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Iberian ibex

The Iberian ibex (Capra pyrenaica), also known as the Spanish ibex, is a species of wild goat endemic to the Iberian Peninsula in southwestern Europe, where it occupies rugged mountainous habitats from sea level to elevations of 3,480 meters across Spain, Portugal, and reintroduced sites in France. Characterized by its remarkable agility in steep, rocky terrain and prominent backward-curving horns present in both sexes, the ibex is a browser that primarily feeds on leaves, twigs, and acorns from Holm oaks, supplemented by forbs and grasses.^[1] Once critically endangered due to hunting, habitat loss, and disease, its global population has rebounded to over 100,000 individuals as of 2021 through protected areas, reintroductions, and hunting regulations, earning a Least Concern status on the IUCN Red List.^[2] Physically, Iberian ibexes are medium-sized ungulates with a shoulder height of 65–75 cm, body length of 100–140 cm, and weights ranging from 35–80 kg, with males typically larger and possessing longer horns up to 75 cm or more.^[1] Their coat is brownish-gray, providing camouflage in their alpine environments of forested slopes with rock outcroppings, coniferous, and deciduous trees, generally above 800 meters elevation.^[1] Socially, they form herds that exhibit sexual segregation outside the breeding season, with dominant males defending territories and harems during the November–December rut; females give birth to one or two kids in mid-May after a gestation of 161–168 days, and individuals can live 12–16 years in the wild.^[1] The species comprises two extant subspecies—C. p. hispanica in southeastern Spain and C. p. victoriae in central and northern regions—while two others, C. p. pyrenaica and C. p. lusitanica, went extinct in 2000 and the late 19th century, respectively.^[2] Ongoing threats include habitat fragmentation, sarcoptic mange, and competition from introduced ungulates, despite overall population increases as of 2021.^[2] Protected under the EU Habitats Directive^[3] and Bern Convention,^[4] the Iberian ibex serves as a flagship species for conservation in the region, supporting ecotourism and regulated trophy hunting while facing localized declines in some areas.

Taxonomy

Classification

The Iberian ibex, scientifically classified as Capra pyrenaica, belongs to the kingdom Animalia, phylum Chordata, class Mammalia, order Artiodactyla, family Bovidae, subfamily Caprinae, and genus Capra.https://animaldiversity.org/accounts/Capra_pyrenaica/ This taxonomic placement positions it among even-toed ungulates, sharing the Capra genus with other wild goats such as the Alpine ibex (Capra ibex) and the markhor (Capra falconeri), all adapted to rugged, mountainous environments.https://www.iucnredlist.org/species/3787/97218336 The species was first described by Swiss naturalist Johann Rudolf Schinz in 1838, based on a male specimen held at the Zürich Museum along with sketches and descriptions from earlier observations in the Pyrenees Mountains.https://www.gbif.org/species/2441054 Schinz's description, published in Neue Denkschriften der allgemeinen Schweizerischen Gesellschaft für die gesammten Naturwissenschaften, highlighted its distinct morphology from other European goats, emphasizing its presence in the Pyrenean region.https://doi.org/10.1007/978-3-319-65038-8_33-2 The specific epithet "pyrenaica" derives from the Latin term for the Pyrenees Mountains, reflecting the type locality of the original specimens in this range.https://doi.org/10.1007/978-3-319-65038-8_33-2 The common name "Iberian ibex" underscores its endemic distribution across the Iberian Peninsula, distinguishing it from other ibex species in Eurasia and North Africa.https://www.iucnredlist.org/species/3787/97218336 Within the Capra genus, C. pyrenaica is differentiated from the domestic goat (Capra hircus)—its closest domesticated relative—through distinct genetic markers and morphological features.https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0269873 Genetically, Iberian ibex populations form separate clusters with no evidence of introgression from domestic goat gene pools in surveyed groups, as confirmed by microsatellite and SNP analyses.https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0269873 Morphologically, key distinctions include lyre-shaped horns in males that curve backward and inward without pronounced ridges, contrasting with the more variable, often scimitar-like horns in domestic goats; additionally, cranial and postcranial bones exhibit specific osteological traits, such as narrower metapodials and distinct horn core cross-sections, enabling reliable identification in archaeological and ecological contexts.https://turia.uv.es/index.php/sjpalaeontology/article/download/17157/15380/56013

Subspecies

The Iberian ibex (Capra pyrenaica) is traditionally classified into four subspecies based on morphological traits such as coat color and horn structure, though recent genetic analyses have questioned the depth of these distinctions. These include C. p. pyrenaica (Pyrenean ibex), C. p. hispanica (Southeastern Spanish ibex), C. p. victoriae (Gredos ibex), and C. p. lusitanica (Portuguese ibex).^[5] Of these, only C. p. hispanica and C. p. victoriae remain extant, while the other two are extinct.^[5] The Pyrenean ibex (C. p. pyrenaica) was characterized by a darker gray-brown coat in males, with prominent black markings on the legs, flanks, and neck, distinguishing it from the lighter pelage of other subspecies.^[6] It inhabited the Pyrenees and became extinct in January 2000, with the last known individual, a female named Celia, dying from injuries caused by a fallen tree.^[7] The Portuguese ibex (C. p. lusitanica), noted for its brown markings and short, wide horns averaging 51 cm in length, went extinct in 1892 due to overhunting and habitat loss in its range across Portugal, Galicia, Asturias, and northern Castile.^[7] In contrast, the Southeastern Spanish ibex (C. p. hispanica) features a lighter brown coat and relatively shorter horns, typically reaching 80-100 cm in males, and is found in southern Iberian ranges such as the Sierra Nevada, Ronda Mountains, and Beceite.^[8] The Gredos ibex (C. p. victoriae), the largest subspecies with a robust body size—males weighing up to 80 kg and standing 70-75 cm at the shoulder—exhibits a darker winter coat and longer, more massive horns, occupying central sierras like Gredos.^[9] Genetic studies using mitochondrial DNA, particularly the cytochrome b gene, have confirmed distinctions among the subspecies, revealing separate lineages for C. p. pyrenaica from the victoriae-hispanica group, with the former showing closer affinity to the Alpine ibex (Capra ibex) in some analyses.^[10] These mtDNA assessments highlight low overall genetic diversity in the species but support the historical taxonomic separation based on sequence polymorphisms.^[5] Efforts to revive the Pyrenean ibex through cloning utilized skin cells cryopreserved from Celia prior to her death. In July 2003, scientists produced the first cloned individual from an extinct subspecies via somatic cell nuclear transfer into domestic goat oocytes, resulting in a live female birth via cesarean section in a surrogate Spanish ibex–domestic goat hybrid. However, the clone survived only minutes before succumbing to severe respiratory failure caused by atelectasis in the left lung and an additional disconnected lung lobe.^[7]

Physical description

Morphology

The Iberian ibex (Capra pyrenaica) exhibits a robust build adapted to montane terrains, with adults typically measuring 100–155 cm in head-body length, standing 65–89 cm at the shoulder, and weighing 30–90 kg, whereby males attain larger dimensions than females.^[1]^[9] Its pelage comprises short, coarse outer hairs overlying a dense underwool layer that provides thermal insulation; coloration shifts seasonally from grayish-brown in summer to a thicker, darker winter coat with pale undertones on the sides.^[9] The species features short, sturdy legs supporting its frame, complemented by large, flexible black hooves that are cloven—divided into distinct front and rear sections—with spongy, rubber-like padding on the soles to enhance grip and traction on steep, rocky inclines.^[1]^[11] Iberian ibex possess keen visual acuity for spotting predators at distance, aiding survival in open, rugged landscapes.^[1]

Horns and sexual dimorphism

The Iberian ibex exhibits pronounced sexual dimorphism, particularly in horn development and overall body size. Males possess prominent horns characterized by a lyre-shaped curvature that extends outward, then upward and backward over the head. These horns begin growing at approximately 2-3 years of age and continue to develop throughout the male's life, with annual growth rings (annuli) forming that serve as reliable indicators of age.^[12] Maximum horn length in males reaches up to 83 cm, though typical lengths range from 36-78 cm, with basal circumference varying from 16-23 cm.^[13]^[14] In male-male combats during the rut, these horns are used to establish dominance, with larger, more symmetrical structures correlating with greater competitive success.^[14] Females possess shorter, thinner horns typically measuring 13.5–28.7 cm in length, contributing to the species' marked sexual dimorphism, though they exhibit subtle differences in body proportions such as relatively shorter limbs compared to males.^[15]^[9] Males are 30-50% heavier than females, with adult males weighing 50-90 kg and females 30-46 kg, reflecting differences in resource allocation for growth and maintenance.^[16] Males also possess larger skulls, with up to 95% of cranial measurements showing significant dimorphism, including greater overall dimensions and robusticity.^[15] This disparity extends to skeletal development, where females exhibit earlier bone ossification, completing appendicular skeleton maturation about two years faster than males, which supports their quicker attainment of reproductive maturity.^[17]

Distribution and habitat

Geographic range

The Iberian ibex (Capra pyrenaica) is native to the Iberian Peninsula, where its historical range encompassed mountainous regions across Spain, Portugal, Andorra, Gibraltar, and the Pyrenees straddling Spain and France.^[18] Key areas included the Sierra Nevada and Sierra de Gredos in central and southern Spain, the Serra da Estrela in northern Portugal, and the Pyrenean highlands.^[18] This distribution spanned elevations from sea level to over 3,400 m, reflecting the species' adaptability to diverse montane terrains.^[18] Currently, the ibex occupies fragmented populations primarily in Spain and Portugal, with major concentrations in the Sierra Nevada (approximately 15,000 individuals), Sierra de Gredos (about 8,000), and the broader Iberian System (over 50,000).^[18] In Portugal, numbers remain low, estimated at over 600 individuals in northern areas such as the Serra da Estrela, following reintroductions from Spanish stock.^[19] The species has been reintroduced to the French Pyrenees starting in 2014, with 254 individuals released by 2021, leading to a thriving population of approximately 900 by 2025, mainly in the Pyrenees National Park and Ariège Regional Natural Park.^[20]^[21] Overall, the global population exceeds 100,000 individuals as of the 2021 IUCN assessment, predominantly in Spain, with trends indicating stability or increase due to conservation efforts.^[18] Significant extirpations have reduced the range, including the complete loss from core areas in northern Portugal by the late 19th century and from the Pyrenees (where the subspecies C. p. pyrenaica went extinct) by 2000.^[18] The species is now considered extinct in Andorra and Gibraltar, though occasional vagrants have been noted in Andorra.^[19] These losses have confined remaining populations to isolated refugia, covering an extent of occurrence of about 90,000–200,000 km².^[18]

Habitat preferences

The Iberian ibex (Capra pyrenaica) primarily inhabits rugged mountainous environments at elevations typically exceeding 800 meters, with summer ranges extending up to 3,500 meters to access cooler alpine meadows and abundant herbaceous vegetation. During winter, individuals exhibit altitudinal migration, descending to lower elevations between 500 and 1,000 meters where snow cover is reduced and milder conditions prevail, often favoring south-facing slopes for enhanced solar exposure.^[1]^[22]^[23]^[24] In terms of terrain, the species strongly prefers steep slopes interspersed with cliffs, rocky outcrops, and escarpments, which provide critical refuge from predators and facilitate agile movement; flatlands and open lowlands are largely avoided due to increased vulnerability. These structural features are particularly vital in semi-arid regions, where even small cliffs contribute significantly to habitat suitability alongside sparse bush cover.^[1]^[23]^[25] Preferred vegetation encompasses a mix of coniferous forests (such as pines), deciduous woodlands (including holm oaks, Quercus ilex), open grasslands dominated by grasses like Festuca spp. and Deschampsia flexuosa, and shrublands offering browse. In southern populations, adaptability to Mediterranean scrublands is evident, where ligneous plants supplement the primarily graminoid diet. Microhabitat selection includes shaded, brushy areas and cliffs for resting and kid protection, with females particularly favoring elevated rocky sites during parturition for anti-predator security.^[1]^[26]^[23]^[27]^[28]

Behavior and ecology

The Iberian ibex displays pronounced sexual segregation in its social organization, with females and their juveniles forming stable, year-round matrilineal herds, while adult males remain solitary or associate in small bachelor groups outside the breeding season.^[1] These female-juvenile groups typically consist of 3 to 20 individuals, though larger aggregations of up to 50 can occur in winter when resources concentrate animals.^[29] Mixed-sex groups form briefly during the autumn rut, after which segregation resumes.^[30] Social hierarchies within these groups are structured around dominance interactions. In female herds, a linear hierarchy prevails, determined primarily by age and body weight, with higher-ranking individuals gaining priority access to resources through agonistic behaviors such as displacements and threats.^[31] Among males, dominance is established via horn clashes and displays, favoring larger, older individuals who lead bachelor groups or solitary wanderings.^[1] Iberian ibex are mainly diurnal, exhibiting bimodal activity peaks at dawn and dusk for foraging and movement, followed by midday resting on cliffs or shaded rocky areas to avoid heat and predation.^[32] Activity patterns shift seasonally; in winter, they are more consistently diurnal with midday peaks, whereas hot summers prompt increased nocturnal and crepuscular behavior to conserve energy.^[32] Individuals maintain nomadic movements within defined home ranges averaging 0.25 to 33 km², varying by sex, season, and location, with males occupying larger areas than females.^[33]^[30] Altitudinal migrations track forage availability, involving ascents to higher elevations in spring for fresh vegetation and descents in autumn to lower slopes, particularly in regions with steep topographic gradients.^[30]

Diet and foraging

The Iberian ibex (Capra pyrenaica) is an opportunistic mixed feeder, with diet composition varying by season and region. Herbaceous plants such as grasses and forbs often comprise 30–80% of intake in spring and summer, while shrubs and woody plants can make up 50–80% or more in autumn and winter, depending on availability. Key grass species include Festuca indigesta, Deschampsia flexuosa, and Agrostis nevadensis, selected for their abundance and moderate digestibility.^[34]^[35]^[36] Seasonal variations in diet reflect changes in vegetation availability and quality. In spring and summer, the ibex favors tender grasses and forbs at higher elevations, where herbaceous plants dominate up to 83% of the diet, driven by increased protein content and digestibility. During autumn and winter, consumption shifts toward woody browse such as Quercus ilex, Phillyrea latifolia, and shrubs like Cytisus spp. and Juniperus oxycedrus, comprising up to 59-78% of intake in harsher conditions. Lichens are occasionally consumed in winter but are not a primary component.^[37]^[36]^[23] Foraging behavior emphasizes selective intake of nutrient-rich plants within available patches, with the ibex prioritizing locally abundant species while secondarily favoring those with higher digestibility and lower fiber content. As ruminants, they engage in rumination during resting periods to efficiently process fiber-rich forage, supporting their adaptation to variable-quality vegetation. Feeding occurs primarily during dawn and dusk crepuscular periods, though this is modulated by habitat vegetation types.^[34]^[36]^[23] Physiologically, the Iberian ibex stores fat in the kidneys to buffer energy deficits during winter, when forage quality declines and snow cover limits access; these stores deplete over six months, reaching lowest levels in early spring, particularly under high population density or severe weather. This adaptation, combined with a ruminant digestive system optimized for low-quality plants, enables survival on sparse winter browse. The ibex derives most of its hydration from moisture in vegetation, reducing the need for frequent drinking in arid mountain habitats.^[38]^[34]

Reproduction and life cycle

The Iberian ibex (Capra pyrenaica) has a distinct seasonal reproductive cycle adapted to its mountainous environment. The rut, or mating season, occurs from November to December, lasting approximately 7 weeks. During this time, adult males leave their bachelor groups to join female herds, where they compete aggressively for mating opportunities through dominance hierarchies. Older males (over 5 years) typically engage in agonistic displays such as testing strength by pushing or clashing horns, while younger males resort to more primitive, direct fights; contests are often between age-similar individuals and favor the older combatant. Courtship involves vocalizations like roaring and physical displays including leaping to attract females, with older males achieving higher reproductive success by expending more energy on mating efforts in the later rut phase.^[39] Gestation lasts 5-6 months (22-24 weeks), resulting in births primarily from late April to early June, synchronized with peak vegetation availability. Females typically produce a single kid, though twinning occurs in fewer than 5% of cases. Newborns are precocial, weighing 2-3 kg at birth and capable of standing and moving within hours, which aids their survival in rugged terrain. Maternal care follows a "hider" strategy: for the first 1-2 weeks, the mother conceals the kid in rocky crevices or sheltered spots, visiting periodically to nurse while foraging alone to minimize detection by predators. After this isolation phase, kids rejoin female groups for protection, with lactation and weaning occurring around 3-4 months. Juvenile mortality is substantial, at 20-40%, largely from exposure, falls, and early predation during the vulnerable hiding period. Sexual maturity is attained earlier in females (at 1.5-2 years) than in males (2-3 years), though full reproductive potential peaks in prime-aged adults (4-13 years for females). In the wild, lifespan averages 10-15 years, limited by environmental stresses, predation, and disease, with females often outliving males due to less intense rut-related risks.

Predation

Predators

The primary natural predators of the Iberian ibex (Capra pyrenaica) target different life stages, with young kids being most vulnerable to raptors and smaller carnivores. The golden eagle (Aquila chrysaetos) preys on juvenile ungulates, including young ibex. In some Pyrenean populations, Bovidae (such as young feral goats) account for up to 14% of prey items and 32% of biomass in the golden eagle diet.^[40] Similarly, the red fox (Vulpes vulpes) attacks ibex kids, contributing to early-life mortality in rugged terrains.^[1] In northern ranges, such as the Cantabrian Mountains and Pyrenees, larger carnivores pose risks to juveniles and adults. The Iberian wolf (Canis lupus signatus) occasionally preys on ibex, representing about 3.1% of identified prey in scat analyses from central Spain, though wolves show a negative selection for this species compared to wild boar and roe deer.^[41] The Iberian lynx (Lynx pardinus), primarily a rabbit specialist, opportunistically hunts young ibex in ecosystems where alternative prey is scarce, as evidenced by taphonomic remains in lynx-modified sites.^[42] Brown bears (Ursus arctos) actively kill wild ungulates in the Cantabrian Mountains, including ibex, through ambush tactics on individuals in open areas, though such predation is opportunistic and supplements their mainly herbivorous diet.^[43] In the French Pyrenees, where ibex have been reintroduced since 2018, expanding bear and wolf populations may pose emerging predation risks to these herds, though specific impacts remain under study as of 2025.^[44] Human hunting has historically dominated as the most significant mortality factor for Iberian ibex across all life stages, driving population declines until regulated conservation measures took effect in the late 20th century.^[45] Predation impacts vary regionally due to differences in predator distributions. In southern Spain, where large carnivore densities are low following historical extirpations like that of the wolf, raptor attacks on kids represent a greater relative threat compared to northern areas.^[41] In the Pyrenees, reintroduced populations of brown bears and wolves have the potential to increase localized predation pressure on ibex herds, particularly juveniles, as these carnivores recolonize former ranges. While not predators, scavengers such as griffon vultures (Gyps fulvus) and cinereous vultures (Aegypius monachus) feed on ibex carrion, aiding in the removal of dead animals from habitats shared with recovering ibex populations in the Iberian Highlands.^[46]

Anti-predator behaviors

Iberian ibex employ a combination of vocal and behavioral signals to detect and respond to threats. Upon sensing danger, typically through visual or olfactory cues, individuals emit distinctive explosive whistling calls that alert nearby herd members and initiate a group response. These alarm calls are frequently observed in reaction to disturbances such as approaching humans and serve to coordinate collective evasion, enhancing survival in rugged environments.^[1]^[47] Escape tactics in Iberian ibex emphasize rapid flight to terrain where their agility provides a decisive advantage. When alarmed, herds form columns and flee toward steep rocky slopes, cliffs, or broken country, utilizing these features as refuges inaccessible to most predators. Experienced adults often lead these movements, guiding the group to familiar safe areas and minimizing disorientation during pursuit. This strategy leverages the species' morphological adaptations for climbing, allowing individuals to outmaneuver threats on precipitous ground.^[48]^[1] Vigilance levels vary contextually to balance anti-predator monitoring with other activities. In open habitats, ibex exhibit higher rates of scanning for threats, with individual vigilance decreasing as group size increases due to shared detection efforts. Lactating females demonstrate elevated vigilance, particularly when guarding young kids hidden in inaccessible rocky crevices during the initial weeks post-parturition to shield them from predators. During the breeding season, males display increased aggression toward potential threats, potentially deterring approaches while defending territories.^[49] Grouping confers significant anti-predator advantages for Iberian ibex, primarily through risk dilution and improved detection. Herds reduce the per capita probability of predation by spreading risk across members, allowing more time for foraging in larger assemblages. Juveniles position themselves centrally within groups for added protection, while solitary individuals—often adult males outside the rut—rely more heavily on personal speed, agility, and terrain familiarity for evasion. These dynamics underscore the adaptive value of sociality in mitigating predation pressure.^[1]^[49]

Conservation

Historical population declines

The Iberian ibex (Capra pyrenaica) underwent severe population declines during the 19th century, primarily driven by overhunting for meat and trophies, as well as habitat conversion to agricultural land, which fragmented its montane ranges across the Iberian Peninsula.^[50] These pressures led to the extinction of the Portuguese subspecies (C. p. lusitanica), once distributed in northern Portugal and Galicia, with the last individuals disappearing around 1890 due to relentless persecution and loss of suitable terrain.^[50] By the mid-19th century, the species had vanished from the French Pyrenees, marking a significant contraction of its overall range.^[23] In the 20th century, unregulated hunting continued to decimate remaining populations, particularly the Pyrenean subspecies (C. p. pyrenaica), which dwindled to fewer than 100 individuals by the early 1900s and just 10 by the late 1980s, culminating in the death of the last known wild female in January 2000.^[50] Disease outbreaks, such as sarcoptic mange (Sarcoptes scabiei), further exacerbated losses, causing over 95% mortality in affected groups like those in the Sierras de Cazorla during the 1990s, though earlier epizootics contributed to broader instability.^[50] Competition with domestic goats for forage and space intensified these declines, as livestock grazing degraded habitats and transmitted pathogens in shared areas.^[23] By the 1970s, the total Iberian ibex population had fallen to fewer than 10,000 individuals across fragmented nuclei, a stark reduction from its historically abundant status in the Middle Ages, underscoring the cumulative impact of anthropogenic factors up to that point.^[51] Specific regional censuses reflected this nadir, such as around 700 in the Maestrazgo mountains in 1970 and over 9,000 in the Gredos range by 1977, highlighting uneven but overall precarious distributions.^[50]

Current status and threats

The Iberian ibex (Capra pyrenaica) is classified as Least Concern on the IUCN Red List, based on a 2021 assessment that highlights its overall stable status across its range. However, vulnerabilities persist at the subspecies level, particularly for the southeastern Spanish ibex (C. p. hispanica), which faces localized pressures despite the species' recovery from historical lows. The total population is estimated at over 50,000 individuals and continues to increase, predominantly in Spain with a small reintroduced group of about 100 in Portugal. Populations are stable or increasing in core Spanish mountain ranges such as the Sierra Nevada and Gredos, but remain fragmented due to geographic barriers and past extirpations, limiting gene flow and resilience.^[19] Sarcoptic mange, caused by the mite Sarcoptes scabiei, remains a primary ongoing threat, with outbreaks causing significant morbidity and mortality in affected herds, though prevalence has declined in some areas due to developing resistance in ibex populations.^[52] Overgrazing by expanding ibex herds and competition with extensive livestock, particularly domestic goats, contributes to habitat degradation, displacing ibex to suboptimal elevations and reducing forage quality in central Spanish ranges.^[53] Hybridization with domestic goats poses a genetic risk, introducing introgressed alleles that threaten the purity and adaptability of wild populations, especially in fragmented habitats near human settlements.^[20] Climate change exacerbates these issues by altering precipitation patterns and vegetation growth, forcing altitudinal shifts that limit access to optimal forage and increase exposure to novel stressors.^[54] Human-related factors further compound risks, including regulated trophy hunting that, while managed through quotas, can impact male demographics if not balanced, alongside persistent poaching that undermines population stability.^[45] Vehicle collisions are increasingly documented in migration corridors and near expanding human infrastructure, contributing to non-natural mortality rates among dispersing individuals.^[55] These threats highlight the need for targeted monitoring to address fragmentation and disease dynamics in this recovering but unevenly distributed species.^[56]

Conservation efforts and reintroductions

The Iberian ibex benefits from protection in several key national parks across its range, including Sierra Nevada National Park in southern Spain, which hosts the largest and most genetically diverse population on the Iberian Peninsula and implements long-term monitoring to support conservation.^[28] Similarly, Ordesa y Monte Perdido National Park in the Spanish Pyrenees serves as a critical habitat, safeguarding ibex through strict preservation measures within its diverse alpine ecosystems.^[57] To balance population growth with habitat sustainability, regulated hunting quotas have been enforced since the 1960s, with national game reserves established under Law 37/1966, transforming hunting from a historical threat into a tool for population control and funding habitat management.^[45] Reintroduction programs have played a pivotal role in restoring the species, particularly in areas where subspecies were historically extinct. An early effort to revive the Pyrenean subspecies through cloning in 2003 ultimately failed due to health issues in the cloned individuals, highlighting challenges in genetic resurrection techniques. In contrast, translocations of ibex from Spanish populations to the French Pyrenees, initiated in 2014, have proven successful, with approximately 900 individuals established as of 2025 through ongoing releases that promote natural dispersal and breeding.^[21] These efforts emphasize sourcing from genetically robust stocks to enhance viability without compromising local adaptations. Active management strategies address overpopulation and health risks to ensure long-term stability. In densely populated areas like Sierra Nevada, selective culling is employed to mitigate overgrazing and reduce disease transmission risks, with targeted removals proving more efficient than broad-scale interventions.^[58] Comprehensive disease monitoring and vaccination protocols target pathogens shared with livestock, such as sarcoptic mange, through regular surveillance and health assessments in both wild and captive populations.^[59] Habitat restoration initiatives focus on rehabilitating grazing lands to minimize competition with domestic goats, including removal of invasive species and fencing to create ibex-preferred zones.^[60] International collaboration bolsters these efforts through EU-funded LIFE projects, which prioritize genetic purity by promoting translocations from diverse source populations and monitoring hybridization risks with domestic goats.^[61] Additionally, ecotourism initiatives in rewilding areas like the Iberian Highlands generate revenue for conservation by attracting visitors to observe ibex in their natural habitats, supporting habitat protection without direct exploitation.^[62]

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[21]
Home range in genus Capra: from polygons to Brownian bridges of ...
Distribution, status and conservation problems of the Spanish Ibex, Capra pyrenaica (Mammalia: Artiodactyla). Mammal Review 32(1):26–39. 10.1046/j.1365 ...<|control11|><|separator|>
[22]
Small cliffs are key predictors of the presence of Iberian wild goat
We model the habitat of Iberian wild goat Capra pyrenaica in a semi-arid area. •. Habitat is predominantly explained by small cliffs and bushes. •. High ...
[23]
Capra pyrenaica Schinz 1838 - Plazi TreatmentBank
May 2, 2022 · The forehead, beard, front and back of the neck, chest, brisket, and front of the shoulder and legs are black; there is a dark mid-dorsal stripe ...
[24]
Role of Various Woody Species in Spanish Mediterranean Forest ...
Aug 30, 2025 · This study analyses the importance of nine woody species in the diet of two wild herbivores with browser and browser-grazer feeding habits: ...
[25]
Long-term monitoring of the Iberian ibex population in the Sierra ...
Jun 25, 2020 · Distribution, status and conservation ... Biology, ecology and status of Iberian ibex Capra pyrenaica: A critical review and research prospectus.
[26]
Spatial structure in groups of Spanish ibex (Capra pyrenaica)
In the first study we characterized the pattern of agonistic interactions during feeding throughout a whole year in groups of Iberian ibex bucks; and in the ...
[27]
Linking seasonal home range size with habitat selection and ...
Jan 5, 2018 · The Iberian ibex (Capra pyrenaica) is an endemic species of the Iberian Peninsula that inhabits mountainous systems [19].Missing: elevation | Show results with:elevation
[28]
Social Dominance and Breeding Activity in Spanish Ibex (Capra ...
Dominance hierarchies were established in both groups by noting agonistic interactions between the individual females. Six high-ranking females of the 10 ...Missing: Iberian | Show results with:Iberian
[29]
Time distribution of activities in the Spanish ibex, Capra pyrenaica
Aspects of the behaviour and circadian rhythms (primarily activity patterns) of the Artiodactyla have been studied quite extensively. It has been ...
[30]
The home-range of the Spanish ibex in spring and fall - ResearchGate
Aug 10, 2025 · Seasonal home ranges varied from 39 km2 (fall/winter) to 1,272 km2 (breeding). Movements during 1987 and 1988 totaled 246 and 1,308 km2, ...
[31]
(PDF) The feeding strategy of Spanish ibex (Capra pyrenaica) in the ...
Aug 6, 2025 · The diet of the Spanish ibex is primarily herbaceous throughout the year and consists essentially of grasses. Diet selection was primarily ...Missing: sources | Show results with:sources
[32]
Summer feeding strategy of Spanish ibexCapra pyrenaica and ...
The ibex consumed cell wall-rich resources such as grasses and shrubs, while the sheep consumed higher quality feed and includes more forbs in its diet.Missing: sources | Show results with:sources
[33]
[PDF] Diet selection by Spanish ibex in early summer in Sierra Nevada
Previous analyses of the Spanish ibex Capra pyrenaica Schinz, 1838 feeding habits have revealed variations between locations, seasons and sexes (Martinez. 1988, ...
[34]
(PDF) Seasonal dietary shifts and selection of Iberian wild goat Capra pyrenaica Schinz, 1838
### Summary of Seasonal Dietary Shifts and Selection of Iberian Wild Goat (Capra pyrenaica)
[35]
[PDF] The effects of winter severity and population density on body stores ...
Abstract We studied the factors that determined kidney fat stores (KFs) and kidney stores (Ks)—defined as the residuals from the linear regression of kidney ...
[36]
https://rcin.org.pl/Content/13024/BI002_2613_Cz-40-2_Acta-T44-nr31-335-346_o.pdf
[37]
(PDF) Diet and Breeding Performance of the Golden Eagle Aquila ...
In the present paper, we report data on the diet and breeding performance of Golden Eagles Aquila chrysaetos monitored during a ten-year period in two semi- ...
[38]
[PDF] Alarm calls and flight behaviour in Spanish ibex (Capra pyrenaica)
Spanish ibex are cryptic in colour but in response to potential predators they give an auditory signal consisting of high-pitched explo- sive whistle, which ...
[39]
Wild Boar and Roe Deer as Key Species for Its Conservation - PMC
The predominant wild fauna species that are susceptible to be preyed upon by wolves were: wild boar, roe deer, and mountain goat (Capra pyrenaica). The ...
[40]
Taphonomic criteria for identifying Iberian lynx dens in quaternary ...
Apr 29, 2020 · pyrenaica), in which rabbits are scarce. Here, the cave-dwelling ibex could have been the prey of lynx. Lynx remains are also very abundant in ...
[41]
Behaviour of brown bears killing wild ungulates in the Cantabrian ...
Aug 9, 2025 · No serious incidents between the two species have so far been recorded, but the presence of wolves is clearly a potential threat to small cubs.
[42]
Hunting Iberian ibex: from a threat factor to a conservation tool
Feb 20, 2025 · In this study we have compiled the information from official databases about Iberian ibex, Capra pyrenaica, hunting yields, hunting trophies and hunting ...
[43]
Iberian Highlands - Rewilding Spain
This has encouraged the comeback of deer, wild boar, mouflon, small groups of Iberian ibex and an abundance of Egyptian and griffon vultures; but the top ...
[44]
Mammals (Chapter 3) - Escaping From Predators
Alarm calls and flight behaviour in Spanish ibex (Capra pyrenaica). Biology of Behaviour, 13, 11–21.Google Scholar. Alendal, E. & Byrkjedal, I. (1976) ...<|control11|><|separator|>
[45]
[PDF] The CIC Caprinae Phenotype System - Regulations.gov
Another exception is Capra pyrenaica victoriae, which occurred originally only in the ... in or near steep, rocky, or broken country and will utilize cliffs to ...
[46]
[PDF] Grouping behaviour of Iberian wild goat during the parturition period ...
Jan 4, 2021 · Group size and composition of the. Spanish ibex (Capra pyrenaica Schinz) in the Sierras of. Cazorla and Segura. Pp: 134-147. In: S. Lovari (ed) ...<|separator|>
[47]
[PDF] Distribution, status and conservation problems of the Spanish Ibex ...
In this paper, the distribution and status of the Spanish Ibex, Capra pyrenaica (Mammalia: Artiodactyla), are revised. The whole Iberian population numbers ...
[48]
wildlife comeback in europe the recovery of selected mammal and ...
... fewer than 10,000 breeding pairs (or 20,000 breeding individuals, or ... Iberian ibex Capra pyrenaica Summary The Iberian ibex declined due to over ...<|control11|><|separator|>
[49]
Long-term monitoring of the Iberian ibex population in the Sierra ...
Jun 25, 2020 · Biology, ecology and status of Iberian ibex Capra pyrenaica: A critical review and research prospectus. Mammal Rev 39, 17–32 (2009). Article ...
[50]
Development of resistance to sarcoptic mange in ibex - Pérez - 2022
Apr 7, 2022 · Our model predicts that the prevalence of sarcoptic mange in the Iberian ibex will continue to decrease over the next 2 years.
[51]
The Iberian ibex is under an expansion trend but displaced to ...
May 16, 2006 · The Iberian ibex is under an expansion trend but displaced to suboptimal habitats by the presence of extensive goat livestock in central Spain.
[52]
Climate change impacts on altitudinal movements of society large ...
Apr 13, 2025 · This study examines the impact of climate change on the altitudinal movement patterns and number of individuals of four large mammal species
[53]
(PDF) Wildlife-vehicle collisions in Spain - ResearchGate
Aug 7, 2025 · In this study, we depict the involved species and evaluate the geographic distribution and economic cost of this human-animal interaction in Spain.
[54]
Genetic signature of blind reintroductions of Iberian ibex (Capra ...
Sep 21, 2022 · Currently the species is extinct in the northernmost areas of its native range, namely in the Pyrenees [2]. Four Iberian ibex subspecies have ...
[55]
Fauna of the Ordesa and Monte Perdido National Park
The Ordesa and Monte Perdido National Park is a protected area located in the province of Huesca, in the autonomous community of Aragon, Spain.
[56]
https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0269873
[57]
Targeted strategies for the management of wildlife diseases
Sep 14, 2021 · Interestingly, the number of ibex euthanized and culled in targeted strategies were lower than in untargeted ones, thus decreasing the ...
[58]
Diseases of Iberian ibex (Capra pyrenaica)
Jun 1, 2023 · This review identifies the gaps in knowledge regarding pathogenesis, immune response, diagnostic methods, treatment, and management of diseases in Iberian ibex.
[59]
(PDF) The Iberian ibex is under an expansion trend but displaced to ...
Aug 6, 2025 · The Iberian ibex is under an expansion trend but displaced to suboptimal habitats by the presence of extensive goat livestock in central Spain.
[60]
LIFE 3.0 - LIFE93 NAT/E/011800
This would be supported by habitat improvement measures to increase the extent and quality of the ibex's grazing land.Missing: Iberian purity ecotourism
[61]
Rewilding Spain to enhance nature-based tourism in the Iberian ...
Feb 1, 2023 · Wild nature has the potential to provide valuable jobs and income for people in and around the Iberian Highlands rewilding landscape.