–Pg preserved most functional diversity (losing only 7% of marine bivalve functional groups) despite comparable taxonomic losses around 64–76%.^[19] This selectivity enabled faster ecosystem restructuring, with mammalian and avian radiations contributing to modern biodiversity patterns within 10 million years. The K–Pg event offers critical analogies for understanding rapid biodiversity crises, particularly the ongoing anthropogenic extinction. A 2025 analysis indicates that the ongoing anthropogenic extinction episode, while concerning, currently falls short of the devastation caused by the K–Pg event, though its eventual magnitude will depend on human responses, with projections suggesting recovery could take millions of years if trends persist.^[20]

Stratigraphy and Chronology

K-Pg boundary characteristics

The Cretaceous–Paleogene (K-Pg) boundary is marked by a distinctive global stratigraphic layer, typically a thin clay bed enriched in iridium (Ir), a rare siderophile element with concentrations reaching up to 30 parts per billion (ppb) in the boundary clay, far exceeding typical crustal abundances of less than 0.1 ppb. This iridium anomaly, first identified in Italian pelagic limestones, has been documented worldwide and is attributed to extraterrestrial material dispersed from an impact event.^[12] Associated with the iridium-rich clay are impact-derived markers such as shocked quartz grains, nickel-rich spinels, and silicate spherules formed from condensed vaporized rock, with the latter often altered to clay minerals in marine settings.^[21] In proximal sites, these include tektites and microtektite glass fragments from impact melt, exhibiting vesicular textures and compositions matching Chicxulub crater ejecta. The global stratotype section and point (GSSP) for the K-Pg boundary is located at El Kef, Tunisia, where the boundary is defined at the base of a 1–3 mm thick, rust-colored ferruginous clay layer containing the iridium peak, overlying marly limestones and underlying reddish clays with palynological evidence of the boundary.^[22]^[23] This section preserves a complete marine record, including deep-sea foraminiferal assemblages that show abrupt changes across the boundary clay. In terrestrial settings, such as the Hell Creek Formation in the western United States, the boundary appears as a similar clay layer within floodplain sediments, often capped by a coal seam in the overlying Paleocene Tullock Member, with the iridium anomaly and spherules integrated into the local stratigraphy.^[24] Additional features in boundary sections include a prominent "fern spike" in pollen and spore records, characterized by a sudden dominance of fern spores (up to 80–100% of assemblages) immediately above the clay layer, reflecting rapid colonization by disturbance-tolerant pioneer vegetation in post-impact ecosystems. Charcoal fragments and soot particles within or just above the boundary clay indicate widespread wildfires, with elevated black carbon levels suggesting biomass burning that contributed to atmospheric loading.^[25] Recent drilling expeditions, including the 2016 IODP-ICDP Expedition 364 to the Chicxulub crater and subsequent ocean coring efforts analyzed in 2025, have provided high-resolution profiles of boundary layers in marine sites, revealing enhanced details on soot from vaporized organic matter and sulfate aerosols from evaporated seawater and evaporites, which were lofted into the stratosphere and deposited globally within the iridium-enriched clay.^[26]^[27] These findings, from sites like the Gulf of Mexico and Atlantic cores, show that sulfate contributions were lower than previously modeled, with soot playing a more significant role in short-term climate forcing, while confirming the synchroneity of the boundary at approximately 66 Ma.^[12]

Precise dating methods

The precise dating of the Cretaceous–Paleogene (K-Pg) boundary has been established through advanced radiometric techniques applied to volcanic ash beds and impact-related deposits interlayered with boundary sediments, providing absolute ages with uncertainties below 0.1 million years. The primary method involves uranium-lead (U-Pb) geochronology on zircon crystals extracted from these ash beds, utilizing chemical abrasion-isotope dilution-thermal ionization mass spectrometry (CA-ID-TIMS) to minimize lead loss and achieve high precision. A seminal study on zircons from ash layers in the Hell Creek Formation, Montana, yielded an interpolated age of 66.021 ± 0.024 Ma for the boundary, directly tying it to the iridium enrichment layer that marks the event.^[28] Supporting this, ⁴⁰Ar/³⁹Ar dating of sanidine crystals from the same ash beds has provided complementary high-resolution results, with an age of 66.052 ± 0.008 Ma for a layer immediately above the iridium anomaly at the Denver Basin site, Colorado, confirming consistency across methods.^[29] These techniques are calibrated against the iridium layer and align the boundary within geomagnetic polarity chron C29r, a reversed polarity interval spanning approximately 66.5 to 65.5 Ma. Astronomically tuned cyclostratigraphy further refines the chronology by correlating sedimentary cycles in marine sections, such as limestone-marl couplets at Zumaia, Spain, to Milankovitch orbital parameters, yielding an age of approximately 66.04 Ma for the boundary and supporting the radiometric framework with sub-10 kyr resolution.^[30] Calibration points include the Deccan Traps volcanism, dated via U-Pb on zircons from interbedded sedimentary ashes to a main eruptive phase between 66.27 and 65.52 Ma, with peak activity around 66.1–66.0 Ma overlapping the boundary timing.^[31] Recent advances as of 2025 integrate rhenium-osmium (Re-Os) dating of organic-rich boundary clays, achieving sub-10 ka precision through isochron analyses that confirm the synchroneity of the Chicxulub impact and the primary extinction pulse, as evidenced by Os isotope excursions in Danish fish clays aligned to 66.05 Ma.^[32]

Extinction duration and phases

The Cretaceous–Paleogene (K-Pg) extinction event unfolded over a temporal framework that included a prelude of environmental and biotic perturbations, an acute crisis at the boundary, and a protracted recovery. Approximately 300,000 years prior to the K-Pg boundary, a period of Maastrichtian warming linked to Deccan Traps volcanism induced biotic stress, evidenced by shifts in planktic foraminiferal assemblages and reduced diversity in certain marine groups.^[33]^[34] This pre-boundary phase featured gradual declines in some taxa, such as ammonites, where species richness decreased by up to 20% in Antarctic sections during the late Maastrichtian due to regional ecological pressures.^[35] However, continental records, including those from New Mexico, indicate that non-avian dinosaurs maintained high diversity and regional endemism until within roughly 340,000 years of the boundary, with no overarching decline.^[36] The core extinction manifested as an instantaneous crisis at the K-Pg boundary itself, constrained by U-Pb dating to approximately 66 million years ago, with the acute phase lasting years to decades following the Chicxulub impact.^[6] This rapid event eliminated about 75% of marine species and 60% of terrestrial vertebrate species, marking a selective purge particularly affecting large-bodied taxa across ecosystems.^[37] Fossil records reveal temporal heterogeneity: terrestrial sections, such as those in the Hell Creek Formation, show abrupt turnovers synchronous with the iridium-rich boundary layer, while deep-sea cores exhibit staggered declines in benthic foraminifera and calcareous nannoplankton, with some pre-boundary perturbations extending tens of thousands of years earlier due to ocean acidification and productivity shifts.^[38]^[39] Recovery proceeded in pulses spanning 10³ to 10⁵ years, characterized by initial opportunistic blooms—such as fern spikes in terrestrial settings and microbial proliferations in marine environments—followed by gradual rediversification of surviving lineages.^[40] Selectivity in timing was pronounced, with large-bodied organisms facing near-immediate collapse from impact-related disruptions, whereas certain marine groups, buffered by deeper habitats, experienced delayed declines over decades to centuries.^[41]^[42] This phased structure underscores the event's complexity, blending prolonged precursors with a cataclysmic finale and uneven biotic rebound.

Extinction Patterns

Microbiota and fungi

The Cretaceous–Paleogene (K-Pg) extinction event profoundly impacted marine microbiota, particularly calcareous nannoplankton, which form the base of oceanic food webs through primary production. Coccolithophores, key producers of these microscopic calcite plates, suffered near-total eradication, with over 90% of species going extinct at the boundary due to the combined effects of darkened skies, ocean acidification, and disrupted nutrient cycling from the asteroid impact.^[43] This collapse halted calcification and export productivity in surface waters, leading to a "strangelove ocean" phase where surviving primary producers were limited. In contrast, certain resilient groups like dinoflagellates exhibited opportunistic responses; their cysts, preserved in sediments, show a marked increase in abundance immediately above the K-Pg boundary, indicating a temporary bloom of bloom-forming species adapted to low-light and nutrient-stressed conditions before gradual diversification resumed.^[1] This pattern underscores the heterogeneous impact on marine unicellular life, where photosynthetically dependent taxa declined sharply while heterotrophic or mixotrophic forms temporarily dominated. Freshwater and terrestrial microbiota faced similar disruptions to algal communities, with primary producers like green algae and early diatom-like forms experiencing reduced diversity and abundance due to prolonged "impact winter" conditions that suppressed photosynthesis across aquatic and soil environments.^[44] In lakes and rivers, this led to collapses in planktonic algal blooms, altering the base of aquatic food webs and promoting shifts toward bacterial dominance in nutrient-poor waters. Fungi, particularly saprotrophic species, proliferated dramatically in the aftermath, as evidenced by a global "fungal spike" in sediments—a thin layer of exceptionally high fungal spore and hyphal abundance at or just above the K-Pg boundary, signaling a dominance of decomposition processes amid widespread plant die-off.^[45] This surge, observed in both terrestrial and marginal marine deposits, reflects fungi's role in breaking down vast quantities of dead vegetation, thereby facilitating nutrient recycling in darkened, post-impact ecosystems where autotrophic life struggled. Ectomycorrhizal fungi, symbiotic with surviving woody plants, demonstrated enhanced persistence compared to non-mycorrhizal groups, aiding host nutrient uptake from decomposing litter during the crisis. Overall, microbiota exhibited lower extinction rates than macroscopic life, with prokaryotic diversity losses estimated below 20% in many lineages, though eukaryotic microbes like nannoplankton saw higher turnover.^[9] Opportunistic bacteria, including sulfate-reducers, thrived in expanding anoxic zones created by stratified waters and reduced oxygen solubility, filling niches left by collapsed algal productivity. Recent analyses (2024–2025) of wildfire-impacted soils analogize this resilience, showing fungi's tolerance to soot deposition and heat, which mirrors the global firestorms and atmospheric particulates at the K-Pg boundary that favored decomposer guilds.^[46]

Marine invertebrates

The Cretaceous–Paleogene (K-Pg) extinction event exhibited stark selectivity among marine invertebrates, with planktonic forms suffering far higher losses than benthic ones, reflecting vulnerabilities tied to habitat, mobility, and dependence on surface productivity. Planktonic groups, which rely on open-ocean conditions and primary production, experienced near-total collapse, while benthic communities in deeper waters showed greater resilience due to more stable environments and access to refractory organic matter. This pattern underscores the event's disruption of the marine food web from the base upward, sparing generalist bottom-dwellers but decimating specialized drifters. Among planktonic invertebrates, ammonites and belemnites—key cephalopod groups—underwent near-total extinction synchronous with the boundary, with no surviving species crossing into the Paleogene. Radiolarians, siliceous planktonic protists, lost approximately 95% of their species diversity across the K-Pg, as evidenced by global deep-sea records showing abrupt turnover in polycystine assemblages. Planktonic foraminifera fared slightly better but still suffered about 75% genus-level extinction, disproportionately affecting photosymbiotic species that hosted algal partners and depended on light-dependent productivity, while non-symbiotic forms had higher survival rates.^[47]^[48]^[49]^[50] In contrast, benthic marine invertebrates displayed lower overall extinction rates, estimated at 30-50% for deep-sea infauna such as benthic foraminifera and protobranch bivalves, which benefited from minimal disruption in abyssal oxygen and carbon flux. However, shallow-shelf communities were devastated, with the entire rudist bivalve clade—dominant reef-builders in tropical carbonates—wiped out, alongside inoceramid bivalves, which had already declined but vanished completely at the boundary. Opportunistic polychaetes, like opportunistic annelids in soft sediments, and certain resilient mollusks survived in refugia, thriving post-event due to their tolerance for low oxygen and organic enrichment.^[51]^[9]^[52]^[47] Extinction intensities varied regionally, with higher losses in tropical latitudes where warm, oligotrophic waters amplified impacts on carbonate-dependent and photosymbiotic taxa, compared to more moderate declines at higher latitudes. This latitudinal gradient is evident in bivalve and brachiopod records, where tropical assemblages lost up to 70% of genera versus 40-50% in temperate zones. Trophic structure further influenced selectivity, with herbivorous and primary consumers—such as suspension-feeding rudists and photosymbiotic foraminifera—hit hardest due to the collapse of phytoplankton bases, while detritivores and scavengers endured better. Recent 2025 International Ocean Discovery Program (IODP) drilling in the Gulf of Mexico and Caribbean reveals pulsed extinction phases in benthic records, correlating with episodic productivity crashes from impact fallout and Deccan volcanism, including a sharp post-impact drop in export flux lasting millennia.^[53]^[9]^[19]^[26]

Terrestrial invertebrates

The Cretaceous–Paleogene (K-Pg) extinction event had a moderate impact on terrestrial insect orders, with estimates of species-level losses around 40-50% based on declines in insect-feeding damage on fossil leaves.^[54] This extinction was particularly evident in specialized herbivorous groups, where damage types such as leaf mines and galls dropped sharply from 16% to 4% of leaves at the boundary.^[55] Social hymenopterans like bees and ants showed greater resilience; while some bee lineages experienced significant turnover near the boundary due to disruptions in plant-pollinator relationships, crown-group bees and ants survived and diversified rapidly in the early Paleogene, buffered by their generalist foraging and social structures.^[56]^[57] Among other terrestrial arthropods, myriapods such as millipedes and arachnids like spiders demonstrated high resilience, with no significant family-level declines recorded across the event.^[58] In contrast, certain beetle families, particularly dung-feeding scarabaeines, suffered notable losses linked to the collapse of large herbivore populations and associated habitat alterations following the extinction.^[59] Extinction patterns were global yet uneven, with greater impacts in forested ecosystems where plant diversity plummeted, reducing food resources for herbivorous and detritivorous forms.^[55] Fossil evidence for these patterns derives primarily from amber inclusions, which preserve detailed arthropod assemblages spanning the latest Cretaceous and earliest Paleogene, revealing continuity in resilient groups like ants and spiders while showing gaps in specialized lineages.^[60] Coprolites from Cretaceous deposits further document insect diets and communities, including termite remains that indicate stable detritivore roles pre- and post-boundary.^[61] Selectivity favored social insects and ecological generalists, which maintained broad diets and nesting behaviors adaptable to disrupted environments; recent analyses highlight how post-impact wildfires and global cooling likely advantaged burrowing species by providing insulated refugia amid surface devastation.^[57]^[62] This plant die-off exacerbated food scarcity for many herbivores but enabled opportunistic generalists to persist.^[55]

Fish and amphibians

The Cretaceous–Paleogene (K-Pg) extinction event resulted in substantial losses among teleost fish, with estimates indicating that 50 to 90% of species went extinct, though family-level extinction was lower at around 10%. Open-ocean species were disproportionately affected, including large predatory groups such as the ichthyodectids, which dominated pelagic ecosystems but vanished entirely at the boundary due to disruptions in marine food webs and environmental stressors like darkened skies from atmospheric dust. In contrast, freshwater teleosts exhibited greater resilience, with lower extinction rates and evidence of survival in continental deposits, likely owing to buffered habitats less exposed to global oceanic perturbations.^[63]^[44]^[64] Chondrichthyans, including sharks and rays, experienced relatively low overall extinction rates of less than 10% at the family level, attributed to their occupation of stable predatory and scavenging niches that persisted through the crisis. Extinction increased gradually with habitat openness, reaching up to 45% among open-ocean forms, while freshwater and nearshore species showed near-zero losses, highlighting the protective role of coastal and riverine environments. Recent analyses confirm no significant pre-boundary declines in chondrichthyan diversity, underscoring the abrupt nature of the event's impact on this group.^[44] Amphibians suffered minimal direct species losses, estimated at around 10%, with frogs and salamanders achieving global survival across the boundary, as evidenced by continuous fossil records in North American Hell Creek Formation sites. Larval stages, however, faced heightened vulnerability from freshwater acidification triggered by atmospheric CO₂ release and sulfate aerosols, potentially disrupting development in pond and stream habitats, though adult ectothermy and burrowing behaviors aided overall persistence. Family-level extinction was negligible, with no documented turnover in caudatan lineages.^[65]^[66]^[44] Extinction patterns among fish and amphibians displayed clear selectivity by size and habitat, favoring small-bodied, estuarine, and freshwater forms that could exploit refugia amid collapsed marine productivity; larger, pelagic species, conversely, succumbed to trophic cascade effects from invertebrate prey disruptions. A 2025 study of boundary sections in Saskatchewan and Montana revealed no significant pre-impact declines in teleost or amphibian assemblages, reinforcing that losses were synchronous with the asteroid impact rather than protracted.^[67]^[44]

Non-avian reptiles and pterosaurs

The non-avian reptiles and pterosaurs experienced varied extinction patterns during the Cretaceous–Paleogene (K-Pg) event, with complete loss of several major marine clades and significant but differential impacts on terrestrial and semi-aquatic groups.^[44] Marine reptiles underwent total extinction at the K-Pg boundary, including the complete loss of mosasaurs and plesiosaurs, which dominated Late Cretaceous oceans as apex predators and had diversified into diverse ecological niches such as open-water hunting and coastal foraging.^[44] Ichthyosaurs, another key group of fish-like marine reptiles, had already faced an abrupt two-phase extinction earlier in the Cretaceous, associated with reduced evolutionary rates and environmental volatility, leaving no survivors by the Maastrichtian.^[68] This 100% loss of advanced marine reptile lineages was likely driven by the collapse of primary productivity following the Chicxulub impact, which disrupted phytoplankton-based food webs essential to these top predators.^[44] Among crocodyliforms, approximately 80% of species went extinct, with all marine forms such as the thalattosuchians and dyrosaurids completely wiped out, while only basal, terrestrial or freshwater-adapted lineages like early alligatoroids and basal eusuchians survived into the Paleogene.^[69] This selective survival favored smaller, generalist forms capable of exploiting disturbed habitats, reflecting the broader pattern where ectothermic reptiles with flexible diets endured better than specialized marine or large-bodied taxa.^[70] Turtles and lepidosaurs (including squamates like lizards and snakes) suffered moderate losses of 20-40% at the species or genus level, with terrestrial and freshwater species faring better than marine ones; for instance, over 80% of Cretaceous turtle species crossed the boundary, enabling survival of basal cryptodires and pleurodires, while squamates saw a severe but not total decline, with post-event diversification filling vacated niches.^[71] Choristoderes, a group of aquatic reptiles akin to champsosaurs, experienced regional extinction in North America but survived elsewhere in Laurasia, with a single family persisting into the Miocene due to their adaptation to freshwater environments less affected by the global marine disruptions.^[44] Pterosaurs faced complete global extinction at the K-Pg boundary, with no post-boundary fossils documented worldwide, marking the end of their 160-million-year reign as the dominant flying vertebrates.^[72] Recent analyses indicate a late Maastrichtian diversity drop, evidenced by reduced morphological disparity and species richness in the final million years before the event, potentially linked to competition in aerial niches and environmental stressors, though some records suggest high local diversity persisted until the abrupt catastrophe. This extinction opened opportunities for avian birds to occupy flying roles in post-K-Pg ecosystems.^[73]

Non-avian dinosaurs

The Cretaceous–Paleogene (K-Pg) extinction event resulted in the complete extinction of all non-avian dinosaur lineages, including theropods, sauropods, ornithischians, and other groups, with no unambiguous body fossils, eggshells, or nesting traces found in post-boundary strata worldwide.^[6] This total loss is evidenced by the abrupt disappearance of diverse non-avian dinosaur assemblages from formations such as Hell Creek (North America), Lameta (India), and Nemegt (Asia), which persisted until the very end of the Maastrichtian stage.^[6] The absence of any surviving non-avian dinosaurs post-impact underscores the event's severity for this clade, leaving their ecological niches—ranging from large herbivores like hadrosaurs and ceratopsians to apex predators such as tyrannosaurids—entirely vacated in terrestrial ecosystems.^[36] Prior to the K-Pg boundary, non-avian dinosaurs showed no signs of significant global decline, maintaining robust populations and high regional diversity in the final hundreds of thousands of years of the Cretaceous.^[36] Recent geochronological revisions of the Naashoibito Member in New Mexico's San Juan Basin, dated to approximately 66.4–66.0 million years ago, reveal thriving dinosaur communities with endemic species, including late-surviving hadrosaurs and other ornithischians, countering earlier notions of faunal impoverishment.^[36] These findings indicate that non-avian dinosaurs were ecologically stable and partitioned into distinct provincial assemblages right up to the asteroid impact, with no evidence of pre-extinction stress from volcanism or other factors.^[36] The K–Pg extinction exhibited strong size selectivity across vertebrate taxa, with most animals exceeding approximately 25 kg in body mass suffering near-total extinction, while smaller-bodied forms (under ~1 kg) in lineages such as mammals, birds, and squamates had higher survival rates. Late Maastrichtian non-avian dinosaurs lacked a diverse array of small-bodied forms (most exceeding 1 kg), rendering the group particularly vulnerable to the environmental perturbations of the K-Pg event.^[74]^[75] This selectivity amplified the clade's demise, as even mid-sized herbivores and carnivores, integral to food webs, were unable to persist. Fossil evidence for the final non-avian dinosaurs is concentrated in uppermost Maastrichtian units like the Hell Creek Formation of the Western Interior Basin, where last occurrences of taxa such as Tyrannosaurus rex, Triceratops horridus, and hadrosaurs cluster just below the boundary clay.^[76] A ceratopsian brow horn discovered 13 cm beneath the iridium-rich K-Pg boundary in southeastern Montana represents one of the stratigraphically highest in situ non-avian dinosaur remains, refuting claims of a multi-meter "fossil gap" and confirming their presence until the impact's immediate prelude.^[76] Bone beds in these deposits, including those at the Tanis site in North Dakota, contain disarticulated dinosaur elements—such as a Thescelosaurus leg with preserved skin—intermingled with impact markers like tektites, shocked quartz, and ejecta spherules, directly linking the animals' deaths to the Chicxulub bolide.^[77] These assemblages demonstrate that non-avian dinosaurs were active and abundant in floodplain environments moments before the catastrophe.^[77]

Birds and mammals

The Cretaceous–Paleogene (K-Pg) extinction event resulted in the complete loss of major archaic bird groups, including enantiornithines and hesperornithines, which were diverse and widespread in the Late Cretaceous but failed to cross the boundary into the Paleogene.^[73] In contrast, the neornithine crown group, representing the lineage leading to modern birds, experienced relatively low extinction rates of approximately 10-20%, allowing multiple lineages to persist through the crisis.^[73] This survival is attributed to ecological selectivity favoring generalist species adapted to open or ground-based habitats, as global forest collapse following the Chicxulub impact eliminated arboreal niches occupied by many stem birds.^[78] Evidence for avian continuity includes fossil tracks and skeletal remains documented on both sides of the K-Pg boundary in formations such as Hell Creek, indicating uninterrupted presence of small, shorebird-like neornithines.^[73] Mammals likewise exhibited high survivorship across the K-Pg boundary, with roughly 90% of small-bodied species enduring the event, particularly favoring insectivorous multituberculates and therian mammals that were nocturnal, burrowing, or otherwise adaptable to disrupted environments.^[79] Large-bodied forms, including early marsupials, were largely absent prior to the extinction and did not factor into post-boundary recovery, underscoring the dominance of diminutive, opportunistic therians and multituberculates in the immediate aftermath.^[80] Key traits such as burrowing behavior and dietary flexibility enabled these survivors to exploit insect resources amid the collapse of larger food webs, with fossil burrows and coprolites preserving evidence of such habits across the boundary in North American sites. Recent 2025 analyses of Late Cretaceous mammal assemblages confirm no significant pre-impact decline in diversity or abundance, reinforcing that the extinction acted as a selective filter rather than a culmination of long-term stress.^[36] The shared emphasis on endothermy, small size, and versatile foraging among surviving birds and mammals highlights convergent selective pressures during the crisis, setting the stage for subsequent size increases and ecological expansions in the Paleogene, though full diversification occurred later.^[81]

Terrestrial plants

The Cretaceous–Paleogene (K–Pg) extinction event resulted in substantial losses among terrestrial plants, with overall diversity declining by approximately 45% in tropical regions based on extensive fossil records from South America. Macrofloral analyses indicate extinction rates ranging from 57% to 66% at the boundary, concentrated in a geologically instantaneous pulse, while palynological records show more moderate losses of around 30% in miospore taxa in North America.^[82] Gymnosperms, including cycads and conifers, were disproportionately affected, with their representation in floras dropping sharply from 2.5% to 0.4% in the Paleocene, reflecting higher vulnerability to the environmental perturbations compared to angiosperms. In contrast, angiosperms exhibited remarkable lineage-level resilience, with phylogenetic analyses revealing no evidence of a mass extinction at the K–Pg boundary and stable extinction rates across geological time, despite regional species turnover up to 75%.^[83] A prominent feature of the post-boundary flora was the "fern spike," a transient dominance of fern spores in sedimentary records from sites worldwide, including North America and New Zealand, signaling their role as opportunistic pioneers in devastated landscapes.^[82] This spike, documented through palynological evidence, lasted briefly before angiosperm recovery, highlighting ferns' ability to exploit disturbed environments with wind-dispersed spores and minimal light requirements. Regional patterns varied, with severe losses in the southern hemisphere—such as in Patagonia, where leaf fossil assemblages indicate substantial vegetation collapse—but also faster initial rebounds compared to northern sites, where palynofloral extinctions reached 30% or more. Tropical forests experienced widespread disruption, yet angiosperm diversity began to recover within a few million years, eventually comprising 84% of Paleocene floras in some areas.^[84] Trophic interactions were altered by the event, with declines in specialized insect herbivores and pollinators contributing to shifts in plant reproduction and damage patterns; for instance, leaf herbivory persisted at high levels (>50%) but with reduced host specificity in the aftermath. Angiosperm survival has been attributed to pre-extinction adaptations such as efficient insect pollination and versatile reproductive strategies, enabling them to outcompete gymnosperms in the post-K–Pg world, as confirmed by 2023 phylogenetic studies.^[83] Evidence from pollen and macrofossil records underscores these patterns, with abrupt diversity drops at the boundary followed by selective recovery.^[82] Additionally, elevated charcoal abundance in boundary sediments points to widespread wildfires, likely amplified by the impact's thermal pulse, which further stressed plant communities and facilitated ecosystem turnover.

Causal Mechanisms

Asteroid impact hypothesis

The asteroid impact hypothesis posits that a large extraterrestrial body struck Earth approximately 66 million years ago, serving as the primary catalyst for the Cretaceous–Paleogene (K-Pg) mass extinction. This event is linked to the Chicxulub impact structure, where a carbonaceous-type asteroid estimated at 10–15 km in diameter collided with the Yucatán Peninsula in Mexico, excavating a transient cavity that rapidly collapsed to form a final crater roughly 180 km in diameter.^[85]^[86] The impact released energy equivalent to billions of atomic bombs, vaporizing rock and ejecting material globally within minutes, with the initial collision and crater excavation completing in about 8 minutes.^[86] Direct evidence for the Chicxulub impact derives from extensive drilling campaigns spanning the 1990s to the 2020s, which penetrated the crater's peak ring and revealed impact melt rock and shocked minerals indicative of extreme pressures exceeding 5–45 GPa.^[87] These include shocked quartz grains with planar deformation features, preserved in ejecta deposits, confirming hypervelocity impact origins.^[86] Complementing this, a thin global ejecta layer at the K-Pg boundary contains elevated iridium concentrations (up to more than 100 ppb in spinel-bearing spherules within peak-ring cores) from the asteroid's chondritic composition, alongside Ni-rich spinels formed during high-temperature condensation of vaporized material.^[12]^[88] This iridium anomaly, detected worldwide, marks the boundary's stratigraphic signature and aligns precisely with the extinction horizon.^[12] The immediate aftermath triggered cascading global disruptions. The impact generated massive tsunamis with wave heights exceeding 100 m near the site, propagating across oceans and inundating continents up to 1,000 km inland.^[89] Re-entering ejecta and thermal radiation ignited widespread firestorms, evidenced by soot and charcoal layers in K-Pg sediments spanning multiple continents, burning up to 70% of global forests in hours to days.^[90] These fires lofted soot into the atmosphere, exacerbating an "impact winter" where stratospheric dust and sulfate aerosols blocked sunlight, causing a rapid global cooling of 5–10°C that persisted for years to decades and collapsed primary productivity.^[91] Recent high-fidelity geophysical and geochemical models, incorporating 2025 data from refined crater stratigraphy, further affirm the Chicxulub event's exact synchroneity with the K-Pg boundary, dated to within decades via osmium isotopes and sedimentation rates.^[12]^[92] Hypotheses of multiple contemporaneous impacts remain unsupported, as no other craters of comparable age and scale have been confirmed, with structures like Popigai predating the boundary by millions of years.^[93]

Volcanic activity role

The Deccan Traps represent a large igneous province consisting of extensive flood basalt eruptions in what is now west-central India, with a preserved volume of approximately 500,000 km³ of basaltic lava extruded over roughly 1 million years, from about 66.4 to 65.5 Ma.^[31] The eruptive activity intensified in distinct pulses, with the main phase peaking between 66.1 and 66.0 Ma, releasing substantial volumes of volcanic gases including sulfur dioxide (SO₂) and carbon dioxide (CO₂).^[31] These emissions contributed to initial global warming through CO₂-induced greenhouse effects, followed by potential short-term cooling from sulfate aerosols formed by SO₂ oxidation in the atmosphere.^[31] The volcanic outgassing from the Deccan Traps is linked to pre-boundary climate instability during the late Maastrichtian, including a long-term warming of about 3°C driven primarily by elevated atmospheric CO₂ levels.^[94] This warming episode, known as the Latest Maastrichtian Warming Event, overlapped with the onset of major Deccan eruptions around 250 kyr before the Cretaceous–Paleogene (K–Pg) boundary and likely exacerbated environmental stress through mechanisms such as acid rain from sulfuric acid deposition and ocean acidification from dissolved CO₂.^[94] Such conditions could have disrupted marine carbonate systems and terrestrial ecosystems, contributing to gradual biotic turnover prior to the boundary.^[31] Recent studies from 2024 and 2025 indicate that Deccan Traps volcanism played a secondary role in the K–Pg extinction through cumulative environmental perturbations like toxicity and habitat alteration, with limited global dispersion of volcanic aerosols and toxins, but insufficient to drive the full mass extinction event.^[95] Mercury isotope records from globally distributed deep-marine sediments reveal elevated Hg concentrations starting ~200–300 kyr before the boundary, consistent with Deccan sourcing, yet the anomalies are smaller than anticipated for the erupted volume, suggesting limited global dispersion of volcanic aerosols and toxins.^[95] High-precision ⁴⁰Ar/³⁹Ar dating confirms alignment of peak Deccan activity with the K–Pg boundary at ~66.04 Ma, indicating temporal overlap with the extinction but no causal connection to the iridium enrichment layer, which reflects extraterrestrial input.^[96] Overall, while Deccan volcanism imposed chronic stress that may have amplified the extinction's severity, its gradual nature contrasts with the abrupt forcing required for the scale of biotic collapse observed.^[95]

Other environmental factors

The Maastrichtian regression, a major global sea-level fall of approximately 50–100 meters during the latest Late Cretaceous, significantly reduced the extent of shallow continental shelf habitats, which were critical for diverse marine communities.^[97] This regression exposed vast areas of seafloor, leading to habitat compression and environmental stress that contributed to pre-extinction biodiversity declines, with estimates suggesting losses of around 10% in affected shelf-dependent taxa due to reduced habitable area and altered biogeochemical conditions.^[98] Such changes intensified selective pressures on benthic and neritic species, setting the stage for heightened vulnerability at the Cretaceous–Paleogene boundary. Oceanic anoxia expanded in the Late Cretaceous, driven by global warming that stratified water columns and promoted the growth of oxygen minimum zones, resulting in widespread dead zones particularly in semi-restricted basins. Concurrently, hypercapnia—elevated dissolved CO₂ levels—induced a pH drop in surface waters, impairing calcification in marine organisms such as foraminifera and coccolithophores by reducing carbonate ion availability and increasing dissolution risk.^[99] These conditions disproportionately affected calcifying plankton and shelled invertebrates, exacerbating physiological stress and contributing to ecological disruptions independent of acute boundary events. Milankovitch cycles, through periodic variations in Earth's orbital parameters, imposed rhythmic climate instability during the Late Cretaceous, with precession (∼20 kyr) and obliquity (∼41 kyr) cycles modulating insolation and monsoon intensity to drive fluctuations in temperature, precipitation, and sea-surface productivity.^[100] These forcings amplified environmental variability, potentially stressing terrestrial and marine ecosystems by altering habitat suitability and resource availability on millennial scales, thus fostering conditions of chronic instability leading into the boundary interval.

Integrated multiple-cause models

Integrated multiple-cause models for the Cretaceous–Paleogene (K-Pg) extinction emphasize the interaction of prolonged environmental stressors and a sudden catastrophic event, where pre-existing conditions from Deccan Traps volcanism and sea-level regression weakened ecosystems, making them vulnerable to the Chicxulub asteroid impact's lethal effects of prolonged darkness and toxic aerosols.^[6]^[101] In this synergistic framework, Deccan eruptions, spanning hundreds of thousands of years prior to the boundary, induced global warming, acid rain, and ocean acidification, with long-term CO₂ release increasing habitat suitability by approximately 27–32% for some taxa like dinosaurs, though short-term aerosol cooling and other perturbations added environmental stress that overall did not drive extinction-level collapse.^[6] Sea-level regression during the late Maastrichtian further fragmented coastal and shallow marine habitats, exacerbating biotic stress and limiting refugia for many species.^[101] The asteroid impact then acted as the decisive "kill shot," triggering an impact winter with up to 15-26% solar dimming, global cooling of -34.7°C, and sulfate aerosols that halted photosynthesis for months to years, collapsing food chains across latitudes.^[6] Quantitative climate and ecological modeling indicates that the Chicxulub impact alone reduced suitable dinosaur habitats to less than 4% of pre-extinction levels, sufficient to drive non-avian dinosaur extinction, while Deccan volcanism's effects were recoverable and mitigated some cooling but amplified long-term disruptions through CO₂-driven warming of +4.7°C to +8.75°C.^[6] These simulations attribute the majority of the extinction's severity—potentially over 75% of species loss—to the impact's acute perturbations, with volcanism serving as an amplifier rather than the primary driver.^[6]^[95] Central to these models is the "press-pulse" concept, which posits that mass extinctions arise from the overlap of chronic "press" disturbances (e.g., Deccan volcanism's multi-generational climate shifts) and acute "pulse" events (e.g., the Chicxulub impact's rapid mortality), as evidenced by elevated extinction rates in geological stages where such coincidences occurred, including the K-Pg boundary.^[102] Recent 2024-2025 studies reinforce this by rejecting volcanism as the sole cause, showing that Deccan activity predated the impact by ~300,000 years and lacked the intensity to independently trigger global collapse, though it primed ecosystems for the pulse's amplified toll.^[95]^[36] Holistically, these models describe a trophic cascade propagating from the base of food webs—where primary producers like phytoplankton and terrestrial plants suffered from darkness and toxins—to higher levels, extinguishing herbivores and predators in a domino effect, with non-avian dinosaurs particularly vulnerable due to their reliance on stable niches.^[70] Variations occurred globally versus regionally, with a latitudinal selectivity gradient showing lower extinction rates at higher latitudes for some groups like bivalves, possibly due to seasonal refugia or differing stressor intensities, though the impact's effects were broadly synchronous and severe across continents.^[52]^[70]

Immediate Effects and Recovery

Global environmental disruptions

The Cretaceous–Paleogene (K-Pg) extinction event, primarily driven by the Chicxulub asteroid impact, triggered profound global environmental disruptions that reshaped planetary conditions in the immediate aftermath. These changes included rapid alterations in climate, atmospheric composition, ocean chemistry, and terrestrial landscapes, persisting for years to decades and contributing to widespread ecological stress.^[103] One of the most immediate effects was a "nuclear winter"-like scenario, characterized by 1–2 years of near-total darkness due to massive injections of dust and soot into the atmosphere, blocking sunlight and halting photosynthesis globally. This was followed by prolonged cooling lasting 5–10 years, with global average temperatures dropping by up to 8°C, primarily from stratospheric sulfate aerosols formed by the impact's vaporization of sulfur-rich target rocks. These aerosols reflected incoming solar radiation, exacerbating the chill and disrupting seasonal patterns across continents and oceans.^[104]^[105]^[106] Atmospheric chemistry underwent drastic shifts, beginning with a short-term spike in carbon dioxide (CO₂) from the impact's thermal decomposition of carbonates and organic matter, followed by a prolonged drop as disrupted biological productivity reduced carbon sequestration. Concurrently, severe ozone depletion occurred in the stratosphere due to soot heating and water vapor injection, resulting in a marked increase in ultraviolet (UV) radiation reaching the surface, which damaged exposed organisms and ecosystems.^[103]^[107] Oceanic systems experienced acidification, with seawater pH dropping by approximately 0.2–0.5 units due to the influx of sulfuric and carbonic acids from atmospheric fallout, potentially contributing to stress on calcium carbonate shells and halting biocalcification in marine organisms like foraminifera and coccolithophores, though the severity's role in extinctions is debated. This acidification also expanded anoxic zones, particularly in intermediate waters, as reduced oxygen solubility in warmer surface layers and decreased overturning circulation stifled ventilation, leading to toxic sulfide buildup in deeper oceans.^[99]^[108] On land, global wildfires ignited by the impact's fireball and heat pulse consumed vast forests, depositing a layer of soot in the K-Pg boundary clay that is evident worldwide and contributed to the initial darkening and cooling. Recent 2025 modeling further links the extinction of dinosaurian megafauna to subsequent facies shifts in continental sediments, as the loss of these large herbivores altered vegetation structure, soil stability, and sediment deposition patterns, promoting closed-canopy forests over open habitats.^[25]^[17]^[109]

Short-term ecological collapse

The Cretaceous–Paleogene (K–Pg) extinction event triggered a rapid breakdown of global ecosystems, primarily through the collapse of primary production that starved higher trophic levels. The impact-induced atmospheric soot and dust blocked sunlight for 2–10 years, halting photosynthesis in both terrestrial and marine environments and leading to widespread food web disruptions.^[17] This resulted in the formation of expansive "dead zones" in oceans, where marine anoxia expanded due to reduced oxygen production and increased organic matter decay, and in soils, where nutrient cycling stalled amid darkened conditions.^[110] Starvation cascades affected plankton, fish, and larger consumers, with over 90% of calcareous nannoplankton and planktic foraminifera perishing within months to years.^[26] Ecological collapse manifested prominently in the inversion of trophic pyramids, with apex predators succumbing first due to the immediate scarcity of prey. Large carnivorous theropods and marine reptiles like mosasaurs, reliant on abundant herbivores and fish, experienced near-total extinction as their food sources dwindled.^[44] Herbivores followed rapidly, as megaherbivores such as ornithischians declined amid the loss of vegetation, further destabilizing networks with reduced connectance in food webs.^[70] In contrast, detritivores proliferated, with fungi and bacteria booming on the influx of dead organic matter; fungal spores dominated boundary sediments, indicating a temporary shift to decomposition-dominated ecosystems.^[111] Habitat alterations exacerbated the crisis, as the Chicxulub impact generated mega-tsunamis that flooded coastal zones, depositing sediments and disrupting nearshore communities.^[112] Inland, sulfuric acid rain from vaporized sulfate aerosols acidified soils and waters, killing forests and contributing to a brief "fern spike" as opportunistic plants briefly recolonized barren landscapes.^[17] The peak of ecological collapse occurred within less than 10 years, driven by the "impact winter," though marine anoxia persisted for thousands of years in some regions, as evidenced by recent International Ocean Discovery Program (IODP) analyses of post-impact sediments.^[26] Overall, these short-term dynamics decimated ~75% of species, reshaping biosphere structure before gradual stabilization.^[6]

Long-term biotic recovery

Following the Cretaceous–Paleogene (K-Pg) boundary, biotic recovery began with opportunistic taxa that rapidly colonized disturbed environments. In terrestrial settings, ferns and other spore-producing plants dominated the initial post-extinction landscapes for the first several thousand years, forming "fern spikes" indicative of pioneer vegetation in a world depleted of woody angiosperms. Marine ecosystems saw repopulation by disaster species such as small, mobile invertebrates and algae, which proliferated in the nutrient-rich, sun-blocked waters during the first 10,000 years. These early colonizers, often generalist survivors, laid the groundwork for subsequent ecological rebuilding by stabilizing soil and nutrient cycles. Recent analyses confirm angiosperms underwent no mass extinction at the K-Pg, sustaining diversity and ecological roles through the Paleocene. Diversification accelerated in the Paleocene, marking the onset of major adaptive radiations among surviving lineages. Mammals underwent a significant radiation, with archaic groups like condylarths—small, herbivorous ungulate-like forms—diversifying into new niches by the early Paleocene, approximately 66–60 million years ago, filling roles vacated by non-avian dinosaurs. Birds, leveraging their prior resilience, expanded in diversity, with early neornithine lineages adapting to vacant aerial and arboreal habitats. In marine realms, bony fish and some mollusks began radiating, while angiosperms, which survived with low extinction rates, continued diversifying and maintaining forest canopies in the early Paleocene. This phase of opportunistic diversification transitioned ecosystems from low-diversity, r-selected pioneer communities to more structured assemblages. By the Eocene, approximately 10 million years after the event, new ecosystem configurations emerged, resembling modern biomes. Modern-like forests, dominated by diverse angiosperms and gymnosperms, spread across continents, fostering complex food webs that included emerging large herbivores and predators. Coral reefs, devastated initially, recovered through the radiation of scleractinian corals and associated symbionts, achieving structural complexity by the mid-Paleocene. Recent analyses from 2025 indicate that full recovery to pre-K-Pg biodiversity levels was delayed, taking up to 10 million years in many clades, with marine diversity lagging behind terrestrial recovery.^[113]^[114] The long-term legacy of the K-Pg recovery included a shift toward K-selected life history strategies, favoring species with slower reproduction and greater parental investment, which became prevalent in post-extinction guilds. Survivor communities exhibited increased evenness, with no single taxon dominating as in the immediate aftermath, promoting stability in the emerging Cenozoic ecosystems. This recovery not only rebuilt biodiversity but also set the stage for the dominance of mammals and birds in modern biota.

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Teleost fishes across the Cretaceous–Paleogene boundary
To assess potential effects of the mass extinction on the teleost assemblages leading up to and across the K-Pg boundary, we combined three datasets for ...
[68]
Extinction of fish-shaped marine reptiles associated with reduced ...
Mar 8, 2016 · Belemnite extinction and the origin of modern cephalopods 35 m.y. prior to the Cretaceous-Paleogene event. Geology 39, 483–486 (2011) ...
[69]
Climate constrains the evolutionary history and biodiversity ... - Nature
Sep 24, 2015 · The effect of the K/Pg mass extinction on crocodylomorphs has previously been perceived as minor or non-existent, with any extinction temporally ...
[70]
Shifts in food webs and niche stability shaped survivorship ... - Science
Dec 7, 2022 · ... Cretaceous–Paleogene extinction: A short primer. Cretac. Res.57,368–390 (2016). GO TO REFERENCE. Crossref · Web of Science · Google Scholar. 90.
[71]
[PDF] Two major extinction events in the evolutionary history of turtles
Jul 21, 2022 · In the K–Pg mass extinction, Earth experienced a significant loss in biodiversity ... The global effect of the K–Pg mass extinction on turtles ...Missing: squamates | Show results with:squamates
[72]
Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary
Summary of each segment:
[73]
Mass extinction of birds at the Cretaceous–Paleogene (K–Pg ...
Aug 30, 2011 · All available fossil evidence is consistent with a major extinction of archaic birds coinciding with the K–Pg boundary, which may have provided ...
[74]
Mass extinction of lizards and snakes at the Cretaceous–Paleogene ...
Species-level extinction was 83%, and the K-Pg event resulted in the elimination of many lizard groups and a dramatic decrease in morphological disparity. ...Missing: percentage lost
[75]
Dinosaur extinction: closing the '3 m gap' - PMC - NIH
The only Hell Creek studies with precise stratigraphic control [10,11] were able to constrain the fossil-barren zone using a non-avian dinosaur fossil found 1. ...Missing: occurrences | Show results with:occurrences
[76]
A seismically induced onshore surge deposit at the KPg boundary ...
Apr 1, 2019 · Shocked quartz from the warm fireball, launched at angles from an assumed 70° to the limit of the forbidden zone at 79°, begins to reach the ...<|separator|>
[77]
Report Early Evolution of Modern Birds Structured by Global Forest ...
Jun 4, 2018 · We propose a new hypothesis regarding the extinction of stem birds and the survival of crown birds across the K-Pg boundary: namely, that global ...
[78]
Severe extinction and rapid recovery of mammals across the ...
May 11, 2016 · Extinction rates are markedly higher than previously estimated: of 59 species, four survived (93% species extinction, 86% of genera). Survival ...
[79]
Therian mammals experience an ecomorphological radiation during ...
Our results suggest that an ecomorphological diversification of therians began 10–20 Myr prior to the K–Pg extinction event, led by disparate metatherians and ...
[80]
https://pmc.ncbi.nlm.nih.gov/articles/PMC4920311/
[81]
Genome and life-history evolution link bird diversification to the end ...
Jul 31, 2024 · Our results suggest that the end-Cretaceous mass extinction triggered integrated patterns of evolution across avian genomes, physiology, and life history.
[82]
https://doi.org/10.1016/j.gloplacha.2014.07.014
[83]
https://doi.org/10.1098/rsbl.2023.0314
[84]
Leaf fossils show severe end-Cretaceous plant extinction in ...
Jan 4, 2021 · Fossil leaf assemblages from Patagonia, Argentina, suggest that vegetation in South America suffered great losses but rebounded quickly.
[85]
Chicxulub Asteroid Impact Sulfur Release Was Less Lethal In ...
Jan 17, 2025 · The sulfur was deposited around the K-Pg boundary layer in sedimentary profiles all over the world. We used the corresponding change in the ...
[86]
Discovering the Impact Site - Chicxulub Impact Event
A large impact crater that is 65 million years old. Seven North American researchers finally located the impact site on the Yucatán Peninsula.Missing: extinction | Show results with:extinction
[87]
Mapping the Chicxulub Impact Stratigraphy and Peak Ring Using ...
Aug 3, 2021 · Drilling at Chicxulub has confirmed the presence of melt rock and melt-rich breccias in the central basin and annular trough (Hildebrand et al., ...Missing: primary | Show results with:primary
[88]
[PDF] Multiple impacts across the Cretaceous–Tertiary boundary
sedimentation and unencumbered by impact ejecta beyond fine scale fallout (e.g. iridium, Ni-rich spinels). There are numerous K/T boundary sections world-.
[89]
The Chicxulub Impact Produced a Powerful Global Tsunami
Oct 4, 2022 · The Chicxulub crater is the site of an asteroid impact linked with the Cretaceous-Paleogene (K-Pg) mass extinction at ∼66 Ma.
[90]
CU study provides new evidence ancient asteroid caused global ...
Mar 27, 2013 · Geological evidence indicates the asteroid collided with Earth about 66 million years ago and carved the Chicxulub crater in Mexico's Yucatan ...
[91]
Rapid short-term cooling following the Chicxulub impact at the ... - NIH
Impact models suggest that the first hours after the impact were characterized by earthquakes and tsunamis and the so-called “fireball stage,” including an ...Missing: firestorms | Show results with:firestorms
[92]
High‐Fidelity Cretaceous‐Paleogene Boundary Investigations ...
Oct 1, 2025 · Kaskes et al. (2025) recently explored one such K-Pg boundary deposit in the Raton Basin, Colorado using high-resolution geochemical techniques ...
[93]
[PDF] Popigai and Chicxulub craters: multiple impacts and their associated ...
Feb 17, 2025 · Both Chicxulub and Popigai craters are interpreted as double or multiple craters, formed by impact-induced tectonics that triggered impact ...
[94]
30 kyr before the Cretaceous–Paleogene mass extinction - PMC
Dec 18, 2024 · Our records show long-term warming of ~3°C, probably driven by Deccan CO 2 emissions, and reveal a transient (<10 kyr) ~5°C cooling event.
[95]
Deep marine records of Deccan Trap volcanism before the ...
Sep 30, 2024 · For example, multiple studies have determined different ages for the K–Pg boundary; the most recent estimates are 66.052 ± 0.008 Ma based on ...
[96]
Determination of rapid Deccan eruptions across the Cretaceous ...
Jun 10, 2009 · This lava pile was erupted in some 30 major eruptive periods or single eruptive events (SEE), each with volumes ranging from 1000 to 20,000 km3 ...
[97]
https://doi.org/10.1126/science.1116412
[98]
https://doi.org/10.1111/j.1365-2117.2008.00354.x
[99]
Severity of ocean acidification following the end-Cretaceous ... - NIH
Another reason for being particularly interested in the Cretaceous/Paleogene (K/Pg) boundary in the context of OA is that many surface-dwelling calcifiers went ...Missing: hypercapnia | Show results with:hypercapnia
[100]
Milankovitch cycles and their effects on species in ecological and ...
Apr 8, 2016 · Milankovitch cycles thus force an instability of climate and other aspects of the biotic and abiotic environment on time scales much less ...
[101]
Dinosaur extinction can explain continental facies shifts at ... - Nature
Sep 15, 2025 · We hypothesize that the geographically widespread Cretaceous–Paleogene facies shifts were driven by the extinction of dinosaur megafauna. Large- ...
[102]
(PDF) Cretaceous Extinctions: Multiple Causes - ResearchGate
Patterns of extinction and survival were varied, pointing to multiple causes at this time—including impact, marine regression, volcanic activity, and changes in ...
[103]
Press-pulse: a general theory of mass extinction? - GeoScienceWorld
Nov 1, 2008 · To test the hypothesis that the coincidence of bolide impact and flood basalt volcanism is associated with elevated extinction, we required ...
[104]
On transient climate change at the Cretaceous−Paleogene ... - PNAS
Aug 21, 2017 · These fires would have been an efficient extinction mechanism for large land animals (7, 34). ... DS Robertson, et al., K-Pg extinction: ...Missing: insects | Show results with:insects
[105]
Rapid short-term cooling following the Chicxulub impact at ... - PNAS
May 12, 2014 · The mass extinction at the Cretaceous–Paleogene boundary, ∼66 Ma, is thought to be caused by the impact of an asteroid at Chicxulub, present-day ...Missing: sulfate | Show results with:sulfate
[106]
[PDF] Climate model simulations of the effects of the asteroid impact at the ...
Here we explore the longer-lasting cooling due to sulfate aerosols using a coupled climate model. Depending on aerosol stratospheric residence time, global ...
[107]
Energy, volatile production, and climatic effects of the Chicxulub ...
Sep 1, 1997 · A comprehensive analysis of volatiles in the Chicxulub impact strongly supports the hypothesis that impact-generated sulfate aerosols caused ...Missing: Paleogene | Show results with:Paleogene
[108]
Global Effects - Chicxulub Impact Event
There were a series of local and regional effects that occurred immediately upon impact, including fireball radiation, an airblast, earthquakes, and tsunamis.Missing: firestorms | Show results with:firestorms
[109]
The impact of the Cretaceous–Paleogene (K–Pg) mass extinction ...
Jun 1, 2018 · The Cretaceous–Paleogene (K–Pg) mass extinction event 66 million years ago led to large changes to the global carbon cycle, primarily via a decrease in primary ...
[110]
(PDF) Dinosaur extinction can explain continental facies shifts at the ...
Sep 19, 2025 · We hypothesize that the geographically widespread Cretaceous–Paleogene facies shifts were driven by the extinction of dinosaur megafauna. Large- ...
[111]
Rapid ocean acidification and protracted Earth system recovery ...
Oct 21, 2019 · Mass extinction at the Cretaceous–Paleogene (K-Pg) boundary coincides with the Chicxulub bolide impact and also falls within the broader time ...Missing: rates | Show results with:rates
[112]
(PDF) Fungal Proliferation at the Cretaceous-Tertiary Boundary
Aug 6, 2025 · ... (K-Pg) boundary (Figs. 2 and 3)-when a bolide collided with Earth, caused firestorms for days to weeks, and shut down photosynthesis for ...Missing: decomposition | Show results with:decomposition
[113]
Expedition 364 summary - IODP Publications
Dec 30, 2017 · Chicxulub's role in the Cretaceous/Paleogene (K-Pg) mass extinction ... Hydrocode simulations of Chicxulub crater collapse and peak-ring ...