Fact-checked by Grok 2 weeks ago

Upper Paleolithic

The Upper Paleolithic represents the final subdivision of the Paleolithic period, spanning approximately 50,000 to 10,000 years ago (with regional variations) and characterized by the proliferation of Homo sapiens across Eurasia and eventually the Americas following their migration out of Africa, alongside the development of refined blade-based stone tools, organic material technologies, and the earliest widespread evidence of symbolic behavior including art and ornamentation. This era, often termed the "Upper Paleolithic Revolution," witnessed the replacement or assimilation of archaic human populations such as Neanderthals in Europe and western Asia, driven by modern humans' adaptive advantages in technology and social organization.^[1] Technologically, it featured innovations like prismatic blade production from prepared cores, burins for engraving and woodworking, and composite tools incorporating bone, antler, and ivory—such as needles for sewing clothing and harpoons for fishing—enabling more efficient exploitation of diverse environments from tundra to coastal zones.^[2] Culturally, the period is renowned for its artistic output, including monumental cave paintings and engravings depicting animals and abstract symbols at sites like Chauvet and Lascaux in France, as well as portable art forms such as carved ivory figurines and Venus statuettes, suggesting ritualistic or communicative functions tied to emerging social complexity.^[3] Subsistence relied on mobile hunter-gatherer strategies, with specialized big-game hunting using spears and atlatls, supplemented by gathering, small-game trapping, and early evidence of long-distance exchange networks for materials like shells and ochre, reflecting population growth and interconnected groups across continents.^[4] By its close around the onset of the Holocene approximately 11,700 years ago, these developments had set the stage for the Mesolithic transition, underscoring the Upper Paleolithic as a pivotal phase in human behavioral modernity.^[5]

Definition and Chronology

Definition and Key Characteristics

The Upper Paleolithic represents the final subdivision of the Paleolithic era, dating from approximately 45,000–50,000 to 10,000 years before present (BP), and is defined by significant technological, behavioral, and cultural advancements among anatomically modern humans.^[6] This period marks a shift toward more sophisticated stone tool production, the integration of organic materials in artifact manufacture, and the onset of symbolic expressions that indicate expanded cognitive and social capacities.^[7] Central to the Upper Paleolithic are key characteristics associated with the emergence of behavioral modernity, including the creation of complex toolkits for diverse tasks, the production of portable and parietal art such as engraved bones and cave murals, evidence of long-distance exchange networks for materials like ochre and shells, and the organization of settlements into semi-permanent sites with distinct functional zones for processing, manufacturing, and ritual activities.^[8] These traits reflect enhanced planning, innovation, and social complexity compared to earlier hominin adaptations.^[9] Unlike the Middle Paleolithic, which emphasized the Levallois prepared-core technique for producing predetermined flakes, the Upper Paleolithic introduced prismatic blade technology—yielding elongated, parallel-sided blanks from conical or cylindrical cores—and burins, specialized tools for incising and grooving.^[10] This technological transition was accompanied by a marked increase in the working of organic substances, with bone, antler, and ivory fashioned into awls, points, and ornaments, enabling finer craftsmanship and adaptation to varied environments.^[11] Notable innovations include eyed sewing needles, with the earliest evidence from Eurasia dating to around 45,000 BP, used for threading sinew or fiber to produce fitted garments from hides, which supported habitation in colder climates.^[12] From early phases onward, the development of atlatls—levered spear-throwers for greater propulsion—and bows with arrows (evidenced as early as ca. 54,000 BP) further revolutionized hunting strategies, allowing for more effective pursuit of large game from safer distances.^[13]

Temporal Boundaries and Phases

The Upper Paleolithic is conventionally dated from approximately 45,000–50,000 to 10,000 years before present (BP, uncalibrated radiocarbon years), spanning the final stages of the Pleistocene epoch and concluding with the transition to the Holocene around 10,000 BP.^[6] The onset is debated, with some scholars placing it at the Middle-Upper Paleolithic transition around 50,000 BP (Initial Upper Paleolithic) and others at around 40,000 BP with the full establishment of blade technologies in Europe. This period represents the last subdivision of the Paleolithic, characterized by the widespread adoption of blade-based technologies by anatomically modern humans, and it ends regionally with the onset of Mesolithic or Epipaleolithic cultures as post-glacial warming altered human adaptations.^[14] Scholars divide the Upper Paleolithic into four main phases based on technological and stratigraphic evidence from key Eurasian sites: the Initial phase (c. 50,000–40,000 BP), marking the transition from Middle Paleolithic traditions; the Early to Middle phase (c. 40,000–25,000 BP), featuring refined blade production and diversification; the Late phase (c. 25,000–15,000 BP), associated with adaptations during the Last Glacial Maximum; and the Terminal phase (c. 15,000–10,000 BP), involving post-glacial recolonization and microlithic innovations.^[15] These divisions, while centered on European and Near Eastern chronologies, provide a framework for correlating global assemblages, though exact boundaries vary due to regional differences in human dispersal and environmental pressures.^[16] The spatial extent of the Upper Paleolithic began primarily in Eurasia and Africa, where early manifestations appear around 50,000 BP, before expanding to other continents.^[6] Human occupation in Australia is dated to between 65,000 and 43,000 BP, though recent studies as of 2025 suggest a later arrival around 50,000 BP or earlier, reflecting maritime dispersal from Southeast Asia.^[17]^[18] In the Americas, the period commenced later, around 20,000–15,000 BP, as populations migrated via Beringia during a period of lowered sea levels.^[19] Regional variations in temporal boundaries are notable, with African Upper Paleolithic industries persisting until approximately 12,000 BP in some areas, blending into Later Stone Age traditions amid differing ecological contexts.^[20] Calibration of dates poses challenges, particularly for assemblages older than 40,000 BP, where radiocarbon reliability diminishes due to low atmospheric carbon-14 levels; in such cases, uranium-thorium dating of associated carbonates or optically stimulated luminescence of sediments provides critical chronological control.^[21]

Paleoenvironmental Context

Climate Fluctuations

The Upper Paleolithic period, spanning approximately 50,000 to 12,000 years before present (BP), occurred within the Last Glacial Period, encompassing Marine Isotope Stages 3 (MIS 3, ~60,000–27,000 BP) and 2 (MIS 2, ~27,000–11,700 BP). During MIS 3, the climate was characterized by significant oscillations, including Dansgaard-Oeschger (D-O) cycles and Heinrich events, which drove rapid alternations between cold stadials and warmer interstadials across the Northern Hemisphere.^[22] D-O cycles involved abrupt warmings of 10–15°C over Greenland in as little as decades, followed by gradual coolings, as evidenced by high-resolution ice core records.^[23] Heinrich events, marked by massive iceberg discharges into the North Atlantic, punctuated these cycles with intense cold phases lasting centuries to millennia, disrupting ocean circulation and amplifying regional cooling.^[24] A notable warm phase within MIS 3 was Greenland Interstadial 8 (GI-8), centered around 38,000 BP, which represented a brief amelioration in the otherwise variable climate, allowing temporary expansions of more temperate conditions in parts of Eurasia.^[25] These interstadials contrasted sharply with the stadials, where temperatures dropped rapidly, fostering aridity and permafrost expansion. Transitioning into MIS 2, climate conditions intensified toward the Last Glacial Maximum (LGM, ~26,000–19,000 BP), a period of peak cold and aridity driven by expanded ice sheets and weakened Atlantic Meridional Overturning Circulation.^[26] Ice core data from the Greenland Ice Sheet Project 2 (GISP2) confirm these swings, with MIS 2 exhibiting sustained low temperatures and amplified D-O-like variability, including shifts of 10–15°C over short timescales.^[23] These fluctuations profoundly shaped paleoenvironments, leading to the widespread expansion of tundra-steppe biomes across Eurasia during stadials and the LGM, as cold, dry conditions supplanted forests and promoted grass-dominated landscapes suited to grazing megafauna.^[27] Global sea levels dropped by up to 120 meters during the LGM due to ice volume buildup, exposing continental shelves and land bridges such as Beringia, which connected Siberia and North America and facilitated biotic exchanges.^[26] In MIS 2, intensified cold further entrenched these arid, open biomes, with pollen and macrofossil records indicating tundra dominance north of 57°N in Eurasia, interspersed with steppe-like vegetation to the south and east of major ice sheets. Such environmental instability underscored the dynamic climatic backdrop against which human populations navigated the Upper Paleolithic.

Geographical and Ecological Changes

During the Upper Paleolithic, extensive glaciations in the northern hemisphere profoundly altered landscapes, with ice sheets covering large portions of northern Europe, Scandinavia, and North America, while periglacial zones—characterized by permafrost, frost action, and tundra-like conditions—formed adjacent to the glaciers.^[28] These changes were particularly pronounced in Eurasia, where the advance of the Fennoscandian Ice Sheet during the Last Glacial Maximum (LGM, approximately 26,000–19,000 years ago) displaced ecosystems southward and created vast open terrains.^[29] The resulting mammoth steppe, a cold, arid grassland biome extending from western Europe across Siberia to Beringia, emerged as one of the largest continuous ecosystems of the Pleistocene, supported by low precipitation and frequent clear skies that limited forest growth and promoted grass-dominated vegetation.^[30] This steppe environment, peaking in extent during glacial episodes, facilitated the movement of large herbivore populations but also imposed harsh conditions on surrounding habitats. Lowered global sea levels, reaching up to 130 meters below present during peak glaciation, exposed extensive continental shelves and created land bridges that connected previously isolated regions, reshaping coastal geographies and terrestrial connectivity.^[31] In the North Sea, Doggerland formed a low-lying plain linking Britain to the European mainland, encompassing rivers, wetlands, and uplands that supported diverse flora and fauna until its gradual inundation began around 10,000 years ago.^[32] Similarly, in Southeast Asia, Sundaland united the Malay Peninsula with Borneo, Sumatra, and Java into a single landmass, while Sahul connected Australia, Tasmania, and New Guinea, enabling biotic exchanges across these now-submerged areas during periods of lowered sea levels from about 50,000 to 10,000 years ago.^[33] The Bering Land Bridge, spanning the Bering Strait between Siberia and Alaska, was exposed relatively late, emerging around 35,700 years ago due to regionally variable sea-level dynamics and remaining viable until approximately 11,000 years ago, when post-glacial flooding submerged it. Ecologically, these geographical transformations drove the expansion of open grasslands across Eurasia and North America, creating ideal foraging grounds for herd animals such as woolly mammoths, reindeer, horses, and bison, which thrived in the nutrient-rich, low-biomass steppe-tundra mosaic.^[34] Grassland proliferation, linked to cooler and drier conditions enabled by glacial cycles, reduced woodland cover and favored megafaunal assemblages that defined the mammoth steppe's high productivity.^[30] During the LGM, southern refugia—such as the Iberian and Italian peninsulas in Europe, along with coastal and highland areas in Africa—served as biodiversity hotspots, harboring temperate species and plant communities amid the contraction of northern habitats.^[28]^[35] Recent modeling from the 2020s, incorporating high-resolution sea-level reconstructions and paleotopographic data, highlights the dynamic nature of these coastlines, revealing fluctuating shorelines that repeatedly opened and closed migration corridors along Pacific and Indian Ocean margins during the Upper Paleolithic.

Human Dispersal and Adaptation

Anatomically Modern Humans

The Upper Paleolithic marks the period during which anatomically modern humans (Homo sapiens) became the dominant hominin species in Eurasia, exhibiting fully modern skeletal morphology characterized by a high-vaulted cranium, reduced facial robusticity, and a globular braincase, as evidenced by fossils such as the Peştera cu Oase 2 cranium from Romania, which displays a suite of derived modern human features contrasting with Neanderthal traits.^[36] These individuals also showed regional variations adapted to local environments, including more robust appendicular skeletons and greater overall body mass in cold-adapted populations of northern and central Europe during the Gravettian phase, reflecting higher mechanical loading from mobility and hunting in periglacial conditions, with humeral diaphyseal robusticity significantly elevated compared to later Mesolithic groups.^[37]^[38] Such adaptations are quantified by genetic models predicting taller statures and increased limb proportions in Early Upper Paleolithic Europeans, decreasing post-Last Glacial Maximum due to dietary and climatic shifts.^[38] Key fossil evidence includes the Oase 1 mandible from Peştera cu Oase, Romania, dated to approximately 40,000 years before present (BP), representing one of the earliest unequivocally modern human remains in Europe with fully modern dental and mandibular morphology.^[39] This specimen underscores the rapid dispersal of H. sapiens into Eurasia, where skeletal remains from sites like Kent's Cavern, England, further confirm the presence of fully modern anatomy by around 41,000–44,000 BP.^[40] Genetic studies reveal that Upper Paleolithic humans carried mitochondrial DNA (mtDNA) lineages derived from macrohaplogroups M and N, which originated from the African L3 haplogroup and dispersed out of Africa around 60,000–70,000 years ago, founding non-African maternal lineages as the only successful mtDNA variants to colonize Eurasia.^[41] Y-chromosome data indicate male-biased migration patterns in some early dispersals, with contrasting variation between Y-chromosome and mtDNA suggesting higher male mobility and gene flow in African source populations contributing to Eurasian expansions.^[42] Non-African populations retain 1–4% Neanderthal DNA from admixture events primarily dated to 50,000–40,000 BP, with genomic analyses of Upper Paleolithic individuals like those from Ranis, Germany, constraining the main introgression to after 45,000 BP and showing elevated Neanderthal ancestry in early European arrivals.^[40]^[43] Recent genomic research (2023–2024) supports multiple Neanderthal admixture events across continents, with the main introgression around 45,000 BP influenced by climate-driven population movements, and evidence from Eurasian ancient DNA indicating recurrent gene flow rather than a single pulse, shaping modern human genetic diversity through adaptive introgressions.^[44]^[43]^[45]

Migration Patterns

The final major wave of anatomically modern human dispersal out of Africa occurred between approximately 70,000 and 50,000 years before present (BP), primarily via a southern coastal route along the Indian Ocean, enabling rapid expansion into Eurasia.^[46] This migration is supported by archaeological and genetic evidence indicating a single primary exit point from East Africa, with populations adapting to diverse coastal and inland environments as they moved eastward.^[47] By around 60,000 BP, these groups had established presence in South Asia, setting the stage for further dispersals across the continent.^[48] In Eurasia, the spread accelerated during the early Upper Paleolithic, with modern humans colonizing Europe around 45,000 BP, as evidenced by Aurignacian sites in western and central regions.^[49] To the east, populations reached Siberia by approximately 40,000 BP, marked by Initial Upper Paleolithic assemblages in the Altai Mountains that reflect technological adaptations to cold steppe environments.^[50] Concurrently, coastal migration routes facilitated the peopling of Sahul (the combined landmass of Australia and New Guinea) around 65,000–50,000 BP, with archaeological sites like Madjedbebe dated to ~65,000 BP demonstrating early seafaring capabilities, though recent genetic evidence as of 2025 suggests a later initial arrival ~50,000 BP.^[46] Evidence from datings of cave art in Sulawesi, Indonesia, including 2021 findings at least 45,500 years old and 2024 evidence of narrative art at least 51,200 years old, further underscores an established human presence in Southeast Asia by this period, including symbolic behaviors indicative of cultural complexity.^[51]^[52] The entry into the Americas represents a later phase of Upper Paleolithic expansion, primarily via the Bering Land Bridge (Beringia) between approximately 23,000 and 15,000 BP, when lowered sea levels exposed a terrestrial corridor connecting Siberia and Alaska.^[53] This inland route was complemented by a Pacific coastal migration pathway, with recent evidence from the Cooper's Ferry site in Idaho, USA, yielding artifacts dated to around 16,000 BP that support pre-Clovis human occupation south of the ice sheets.^[54] Oceanographic and archaeological data from 2025 analyses reinforce this coastal model, highlighting how kelp forest ecosystems along the North Pacific Rim could have sustained early maritime voyagers.^[53] During the Last Glacial Maximum (LGM, c. 26,500–19,000 BP), these dispersing populations experienced genetic bottlenecks, likely due to climatic extremes and habitat contraction, which reduced effective population sizes and influenced regional genetic diversity.^[55] Global human population estimates for the Upper Paleolithic suggest growth to between 1 and 3 million individuals by around 20,000 BP, driven by successful adaptations and expansions despite periodic environmental stresses.^[56] These demographic increases reflect the cumulative impact of migrations, with densities varying from sparse hunter-gatherer bands in newly colonized regions to more stable groups in resource-rich areas of Eurasia and Africa.^[57]

Technological Advancements

Stone and Organic Tool Technologies

The Upper Paleolithic marked a significant advancement in lithic technologies, characterized by the systematic production of prismatic blades and specialized tools like burins, which facilitated more precise and versatile applications compared to earlier periods. Prismatic blades, struck from prepared cores to yield long, parallel-sided flakes with multiple dorsal ridges, became a hallmark of Upper Paleolithic toolmaking, enabling efficient material use and standardization across assemblages.^[10] Burins, typically manufactured on prismatic blades through deliberate spalling to create chisel-like edges, were primarily used for engraving and working soft materials such as bone or wood, reflecting increased specialization in tool forms.^[58] Innovations in manufacturing techniques further enhanced lithic efficiency. Pressure flaking, involving the application of controlled force with a soft tool like antler or bone to detach small flakes and refine edges, was widely adopted during the Upper Paleolithic, allowing for sharper and more durable tools.^[59] Heat treatment of stones, achieved by controlled heating to alter material properties and reduce fracturing during knapping, improved flake predictability and tool quality, as evidenced in various European and Asian sites.^[59] The Bohunician industry, dated to approximately 48,000–40,000 BP in Central Europe, represents a transitional phase bridging Middle and Upper Paleolithic technologies, featuring elongated Levallois points and early blade production that foreshadowed later developments.^[60] In late Upper Paleolithic phases, particularly in Siberian traditions around 18,000 BP and later, microblade technology emerged, involving the production of small, wedge-shaped blades detached via pressure techniques from microblade cores, which supported composite tools adapted to cold environments.^[61] Global variations included backed blades in African contexts, where one edge was blunted through steep retouch to create robust, multifunctional implements, appearing in early Upper Paleolithic-equivalent assemblages.^[62] Parallel to lithic advancements, organic tool technologies diversified, incorporating bone, antler, and ivory into durable implements. Bone points and awls, shaped through grinding and incision, served for piercing and carving, while antler harpoons and ivory carvings demonstrated skilled working of hard organics for practical and symbolic purposes.^[63] Recent evidence from Dzudzuana Cave in Georgia reveals plant fiber processing around 34,000 BP, with microscopic residues of indigo plant (Isatis tinctoria) on grinding tools indicating intentional manipulation of non-nutritional fibers, possibly for cordage or dyeing.^[64] By approximately 30,000 BP, Upper Paleolithic toolkits had diversified extensively, encompassing over 100 distinct types across lithic and organic categories, reflecting adaptations to varied subsistence needs and environmental pressures.^[59] This proliferation underscores a broader technological repertoire that integrated stone and organic materials for enhanced functionality.

Projectile and Hunting Innovations

The Upper Paleolithic period marked significant advancements in projectile weaponry, enabling anatomically modern humans to pursue large game more effectively from distances that minimized risk. Spear-throwers, known as atlatls, appeared in Europe around 25,000 years before present (BP), with early examples from the Solutrean culture in sites like Laugerie Haute in France, where antler implements facilitated greater leverage and velocity compared to handheld spears.^[65] These devices extended throwing range and force, with experimental reconstructions demonstrating dart velocities up to 100 km/h, sufficient to penetrate thick hides of megafauna.^[66] Bow-and-arrow technology likely emerged by approximately 20,000 BP in Eurasia, though direct evidence from Grotte Mandrin in southern France pushes its origins to around 54,000 BP, associated with the earliest modern human occupations in Europe.^[67] Small lithic points, including micropoints under 10 mm wide, exhibit impact fractures consistent with arrow use, suggesting poisoned tips for enhanced lethality against herd animals. At Franchthi Cave in Greece, assemblages from circa 13,000 BP include microlithic inserts indicative of composite arrows, reflecting refined archery during the late Upper Paleolithic transition to the Mesolithic.^[67] Composite projectile points with barbs further innovated hunting tactics, combining osseous shafts with multiple stone insets to cause greater tissue damage and hinder prey escape. A notable example from Les Prés de Laure in France, dated to 23,500 BP, consists of a bone point armed with 11 backed lithic barbs made from chert, used in horse hunting as evidenced by associated residues and wear patterns.^[68] These designs increased wounding efficiency, allowing smaller groups to target dangerous predators like cave lions.^[69] Hunting focused predominantly on large herbivores such as mammoths and reindeer, which dominated faunal assemblages across European sites, reflecting specialized strategies adapted to glacial steppe environments. At Kraków Spadzista in Poland, Gravettian layers (circa 25,000 BP) yield cut-marked mammoth bones, providing direct evidence of organized communal hunts that yielded substantial caloric returns.^[70] Reindeer exploitation, evident in high-density bone beds at sites like Předmostí in the Czech Republic, shows seasonal migrations tracked by humans, with sex ratios in assemblages suggesting selective culling of prime adults to maximize efficiency.^[71] These innovations boosted predatory success, as seen in faunal profiles from kill sites where single-species dominance—often exceeding 80% of remains—indicates higher kill rates and reduced energy expenditure per kilogram of meat procured. Seasonal hunting camps, such as Vale Boi in Portugal (circa 20,000 BP), feature dense red deer assemblages alongside processing tools, pointing to temporary aggregations for exploiting predictable migrations during optimal seasons like autumn. Overall, such advancements supported population growth among modern humans dispersing after 40,000 BP, contrasting with contemporaneous Neanderthal practices while overlapping in targeted megafauna.^[72]

Subsistence and Daily Life

Hunting and Foraging Strategies

During the Upper Paleolithic, hunting strategies emphasized the procurement of large game, particularly megafauna such as mammoths, reindeer, and bison, through communal drives and the use of traps to facilitate mass kills. These methods allowed groups to target herds efficiently, as evidenced by faunal assemblages showing age profiles indicative of both selective culling of prime adults and opportunistic captures of juveniles during seasonal migrations.^[73]^[74] Focus on megafauna persisted until the Last Glacial Maximum (approximately 26,000–19,000 years ago), after which extinctions and climatic shifts prompted diversification toward smaller herbivores.^[75] Foraging complemented hunting with seasonal collection of plant resources, including berries, roots, and tubers, which were gathered in open landscapes and forested edges to supplement protein-rich diets. Evidence from sites like Dolní Věstonice in Moravia reveals processing of starchy parenchyma tissues from wild plants, indicating targeted exploitation during peak availability periods.^[76] A notable 2025 discovery at Dzudzuana Cave in Georgia uncovered 34,000-year-old grinding tools bearing residues of Isatis tinctoria (woad), processed for indigo dye rather than food, suggesting early non-nutritional plant use for coloration or medicinal purposes.^[77] Isotopic analyses of human remains indicate that diets in Europe derived 60–80% of protein from animal sources, reflecting heavy reliance on terrestrial and freshwater prey, though broadening occurred mid-period with increased aquatic intake.^[78] In contrast, Early Upper Paleolithic sites on the Cantabrian Coast in northern Spain show evidence of shellfish consumption year-round, as seen in coastal middens.^[79] Specific hunting evidence includes the Mezhirich site in Ukraine, where four mammoth-bone huts constructed around 15,000 years ago incorporated remains from at least 100 individuals, implying large-scale procurement and processing.^[80] Tool marks on endscrapers from early Upper Paleolithic layers, such as those at Fumane Cave, further attest to systematic hide processing for clothing and shelter.^[81]

Settlement and Mobility Patterns

During the Upper Paleolithic, human groups utilized a diverse array of settlement types adapted to varying environmental conditions across Eurasia and beyond. Open-air camps were common in lowland and riverine settings, often featuring semi-subterranean pits lined with mammoth bones or hides for insulation, as evidenced by structures at sites like Mezhirich in Ukraine dating to around 15,000 BP. Rock shelters and cave entrances served as protected bases, particularly in rugged terrains, providing natural windbreaks and facilitating repeated occupations, such as at the Abri Pataud in France. These site types reflect strategic selections for resource proximity, with open-air locales dominating in open landscapes and shelters in more variable topographies. Mobility patterns in the Upper Paleolithic shifted over time, influenced by resource distribution and climatic fluctuations. Early phases, such as the Aurignacian (ca. 43,000–35,000 BP), were characterized by high residential mobility, where groups frequently relocated entire camps to track migratory herds, aligning with Binford's forager model of logistically simple, wide-ranging movements. By the terminal Upper Paleolithic, around 15,000–11,000 BP, reduced residential mobility emerged in some regions, marked by the appearance of storage pits for dried meats and plants, indicating more collector-like strategies with logistical forays from semi-permanent bases to buffer seasonal scarcities. These adaptations were driven by environmental pressures, such as glacial retreats that stabilized local resources.^[82]^[83] Archaeological evidence from hearth arrangements and artifact distributions suggests social organization centered on small family or kin-based units within larger aggregations. At many sites, hearths were clustered in discrete activity areas, with surrounding debris patterns implying nuclear family divisions of labor, such as tool maintenance near sleeping zones, as reconstructed from spatial analyses at open-air locales in southwest France. Trade networks further indicate interconnected social structures, with flint and obsidian sourced and exchanged over distances exceeding 500 km, as seen in Gravettian assemblages where non-local cherts from the Carpathians appear in Danube Valley sites, fostering alliances and information sharing.^[84]^[85]^[86] A prominent example is the Dolní Věstonice complex in Czechia, dated to approximately 26,000 BP, which represents one of the largest known seasonal settlements of the Gravettian period, featuring organized clusters of hearths and pit houses housing perhaps 100–150 individuals during winter aggregations for mammoth hunting and social rituals, with evidence of specialized activity zones like kilns for firing figurines. Recent GIS-based modeling has reconstructed mobility routes around such sites, using least-cost path analyses to map optimal paths between flint outcrops and settlements, revealing networks that integrated resource procurement with seasonal migrations across Central Europe.^[87]^[88]

Symbolic Culture and Cognition

Artistic Expressions

The Upper Paleolithic period marks a profound development in human artistic expression, characterized by the emergence of both fixed and portable forms of visual art that reflect advanced cognitive abilities, such as symbolic thinking and narrative representation. These artworks, primarily from Europe and extending to Southeast Asia, include depictions of animals, human figures, and abstract motifs, often created using natural pigments and engraving techniques. This artistic florescence, beginning around 45,000 years ago, underscores a shift toward complex cultural behaviors among anatomically modern humans.^[3] Cave art represents one of the most iconic manifestations of Upper Paleolithic creativity, featuring paintings, engravings, and reliefs predominantly in deep caves across western Europe. Over 360 known rock art sites in this region, concentrated in France, Spain, and southern Germany, contain thousands of images dating from approximately 40,000 to 12,000 years ago.^[89] Iconic examples include the Chauvet Cave in France, where engravings and paintings of lions, rhinoceroses, mammoths, and horses, executed around 36,000–30,000 years before present (BP), demonstrate sophisticated shading and movement.^[90] Hand stencils, created by blowing pigment around the hand pressed against the rock surface, appear frequently, symbolizing presence or identity; notable instances are found in Chauvet and other Aurignacian sites.^[90] These animal representations often dominate, comprising over 80% of motifs in many caves, suggesting themes related to hunting, mythology, or environmental observation.^[3] Portable art, movable objects crafted from stone, bone, ivory, and shells, further illustrates the period's artistic versatility and widespread distribution. Venus figurines, small carved female forms emphasizing exaggerated breasts, hips, and abdomens, exemplify this category; the Venus of Willendorf, a limestone statuette about 11 cm tall from Austria, dates to approximately 25,000 BP and is among the most renowned.^[91] Over 200 such figurines have been identified across Eurasia, often interpreted as symbols of fertility or idealized femininity.^[92] Decorated tools and personal ornaments, including engraved ivory spears, bone needles with incised patterns, and beads made from pierced shells or mammoth ivory, were produced in sites like the Swabian Jura in Germany, with examples from 40,000 BP onward.^[93] These items, found in habitation contexts, indicate personal adornment and possibly social signaling.^[94] Artistic techniques relied on locally sourced materials, highlighting ingenuity in pigment preparation and application. Primary pigments included red and yellow ochre (iron oxides), black charcoal or manganese oxide, and white kaolin clay, ground into powders and mixed with binders like water, fat, or urine for painting.^[95] Engravings were made using flint tools or fingers on soft surfaces, while blowing techniques produced stencils. For example, a cave painting in Sulawesi, Indonesia, depicting a narrative hunting scene with therianthropes and pigs, dated to at least 51,200 years ago via uranium-thorium dating, represents the oldest known figurative art to date, extending these practices beyond Europe and confirming early pigment use in Southeast Asia.^[52] Recent analyses, including 2023 studies on ochre processing at sites like Blombos Cave (though Middle Stone Age, informing Upper Paleolithic continuity), reveal evidence of grinding and heating for body painting and adornment, suggesting multifunctional pigment applications in daily and ceremonial life.^[96]

Burials and Ritual Practices

The Upper Paleolithic period is marked by the earliest clear evidence of intentional human burials, suggesting the emergence of complex belief systems, social structures, and ritual behaviors among anatomically modern humans. These practices often involved the deposition of grave goods, use of pigments, and careful body positioning, indicating concerns with the afterlife or social status. Sites across Eurasia provide key examples, with burials dating from approximately 40,000 to 10,000 years before present (BP).^[97]^[98] One of the most elaborate burials comes from Sungir in central Russia, dated to around 34,000 BP during the mid-Upper Paleolithic. Here, an adult male and two children were interred in separate graves with thousands of ivory beads—over 13,000 in total—strung as pendants and sewn onto clothing, alongside fox teeth pendants, bone points, and red ochre. The presence of such labor-intensive grave goods implies significant social investment and possible status differentiation, with the children's burials particularly rich, suggesting inherited or ritual significance. Red ochre, a mineral pigment, was liberally applied to the bodies and grave floors at Sungir and other sites, likely symbolizing blood, transformation, or protection in funerary rites; its processing and use reflect advanced symbolic cognition.^[99]^[100]^[101] Rare instances of post-mortem body processing hint at ritual cannibalism or defleshing. At El Mirón Cave in northern Spain, dated to about 18,000 BP in the Magdalenian period, a female individual's remains show cut marks, breakage, and exposure consistent with filleting and marrow extraction, interpreted as culturally motivated cannibalism rather than nutritional stress. Such practices may have served ceremonial purposes, like communal feasting or ancestor veneration, underscoring the ritual treatment of the dead.^[102] Other evidence points to broader ritual activities beyond burials, including possible shamanistic practices inferred from animal remains in caves. In Chauvet Cave, France (dated 36,000–30,000 BP), a cave bear skull was deliberately placed on a large stone slab in a deep chamber, alongside bear bones and engravings, suggesting ceremonial deposition tied to spiritual mediation or bear cults—bears often symbolized power in later ethnographic analogies. Additionally, early musical instruments like the ivory and bone flutes from Geissenklösterle Cave in Germany, dated to around 40,000 BP in the Aurignacian, indicate ritual sound-making, possibly for ceremonies invoking altered states or communal rites.^[103]^[104] Recent analyses of Upper Paleolithic burials reveal interpretations of gender roles through differential grave goods and positioning. For instance, at Sungir, the adult male's grave included projectile points suggestive of hunting status, while the children's adornments emphasized ornamentation, potentially reflecting emerging social norms around gender and age in ritual contexts. These patterns highlight increasing social complexity, though interpretations remain debated due to limited skeletal preservation.^[105]^[94]

Regional Cultural Traditions

European Developments

The Upper Paleolithic in Europe, spanning approximately 43,000 to 12,000 years before present (BP), represents one of the most intensively studied periods of human prehistory, characterized by a succession of distinct cultural techno-complexes that reflect adaptations to diverse environmental conditions, including the harsh climates of the Last Glacial Maximum (LGM). These sequences, primarily associated with anatomically modern humans who dispersed into the continent around 45,000 BP, showcase progressive technological refinements and cultural elaborations, with evidence drawn from thousands of sites across Western, Central, and Eastern Europe. The Aurignacian, marking the onset of the full Upper Paleolithic, emerged around 43,000–35,000 BP and is renowned for its innovative blade technologies, including prismatic blades struck from prepared cores, which enabled more efficient tool production and hafting for composite implements.^[106]^[107] Following the Aurignacian, the Gravettian period (33,000–22,000 BP) dominated much of Europe, featuring backed bladelets and shouldered points suited for hunting large game, alongside iconic symbolic artifacts such as the Venus figurines—small carved female forms interpreted as representations of fertility or ideal body types, found at sites like Dolní Věstonice in Moravia.^[108]^[109] The subsequent Solutrean (22,000–17,000 BP), concentrated in southwestern Europe, introduced exquisite bifacial flintworking techniques, exemplified by the thin, symmetrical laurel leaf points used as spear tips or knives, reflecting a high degree of lithic expertise amid retreating ice sheets.^[106]^[110] The era culminated in the Magdalenian (17,000–12,000 BP), which emphasized osseous technologies like antler harpoons and bone needles for fishing and sewing, alongside expansive seasonal settlements that supported post-LGM recolonization.^[106]^[111] A hallmark innovation of the European Upper Paleolithic was the explosion of symbolic expression through cave art, particularly during the Gravettian and Magdalenian phases, with parietal paintings depicting megafauna in vivid detail at sites such as Altamira in northern Spain (dated to around 36,000–12,000 BP) and Lascaux in southwestern France (approximately 17,000 BP). These artworks, executed with pigments like ochre and charcoal, suggest complex cognitive capacities for abstraction and possibly ritualistic purposes, coinciding with broader evidence of portable art and ornamentation.^[112]^[3] During the LGM (circa 26,000–19,000 BP), European hunter-gatherers adapted to extreme cold by centering their economies on reindeer (Rangifer tarandus), whose migrations dictated seasonal mobility and provided meat, hides for clothing, and bone for tools, as evidenced by dense faunal assemblages at open-air sites in France and Germany.^[113]^[111] Recent research, including a 2023 reassessment of Central European sites, has clarified the Aurignacian's divergence from the earlier Bohunician industry—previously linked to Neanderthals or transitional populations—demonstrating through refined chronology and lithic analysis that the Aurignacian arose independently among modern humans around 43,000 BP, without direct continuity from the Bohunician.^[114] Amid the LGM's glacial advance, human populations contracted into southern refugia, with genomic and archaeological evidence indicating persistent occupations in Iberia and the Italian peninsula, where milder climates and coastal resources sustained groups like the Solutrean in Iberia and proto-Gravettian variants in Italy until post-glacial expansion.^[55]^[115]

African and Near Eastern Variations

In Africa, the Upper Paleolithic period, often termed the Later Stone Age (LSA) in sub-Saharan contexts, is characterized by transitional industries emerging after 50,000 BP, with evidence of specialized toolkits for resource exploitation, such as post-Howiesons Poort assemblages around 44,000–42,000 BP at sites like Border Cave, incorporating blade-based technologies and small tools indicative of evolving microlithic trends.^[116] North African variations, particularly the Iberomaurusian industry starting around 25,000 BP but with roots in earlier bladelet production, emphasize backed microlithic bladelets for hafted tools, reflecting adaptations to coastal and arid zones in the Maghreb.^[117] In the Near East, the Levantine Upper Paleolithic begins with the Emiran culture around 47,000–42,000 BP, marked by Emireh points—thin, pointed blades with retouched bases used as spear tips or knives—signaling a shift from Levallois techniques of the Middle Paleolithic.^[118] This phase shows continuity from Middle Paleolithic traditions, as evidenced by early modern human burials at Skhul and Qafzeh caves dated to approximately 120,000–90,000 BP, where flexed skeletons with ochre and shells suggest ritual practices predating full Upper Paleolithic expressions, though these sites are stratigraphically borderline.^[119] Recent 2020s research, including surveys at Manot Cave and Nahal Rahaf 2, has expanded evidence of blade and bladelet use across the Levant from 40,000–33,000 BP, highlighting regional variability in lithic reduction strategies and heat treatment for tool enhancement.^[120] Key innovations in these regions include the production of ostrich eggshell beads around 44,000–41,000 BP at Border Cave in South Africa, where perforated shells indicate deliberate ornamentation and possible trade networks among LSA groups.^[121] At the same site, compound adhesives made from beeswax, plant resin, and ochre—dated to about 44,000 BP—were used to haft stone segments onto wooden shafts, demonstrating advanced hafting technology for composite tools.^[121] Evidence of symbolic behavior is limited but growing, with precursors to Upper Paleolithic art in MSA contexts like Blombos Cave (~100,000–70,000 BP), where engraved ochre pieces and shell beads suggest early abstract thinking that likely persisted into later periods.^[122]

Asian and Australian Expansions

The Upper Paleolithic in Asia is marked by early human adaptations to diverse environments, beginning with evidence of settlement in Siberia at the Yana Rhinoceros Horn Site (Yana RHS), dated to approximately 32,000 calibrated years before present (cal BP). This site, located north of the Arctic Circle, reveals a toolkit including lithic artifacts, bone tools, and ivory ornaments, indicating human occupation in extreme cold conditions well before the Last Glacial Maximum.^[123] In East Asia, the emergence of microblade technology around 27,000–26,000 cal BP in North China represents a key innovation, characterized by small, standardized bladelets for composite tools that enhanced hunting efficiency in forested and steppe landscapes.^[124] These developments reflect rapid technological diversification as modern humans expanded eastward from earlier Eurasian populations. Human arrival in Australia, part of the Sahul continent, is estimated at around 65,000 years ago based on archaeological evidence such as at Madjedbebe, though recent genetic analyses as of 2025 suggest approximately 50,000 years ago, reflecting ongoing debate between these lines of evidence; this migration was facilitated by seafaring across Wallacea involving intentional watercraft use to navigate island chains.^[125]^[126] Early colonists adapted to varied ecosystems, with archaeological sites showing ground-edge axes between 44,000 and 49,000 BP used for hunting and possibly tool retouching.^[127] Rock art in Kakadu National Park, including dynamic hunting scenes, dates to over 20,000 years ago, providing insights into symbolic expression and environmental knowledge during the Upper Paleolithic.^[128] Coastal adaptations included advanced fishing gear, such as shell fishhooks and nets evidenced at sites like Jerimalai in East Timor around 42,000 BP, demonstrating pelagic fishing capabilities that supported maritime expansion into Sahul.^[129] Figurative art emerged prominently in Island Southeast Asia, with a warty pig painting in Leang Tedongnge cave on Sulawesi dated to at least 45,500 years ago, featuring therianthropic figures and marking one of the earliest known narrative depictions in the region.^[51] Genetic evidence reveals Denisovan admixture in Asian and Australian populations, contributing up to 4–5% of ancestry in Indigenous Australians and Papuans, likely from interbreeding events during migrations through Southeast Asia around 50,000–40,000 BP.^[130] Recent experimental archaeology supports early seafaring in East Asia, with evidence of human occupation on Japan's Ryukyu Islands by 38,000 BP, achieved via dugout canoes capable of crossing straits despite strong currents.^[131] These expansions highlight adaptive innovations that enabled colonization of isolated landmasses.

American Upper Paleolithic

The American Upper Paleolithic spans approximately 23,000 to 10,000 years before present (BP), a period characterized by the initial human colonization of the Americas and the development of distinct lithic technologies adapted to New World environments. This phase, formally termed in recent scholarship, reflects technological continuities with late Upper Paleolithic traditions in Northeast Asia, particularly the use of microblade and bifacial reduction techniques originating from Siberian contexts. Evidence suggests entry via a Beringian land bridge followed by rapid dispersal, with coastal routes playing a key role in southern migrations during the Last Glacial Maximum.^[19] Key archaeological sites provide direct evidence of early occupations. At Cooper's Ferry in Idaho, artifacts including stemmed projectile points and microblade-like tools date to around 16,000 BP, indicating a late Upper Paleolithic assemblage predating the Clovis complex. The White Sands National Park in New Mexico preserves human footprints dated between 23,000 and 21,000 years ago, confirmed through multiple radiocarbon analyses of associated seeds and pollen, representing the earliest verified human presence in North America. The Gault site in Texas yields pre-Clovis artifacts, such as lanceolate projectile points and blades, with sedimentary ages estimating occupation around 20,000 BP, underscoring a diverse toolkit before the dominance of fluted technologies. Clovis, marking the terminal phase around 13,000 BP, features iconic fluted points but builds on earlier stemmed traditions evident across these locales.^[54]^[132]^[133]^[134] Technological hallmarks include bifacially flaked stemmed points and fluted spear points, often produced through combined core-and-blade reduction methods that parallel Siberian microblade industries. A 2025 analysis of Pacific coastal stone tools from sites like Cooper's Ferry reveals morphological similarities to Hokkaido's late Upper Paleolithic bifaces, supporting a coastal migration route along the Pacific Rim that bypassed ice sheets. Genetic studies further indicate affinities between early American populations and Australasians, with an Australasian ancestry signal detected in coastal South American groups, likely introduced via this maritime pathway during initial dispersals. These elements highlight the American Upper Paleolithic as a period of innovation and adaptation in isolation from Old World traditions.^[19]^[135]^[136]

References

[1]
The absolute chronology of Boker Tachtit (Israel) and implications ...
Jun 14, 2021 · The Initial Upper Paleolithic (IUP) marks a distinct cultural change possibly related to Homo sapiens dispersals into Eurasia.Missing: timeline | Show results with:timeline
[2]
Population size does not explain past changes in cultural complexity
We conclude that new theoretical and empirical research is required to identify the factors that drove the changes in cultural complexity.Models Of Henrich And Powell... · Henrich's Model · 2.2. Population Size...
[3]
Upper Paleolithic - an overview | ScienceDirect Topics
Upper Palaeolithic refers to a period in prehistory characterized by extreme variability in techno-economic patterns, involving the long-distance transport of ...Missing: timeline | Show results with:timeline
[4]
Paleolithic art, an introduction - Smarthistory
Some of the oldest known representational imagery comes from the Aurignacian culture of the Upper Paleolithic period (Paleolithic means old stone age).Paleolithic Art, An... · The Oldest Art... · The CavesMissing: features | Show results with:features
[5]
A Primer on Paleolithic Technology | Learn Science at Scitable
Why study Paleolithic technology? What can old stone tools, ancient fire pits, and painted cave walls tell us about our evolutionary past?Earlier Stone Age (esa) /... · Middle Stone Age (msa) /... · Later Stone Age (lsa) /...
[6]
Early culture shaped by migration and population growth
May 2, 2017 · The Paleolithic was a time marked by periods of slow change separated by bursts of cultural innovation.
[7]
The Chronological Factor in Understanding the Middle and Upper ...
The Middle to Upper Paleolithic transition, dating broadly to between 50,000 and 30,000 years BP, marks a pivotal point in late human evolution. In Europe and ...Missing: definition features
[8]
An Anthropological Review of the Upper Paleolithic in the Southern ...
Traditionally, the local Upper Paleolithic is considered to last about 25,000 years (from ca. 50,000 to 25,000 cal. BP), comprising several archaeological ...
[9]
Homo sapiens Is as Homo sapiens Was : Behavioral Variability ...
Behavioral modernity is a condition characterized qualitatively in terms of the inferred presence or absence of particular “modern human behaviors.” For ...What Is Behavioral Modernity? · Why Is “Behavioral Modernity... · Conclusion
[10]
From hominins to humans: how sapiens became behaviourally ...
Humans became behaviourally modern when they could reliably transmit accumulated informational capital to the next generation.
[11]
Upper Paleolithic Tool Technologies
Jan 3, 2003 · These two tools, blades and burins, opened up a whole new world of wood and bone working with an ease and efficiency never previously matched. A ...
[12]
The revolution that still isn't: The origins of behavioral complexity in ...
At a time when human behavioral 'modernity' was inexorably linked to the Middle to Upper Paleolithic transition in Europe, McBrearty and Brooks (2000) blazed ...
[13]
Recovering Perishable Technologies through Use Wear on Tools
At Vogelherd, dating to at least 32,000 b.p., mammoth ribs were fashioned into long thin needles likely used to sew mats, while some beveled bone points show ...
[14]
Bow-and-arrow, technology of the first modern humans in Europe ...
Feb 22, 2023 · We present the earliest evidence for mechanically propelled projectile technology in Eurasia from Layer E of Grotte Mandrin 54 ka ago in Mediterranean France.
[15]
Big Era 2 - World History for Us All - UCLA
In any case, between about 50,000 to 10,000 BP, the period scholars call the Upper Paleolithic, artistic expression burst forth in many parts of the world. ...
[16]
[PDF] Introduction to Anthropology: Holistic and Applied Research on ... - IUP
The Upper. Paleolithic cultural period, spanning approximately 40,000 to 10,000 years ago, is the last subdivision of the Paleolithic period, also known as the ...Missing: definition timeline
[17]
12 The European Upper Palaeolithic - Oxford Academic
Five broad stages can be discerned over the 35 000 years of the European Upper Palaeolithic; the Initial, Early, Mid, Late and Terminal Upper Palaeolithic.
[18]
Rethinking the structure of the first Upper Paleolithic in Western ...
There remains the enigma of the middle valley of the Rhône, not only having recorded the first migration of H. sapiens into Europe, 10 to 12 millennia prior to ...
[19]
Characterizing the American Upper Paleolithic | Science Advances
Oct 22, 2025 · Between ~22,000 and ~18,000 cal yr B.P., a subset of this population migrated along the southern Beringian and Northwest coasts into the ...<|control11|><|separator|>
[20]
An overview of the West African Paleolithic over the last 200000 years
Feb 1, 2018 · Human responses to changing environments in central Africa between 40,000 and 12,000 B.P ... Upper Paleolithic in the 1990s (Mellars, 1989 ...
[21]
Dating the Paleolithic: Trapped charge methods and amino ... - PNAS
Oct 17, 2022 · This review focuses on two different dating approaches: trapped charge and amino acid geochronology. While differing in their fundamental principles, both ...<|control11|><|separator|>
[22]
Regional impacts of climate change and its relevance to human ...
It is so far the best documented suborbital climate variability and marked by a succession of regional warming–cooling events called Dansgaard–Oeschger (D–O) ...
[23]
Decadal-scale progression of the onset of Dansgaard–Oeschger ...
During their onset, temperatures in Greenland increased rapidly by 10–15 ∘C from cold stadial (GS, Greenland Stadial) to warmer interstadial (GI, Greenland ...
[24]
“Heinrich events” (& sediments): A history of terminology and ...
May 1, 2018 · We document the history of terms used to describe Heinrich (H-) layers and events and which mark major glaciological iceberg discharge ...Missing: Paleolithic | Show results with:Paleolithic
[25]
the Campanian Ignimbrite eruption and the Middle to Upper ...
Early Upper Paleolithic layers above the CI tephra are correlated with HE4 and warm intervals that occurred during 38,000–30,000 cal BP (GI 8–GI 5), and ...
[26]
A new global ice sheet reconstruction for the past 80 000 years
Feb 23, 2021 · One example is the missing ice problem during the Last Glacial Maximum (LGM, 26 000-19 000 years before present) – an apparent 8-28 m ...
[27]
Vegetation of Eurasia from the last glacial maximum to present: Key ...
Feb 1, 2017 · Tundra and steppe biomes dominated 21–14 ka BP, but forest biomes were also present. •. From 14 ka BP woody biomes expanded and wetland ...
[28]
Human population dynamics in Upper Paleolithic Europe inferred ...
Aug 16, 2024 · Some LGM refugia might have been more favorable than others (15 ... Conversely, post-LGM populations in southern and southeastern Europe ...
[29]
Human population dynamics in Europe over the Last Glacial Maximum
Jun 22, 2015 · The Crimean Peninsula contains several important Middle and Upper Paleolithic sites, including Starosele, Kabazi II, and Siuren I. The region ...
[30]
Origin and causes of the mammoth steppe: a story of cloud cover ...
January 2001, Pages 549-574. Quaternary Science Reviews. Origin and causes of the mammoth steppe: a story of cloud cover, woolly mammal tooth pits, buckles ...
[31]
Global sea-level rise in the early Holocene revealed from ... - Nature
Mar 19, 2025 · Here we present an early Holocene sea-level curve based on 88 sea-level data points (13.7–6.2 thousand years ago (ka)) from the North Sea (Doggerland).Missing: Paleolithic | Show results with:Paleolithic
[32]
Doggerland: a Speculative Survey | Proceedings of the Prehistoric ...
Feb 18, 2014 · A speculative reconstruction of the topography, river systems, coastline, vegetation, fauna, and human occupation of Doggerland from the Devensian/Weichselian ...Missing: Paleolithic | Show results with:Paleolithic
[33]
Human settlement history between Sunda and Sahul: a focus on ...
Feb 14, 2015 · The presence of Upper Paleolithic settlers in the region is confirmed by findings in East Timor (>42 kya) [20,27], Borneo (~46 kya) [33] ...
[34]
High-resolution ecosystem changes pacing the millennial climate ...
Aug 1, 2023 · Human-climate interaction during the Early Upper Paleolithic: Testing the hypothesis of an adaptive shift between the Proto-Aurignacian and ...
[35]
Evaluating refugia in recent human evolution in Africa - PMC
We present two alternate models suggesting that between 27% and 66% of Africa may have provided refugia to Late Pleistocene human populations.
[36]
Peştera cu Oase 2 and the cranial morphology of early modern ...
Jan 23, 2007 · The Oase 2 cranium exhibits a suite of derived features of modern humans that contrast with those of the Neandertals and align it with preceding ...
[37]
Appendicular robusticity and the paleobiology of modern human ...
The pattern of changes in humeral diaphyseal robusticity is in good correlation with several other human biological indicators of the degree to which the human ...Materials And Methods · Patterns Of Humeral... · Patterns Of Femoral...
[38]
Genetic contributions to variation in human stature in ... - PNAS
We show that the observed decrease in height between the Early Upper Paleolithic and the Mesolithic is qualitatively predicted by genetics.
[39]
The spread of modern humans in Europe - PNAS
Advocates of the “assimilation model” emphasize human fossils that appear to exhibit a combination of Neanderthal and modern human anatomical traits and ...
[40]
On the origin of modern humans: Asian perspectives - Science
Dec 8, 2017 · The earliest example of fully modern human anatomy in Europe is the mandibular fragment from Kent's Cavern in England and the Cavallo teeth ...
[41]
Bioenergetics, the origins of complexity, and the ascent of man - PNAS
May 5, 2010 · Of all of the African mtDNA variants, only two mtDNAs successfully left Africa and colonized Eurasia, founding macrohaplogroups M and N.
[42]
Contrasting patterns of Y chromosome and mtDNA variation in Africa
Apr 27, 2005 · These data suggest that patterns of differentiation and gene flow in Africa have differed for men and women in the recent evolutionary past.
[43]
Earliest modern human genomes constrain timing of Neanderthal ...
Dec 12, 2024 · Ranis genomes harbour Neanderthal segments that originate from a single admixture event ... Neanderthals that may have continued over several ...
[44]
Climate shifts orchestrated hominin interbreeding events ... - Science
Aug 10, 2023 · Whereas bidirectional gene flow between Denisovans and Neanderthals occurred in Siberia, the few European Neanderthal genomes have shown little ...
[45]
Out-of-Africa, the peopling of continents and islands - PubMed Central
AMH left Africa via a single southern exit about 70 000 years ago and rapidly spread around the Indian Ocean towards the Antipodes.
[46]
Major expansion in the human niche preceded out of Africa dispersal
Jun 18, 2025 · We found that the human niche began to expand substantially from 70 ka and that this expansion was driven by humans increasing their use of diverse habitat ...
[47]
After Last Glacial Maximum: The third migration - ScienceDirect.com
It seems history of population migration repeated itself for the same reasons. Between 70,000 BP and 50,000 BP, modern humans did not move towards west because ...
[48]
Expanding the northern boundary of initial upper Paleolithic dispersal
Aug 6, 2025 · Around 55,000 BP Homo sapiens reached southern parts of China (Hublin, 2021, p. 3), around 45,000 BP modern humans appeared in Europe 19 ...
[49]
The Initial Upper Paleolithic of the Altai: New radiocarbon ...
Selected artefacts from Initial Upper Paleolithic and Middle ... A comparison of the terminal Middle Paleolithic and early Upper Paleolithic in the Levant.
[50]
Oldest cave art found in Sulawesi | Science Advances
Jan 13, 2021 · The oldest, with a minimum age of 45.5 ka, is from Leang Tedongnge. The second image, from Leang Balangajia 1, dates to at least 32 ka.
[51]
Characterizing the American Upper Paleolithic - PMC
Oct 22, 2025 · In 2013, Collins et al. (1) defined seven cultural complexes or “patterns” in the Americas predating the Paleoindian period, which begins with ...Missing: timeline | Show results with:timeline
[52]
Late Upper Paleolithic occupation at Cooper's Ferry, Idaho, USA
Aug 30, 2019 · Cooper's Ferry provides direct evidence for human settlement south of Late Wisconsinan ice sheets in the upper Columbia River basin before the ...
[53]
Palaeogenomics of Upper Palaeolithic to Neolithic European hunter ...
Mar 1, 2023 · These data cover a time span of around 30,000 years from the Upper Palaeolithic to the Late Neolithic (defined here by the presence of pottery ...<|separator|>
[54]
How Many People Have Ever Lived on Earth? | PRB
The global population milestone of 8 billion represents nearly 7% of the total number of people who have ever lived on Earth. Tap to unmute.Missing: Upper Paleolithic 20000 BP
[55]
Historical Estimates of World Population - U.S. Census Bureau
Dec 5, 2022 · View table on historical estimates of the world population.Missing: Paleolithic | Show results with:Paleolithic
[56]
[PDF] A Guide to the Identification of Burins in Prehistoric Chipped Stone ...
They are copies of Paleolithic burins made on prismatic blades and are from European collections. It is very lIDfortunate that reproductions of burins from ...
[57]
Thermal engineering of stone increased prehistoric toolmaking skill
Oct 10, 2019 · Our results confirm that heat treatment does reduce the force needed for flake propagation, they also demonstrate that such treatment has no significant effect ...
[58]
Bohunician technology and thermoluminescence dating of the type ...
Aug 6, 2025 · The Bohunician is the first early Upper Paleolithic technocomplex in the Middle Danube of Central Europe and, therefore, is implicated in ...
[59]
Rethinking the origin of microblade technology: A chronological and ...
May 2, 2016 · Microblade technology was involved in core preparation, systematic knapping, soft hammer and indirect/pressure flaking techniques, which were ...
[60]
Back(s) to basics: The concept of backing in stone tool technologies ...
This term, primarily used for Late MSA, Early LSA, and Early Upper Paleolithic assemblages of Africa and South Asia, generally characterizes assemblages that ...
[61]
[PDF] A Review of Late Pleistocene North American Bone and Ivory Tools
Early Upper Paleolithic approaches to bone and antler projectile technology. In G. L. Peterkin, H. M. Bricker, &. P. Mellars (Eds.), Hunting and animal ...
[62]
Direct evidence for processing Isatis tinctoria L., a non-nutritional ...
May 9, 2025 · By demonstrating that plant processing was not restricted to edible plants, our research provides a new dimension to human behaviour and ...
[63]
About Atlatls - World Atlatl Association
The first known spear throwers come from European Upper Paleolithic sites in France and Spain. Most are from the Magdalenian period (ca 15,000 B.C.), with at ...
[64]
(PDF) Atlatl Dart Velocity: Accurate Measurements and Implications ...
Aug 8, 2025 · We present several different series of experiments with radar gun, film, and chronograph to measure dart velocities.
[65]
Bow-and-arrow, technology of the first modern humans in Europe ...
Feb 22, 2023 · Here, we present the earliest evidence for bow-and-arrow technology in Eurasia from Layer E of Grotte Mandrin in Mediterranean France. These ...
[66]
https://www.researchgate.net/publication/315991940_Atlatl_Dart_Velocity_Accurate_Measurements_and_Implications_for_Paleoindian_and_Archaic_Archaeology
[67]
https://www.science.org/doi/10.1126/sciadv.add4675
[68]
The earliest direct evidence of mammoth hunting in Central Europe ...
The oldest unequivocal evidence of mammoth hunting in prehistoric Central Europe has been found in the Gravettian archaeological site Kraków Spadzista (Poland).
[69]
Reindeer prey mobility and seasonal hunting strategies in the late ...
Jul 22, 2024 · Our reconstructions of prey mobility reveal, for the first time, how reindeer adapted their mobility to the mammoth steppe ecosystem in Central ...
[70]
[PDF] Middle and Upper Palaeolithic Hunting Behaviors in the Southern ...
The study found that Neanderthals and modern humans practiced largely identical hunting tactics, and were equally capable of acquiring biogeographical ...
[71]
https://link.springer.com/article/10.1007/s12520-024-02019-z
[72]
(PDF) Upper Paleolithic Net-Hunting, Small Prey Exploitation, and ...
Aug 7, 2025 · Recent interest in Upper Paleolithic small prey acquisition focuses on the significance of fiber-based hunting technologies.
[73]
The Late Quaternary Megafaunal Extinction and Upper Paleolithic ...
Oct 11, 2025 · We suggest that Upper Paleolithic developments including complex projectile technology, dog domestication, geographic expansion, accelerated ...Missing: culling | Show results with:culling
[74]
Plant foods in the Upper Palaeolithic at Dolní Vӗstonice ...
Aug 10, 2025 · The classic image of Upper Palaeolithic hunter-gatherers in Europe envisages them hunting large mammals in largely treeless landscapes.
[75]
https://www.researchgate.net/publication/396209122_The_Late_Quaternary_Megafaunal_Extinction_and_Upper_Paleolithic_Cultural_Changes_A_Hypothesis_for_Bioenergetic-Driven_Human_Adaptations
[76]
Stable isotope evidence for increasing dietary breadth in the ... - PNAS
New carbon and nitrogen stable isotope values for human remains dating to the mid-Upper Paleolithic in Europe indicate significant amounts of aquatic (fish, ...
[77]
Shellfish consumption in the Early Upper Palaeolithic on the ...
This study shows that the consumption of marine resources (used for diverse purposes) is usual since very ancient chronologies.
[78]
A revised radiocarbon chronology for the mammoth bone structures ...
Jul 25, 2025 · Here we present new radiocarbon dates for a mammoth bone structure at Mezhyrich, Ukraine, refining its timeline. By dating small mammal remains ...Missing: huts | Show results with:huts
[79]
Scraping hide in the early Upper Paleolithic: Insights into the life and ...
Jul 17, 2021 · The 74 endscrapers used in the processing of hide show clear and well-developed traces of use. Rounding, a typical feature of hide use-wear, was ...
[80]
Changes in adaptation from Upper Paleolithic to Incipient Jomon ...
Our study evaluates long-term change in hunter-gatherer mobility on Tanegashima Island from the Upper Paleolithic to Incipient Jomon (ca.36,000–12,800 cal BP).
[81]
(PDF) Responses of Upper Palaeolithic humans to spatio-temporal ...
Responses of Upper Palaeolithic humans to spatio-temporal variations in resources: inequality, storage and mobility. February 2020. DOI:10.17863/CAM.60629.
[82]
The palimpsest problem, hearth pattern analysis, and Middle ...
A principal behavioral issue in paleoanthropology centers on the degree to which the behaviors of Neanderthals and modern humans may have differed.
[83]
Long-distance contacts and social networks of the Upper ...
The occurrence of rare lithic artifacts made from exotic rocks, i.e. brought from distance over 100–200 km to the site, is a highly informative source ...
[84]
(PDF) Long-Distance Imported Lithic Raw Materials at the Upper ...
Jun 11, 2015 · Previous research carried out in this region has indicated the possible use of flint imported from the Moldavian Plateau (over 100km away) ...
[85]
(PDF) Dolní Věstonice-Pavlov. Explaining Paleolithic settlements in ...
Nov 23, 2020 · The case study concerns the site of Milovice IV, a multi-layered Upper Paleolithic settlement belonging to the Gravettian settlement area of ...
[86]
(PDF) Geology and lithic procurement in Upper Palaeolithic Europe
Aug 10, 2025 · Geology and lithic procurement in Upper Palaeolithic Europe: A weights-of-evidence based GIS model of lithic resource potential. April 2010 ...
[87]
Where and When? The distribution of Upper Palaeolithic rock art ...
There are more than 360 known Upper Palaeolithic rock art sites in western Europe. These are widely distributed from Gibraltar to central England and from ...
[88]
Chauvet Cave (ca. 30,000 B.C.) - The Metropolitan Museum of Art
Oct 1, 2002 · Many images of large red dots are, indeed, partial handprints made with the palm of the hand. Red hand stencils and complete handprints have ...Missing: Upper age
[89]
Venus of Willendorf - Smarthistory
The artifact known as the Venus of Willendorf dates to between 24,000–22,000 B.C.E., making it one of the oldest and most famous surviving works of art.
[90]
Perspective: Upper Paleolithic Figurines Showing Women with ...
Dec 1, 2020 · Figurines of obese or pregnant women (“Venus figurines”) from Upper Paleolithic Europe rank among the earliest art, and endured from 38,000 to ...Missing: portable | Show results with:portable
[91]
Portable Art From Upper Paleolithic Period - ThoughtCo
Jun 10, 2025 · Excavations at the Vogelherd Cave recovered several small ivory figurines of several animals; Geissenklösterle cave contained more than 40 ...
[92]
Symbolic innovation at the onset of the Upper Paleolithic in Eurasia ...
Jun 12, 2023 · Modern human incursion into Neanderthal territories 54,000 years ago at Mandrin, France. ... Denisova admixture and the first modern human ...Introduction · Results · Pendant Morphotype<|separator|>
[93]
Prehistoric pigments | Resource - RSC Education
The most notable thing about cave art is that the predominant colours used are black (often from charcoal, soot, or manganese oxide), yellow ochre (often from ...
[94]
HOT (Human Origins Today) Topic - A Colorful Path: What the ...
Dec 14, 2023 · Ochre and mineral pigments have been used for well over 100,000 years by our species, Homo sapiens. These iron-rich, brightly colored rocks ...
[95]
Upper Paleolithic mortuary practices in Eurasia: A critical look at the ...
This paper reviews the available evidence for Upper Paleolithic burials in order to determine what a 'typical' burial from that period looks like.
[96]
Grave Markers : Middle and Early Upper Paleolithic Burials and the ...
Slightly more than threequarters of the burials had associated features (pits, hearths, mounds, stone casings). Most were associated with pits. Mounds and ...
[97]
The Age of the Sunghir Upper Paleolithic Human Burials
ABSTRACT: The earlier Upper Paleolithic site of Sunghir, northern Russia yielded elaborate burials of an adult and of two immature individuals, dug into the ...
[98]
(PDF) New Hydroxyproline Radiocarbon Dates from Sungir, Russia ...
Sungir (Russia) is a key Mid-Upper Palaeolithic site in Eurasia, containing several spectacular burials that disclose early evidence for complex burial ...
[99]
Core-Shell Processing of Natural Pigment: Upper Palaeolithic Red ...
Jul 6, 2015 · From the Upper Palaeolithic record, red ochres are best known for their use in cave paintings and ritual burial contexts.
[100]
Dual ancestries and ecologies of the Late Glacial Palaeolithic in ...
Oct 24, 2022 · Genetic investigations of Upper Palaeolithic Europe have revealed a complex and transformative history of human population movements and ...Missing: variations robust
[101]
Is It Ritual? Or Is It Children? Distinguishing Consequences of Play ...
The cave bear skull was placed on a rock, initiating discussions of a “cave bear cult,” which have long been dismissed. As Bahn (2012) highlights, this ...
[102]
Bone aerophones from Eynan-Mallaha (Israel) indicate imitation of ...
Jun 9, 2023 · Here we report new evidence for Palaeolithic sound-making instruments from the Levant with the discovery of seven aerophones made of perforated bird bones.<|separator|>
[103]
Double child burial from sunghir (Russia): Pathology and inferences ...
The grave goods at Sunghir surpass those found in other known Upper Paleolithic burials, indicating a distinct cultural significance, particularly for child ...
[104]
Overview of the European Upper Palaeolithic: The Homo sapiens ...
Massive migration from the steppe was a source for Indo-European languages in Europe ... Initial Upper Palaeolithic Homo sapiens from Bacho Kiro Cave, Bulgaria.
[105]
Upper Palaeolithic population histories of Southwestern France: a ...
Archaeologically, the Upper Palaeolithic of Southwestern France is divided into five main successive chrono-typological periods or industries; Aurignacian, ...
[106]
A refined chronology for the Gravettian sequence of Abri Pataud
The transition from the EUP to the mid-Upper Paleolithic in Europe (broadly referred to as the Gravettian) is characterized by distinct changes in lithic and ...<|separator|>
[107]
The demography of the Upper Palaeolithic hunter–gatherers of ...
Archaeologically, the Upper Palaeolithic of Southwestern France is divided into five broad successive periods; Aurignacian (∼39,500–34,000 cal BP), Gravettian ( ...
[108]
Humans confront the Last Glacial Maximum in Western Europe
Beginning in the Early Upper Paleolithic of Europe ... Various forms thereof are present in Aurignacian, Gravettian, Solutrean and Magdalenian assemblages.
[109]
Territories and economies of hunter–gatherer groups during the last ...
After the last glacial maximum, in a cold and dry climate, the European hunter–gatherers produced a Magdalenian industry in Western and Central Europe and an ' ...
[110]
The Cave Art Paintings of the Lascaux Cave - Bradshaw Foundation
Lascaux is famous for its Palaeolithic cave paintings, found in a complex of caves in the Dordogne region of southwestern France.
[111]
Reindeer Hunters of the Ice Age in Europe: Economy, Ecology, and ...
Economy of animal resources and mobility of human groups during Upper Pleniglacial and Late Glacial (30,000 - 14,000 cal BP) in France: a « Reindeer system » at ...
[112]
The Beginning of the Early Upper Paleolithic in Poland
May 1, 2023 · In this paper, we examine this technical diversity by reassessing and revising the chronological and technological information of the key Aurignacian sites in ...
[113]
Nobody's land? The oldest evidence of early Upper Paleolithic ...
Jun 26, 2024 · The Abrigo de la Malia provides irrefutable evidence of human settlements dating back to 36,200 to 31,760 calibrated years before the present ( ...
[114]
Riddles wrapped inside an enigma. Lupemban MSA technology as ...
Mar 7, 2022 · Early Stone Age (ESA) archaeology is effectively absent from the rainforest zone, with the early Middle Stone Age (MSA) Lupemban industry ...
[115]
Border Cave and the beginning of the Later Stone Age in South Africa
... technology and tool types of the post-Howiesons Poort period and can be considered transitional industries; (ii) the 44-42 ka cal BP assemblages are based ...Missing: Lupemban Stillbay bladelet North
[116]
Origins of the Iberomaurusian in NW Africa: New AMS radiocarbon ...
The Iberomaurusian lithic industry is typified by microlithic backed bladelets and, apart from its geographically wide distribution, is significant because it ...
[117]
The Upper Paleolithic (Chapter 5)
“Microliths” or “geometric microliths” are blades/bladelets that have been extensively modified by backing and/or truncation. Micro-liths occur in small numbers ...
[118]
[PDF] ESR dates for the hominid burial site of Qafzeh in Israel - In Africa
Early modern hominids are found buried at the mouth of a cave at Qafzeh near Nazareth, Israel. They are associated with a Middle Paleolithic lithic.
[119]
Cultural Dynamics in the Levantine Upper Paleolithic, ca. 40–33 ky BP
Apr 29, 2024 · This paper presents a research synthesis of two case studies: Manot Cave, located in western Galilee, Israel, and Nahal Rahaf 2 Rockshelter in the Judean ...
[120]
Early evidence of San material culture represented by organic ...
... Africa, as well as the Near East, had developed key cultural innovations by at least 80,000 y ago. These innovations include systematic use of red pigment ...
[121]
Nassarius kraussianus shell beads from Blombos Cave: evidence ...
Since 1991, excavations at Blombos Cave have yielded a well-preserved sample of faunal and cultural material in Middle Stone Age (MSA) levels.Missing: though | Show results with:though
[122]
The Yana RHS site: humans in the Arctic before the last ... - PubMed
A newly discovered Paleolithic site on the Yana River, Siberia, at 71 degrees N, lies well above the Arctic circle and dates to 27,000 radiocarbon years ...
[123]
The development of early microblade technology in North China
Lithic analysis reveals that the earliest microblade technology emerged in North China at ∼27–26 cal. ka BP. The earliest microblades exhibit precocious ...
[124]
Neanderthal DNA may refute 65,000-year-old date for ... - Live Science
Jul 3, 2025 · A new DNA model suggests humans didn't reach Australia until 50000 years ago, but archaeological data disagrees.Australia's oldest rock painting... · 'Huge surprise' reveals how...
[125]
Retouch-induced marks on Australian hardwood boomerangs
Our findings constitute the first traceological identification of hardwood boomerangs being used as retouchers in various Aboriginal Australian contexts.
[126]
Carvings and paintings, Kakadu - Smarthistory
Kakadu's carvings and paintings, some up to 20,000 years old, record the region's history, with some using 'x-ray painting' at Ubirr.Missing: Upper | Show results with:Upper
[127]
(PDF) Pelagic Fishing at 42,000 Years Before the Present and the ...
Aug 6, 2025 · Here we report remains of a variety of pelagic and other fish species dating to 42,000 years before the present from Jerimalai shelter in East ...
[128]
Widespread Denisovan ancestry in Island Southeast Asia but no ...
Mar 22, 2021 · Our results corroborate widespread Denisovan ancestry in ISEA populations, but fail to detect any substantial super-archaic admixture signals.
[129]
Paleolithic seafaring in East Asia: An experimental test of the dugout ...
Jun 25, 2025 · In this study, we test if a non-sailing dugout canoe can be produced with Upper Paleolithic tools, and if it can cross the 110-kilometer-wide strait.
[130]
Evidence of humans in North America during the Last Glacial ...
Sep 23, 2021 · If 25% or more of the Ruppia seeds were redeposited, the age of these human footprints might be exaggerated by several millennia—with ages more ...
[131]
Evidence of an early projectile point technology in North America at ...
Jul 11, 2018 · This paper presents age estimates from a series of alluvial sedimentary samples from the earliest cultural assemblage at the Gault Site, Central Texas.
[132]
The age of Clovis—13,050 to 12,750 cal yr B.P - PMC - NIH
Oct 21, 2020 · The Clovis complex dates from 13050 to 12750 cal yr B.P. during a time of major environmental change.Missing: phase | Show results with:phase
[133]
Stone tools trace Paleolithic Pacific migration - Oregon State University
Oct 23, 2025 · ... American Upper Paleolithic. They showed that the earliest of this style of projectile points first appear around 20,000 years ago in Hokkaido.Missing: Clovis | Show results with:Clovis
[134]
Pacific & Amazonian Natives: Genetic Affinity & Australasian Ancestry
Mar 29, 2021 · We show the Australasian genetic signal is present in the Pacific coast region, indicating a more widespread signal distribution within South America.