Fact-checked by Grok 2 weeks ago

Capitulum of the humerus

The capitulum of the humerus, also known as the capitellum, is a smooth, rounded eminence forming the lateral portion of the distal articular surface of the , which articulates with the head of the to create the as part of the elbow complex. Located on the inferolateral aspect of the humeral condyle, the capitulum presents a convex, knob-like projection that covers the anterior and inferior surfaces of the bone but spares the posterior aspect, and it is separated from the adjacent medial trochlea by a shallow groove. This structure facilitates forearm flexion and extension by enabling the to pivot smoothly against the during motion, contributing to overall joint stability and the transmission of forces in the . Above the capitulum lies the radial , a shallow depression that accommodates the radial head during full flexion of the . Clinically, the capitulum is significant due to its vulnerability to intra-articular fractures, which often result from high-energy such as falls on an outstretched hand or direct impacts, particularly affecting adolescents and young adults whose centers are developing. These fractures, classified into types based on fragment size and (e.g., Hahn-Steinthal or Kocher-Lorenz patterns), can lead to complications like , , or elbow stiffness if not managed with timely surgical intervention such as open reduction and . As the first in the distal —typically appearing between ages 1 and 2—the capitulum serves as a key radiographic landmark for assessing skeletal maturity and diagnosing pediatric injuries.

Overview and Definition

Structure and Location

The is a smooth, rounded eminence situated on the lateral portion of the distal articular surface of the , forming a convex projection that contributes to the overall condylar structure. In tetrapods, this feature is consistently positioned on the anterior aspect of the distal , providing a key component of the elbow region's bony architecture. In humans, the capitulum forms part of the humeral condyle and is oriented anteriorly and inferiorly at approximately 30 degrees relative to the humeral shaft, with a of about 1-2 in adults. Medially, it is adjacent to the trochlea, from which it is separated by a shallow groove that delineates the two articular surfaces. Proximally, the capitulum is continuous with the radial fossa, a depression on the anterior humerus. Anatomically, the capitulum overlies the radial fossa anteriorly, accommodating the proximal radius during joint flexion, and lies medial to the lateral epicondyle, which serves as an attachment site for muscles.

Etymology and Terminology

The term capitulum originates from the Latin capitulum, a form of caput meaning "head," thereby denoting a "small head." This directly reflects the anatomical feature's characteristic rounded, knob-like prominence on the distal , evoking the image of a diminutive cranial structure. In anatomical , capitulum humeri is the preferred Latin term, with the synonymous capitellum employed interchangeably in various texts to describe the same structure. The structure is codified in modern standardized systems, including the (TA98 identifier: A02.4.04.022; TA2 identifier: 1202) and the (FMA identifier: 23373), ensuring consistent usage across international anatomical references. Historically, the capitulum was detailed as a component of the distal in (20th edition, 1918), where it is portrayed as the lateral, rounded eminence articulating with the . Earlier allusions trace to 16th-century anatomy, notably in ' De humani corporis fabrica (1543), which advanced descriptive precision for humeral features within the tradition. To avoid confusion, the humeral capitulum must be differentiated from homonymous terms in other contexts, as well as extraneous applications like the botanical capitulum (a compact flower head) or liturgical (a short scriptural reading).

Anatomy in Mammals

Human Anatomy

In , the capitulum of the humerus, also known as the capitellum, is a smooth, rounded eminence forming the lateral portion of the distal humerus's articular surface. It measures approximately 15-20 mm in anteroposterior diameter in adults, with studies reporting average widths of about 15.55 mm and depths around 9 mm, contributing to its role in precise without delving into functional aspects. begins from a secondary center that appears between 1 and 2 years of age, typically around 1 year, and fuses with the humeral shaft by 12-14 years, though fusion can occur as early as 10-12 years in some cases. The vascular supply to the capitulum primarily arises from perforating vessels of the recurrent , with contributions from the radial collateral and interosseous recurrent arteries entering posteriorly, ensuring nutrient delivery to the epiphyseal region. Innervation is provided by branches of the , which supply the surrounding lateral epicondylar region and periarticular tissues. Gender variations show the capitulum to be slightly larger in males, consistent with overall distal humeral dimensions being more prominent in males across multiple morphometric parameters. With age, the structure exhibits minimal atrophy, maintaining relative stability into adulthood, though it remains susceptible to in adolescents, particularly affecting the dominant arm in active individuals. On , the capitulum appears as a rounded osseous density on lateral radiographs of the , best visualized in true lateral projections where it projects anteriorly relative to the humeral shaft. (MRI) reveals its coverage by articular , typically 1-2 mm thick, with T2-weighted sequences highlighting the cartilage's high signal intensity against the low-signal bone, aiding in assessment of surface integrity.

Comparative Features in Other Mammals

In quadrupedal mammals such as , the capitulum of the humerus is typically more elongated and cylindrical compared to humans, facilitating greater stability during . This supports a stable at the , with the capitulum articulating less prominently with the to emphasize load distribution across the . Among non-human like chimpanzees, the capitulum exhibits a more globular akin to that in humans, but with enhanced and smoother contours that permit greater radial and pronation-supination for arboreal and manipulative activities. This reflects the demands of suspensory and behaviors, where the capitulum's rounded form allows for increased forearm rotation without compromising joint integrity. Three-dimensional analyses confirm that the capitulum in great apes, including chimpanzees, has a larger articular area and more acute angles relative to cercopithecoids, correlating with orthograde postures. In mammals such as whales, the capitulum is markedly reduced and often flattened or rudimentary, reflecting the of flipper-like s with minimal mobility. The in cetaceans is short and robust, with the capitulum and associated structures fused to the and , eliminating distinct articulations and prioritizing hydrodynamic stability over terrestrial function. This vestigial form aligns with the loss of medullar cavity and overall forelimb simplification in fully lifestyles. Arboreal species like squirrels display a prominent capitulum that is small yet rounded and well-developed for enhanced rotation and flexibility in . The humerus in these is long and slender, with the capitulum supporting agile pronation-supination to navigate branching substrates, distinguishing it from more rigid forms in ground-dwelling relatives. Morphometric studies highlight how this capitular prominence scales with locomotor , promoting rotational freedom in tree-dwelling habits. Across mammals, capitulum size generally scales positively with body mass, as seen in comparative analyses of humeral geometry where larger species exhibit proportionally broader articular surfaces for load-bearing efficiency.

Function and Biomechanics

Articulation with Radius

The is formed by the articulation between the rounded capitulum on the lateral aspect of the distal and the fovea of the head of the , creating a synovial with ball-and-socket-like characteristics that facilitates forearm flexion-extension and . This is part of the broader complex and contributes to the overall stability and mobility of the proximal . The convex curvature of the capitulum precisely matches the shallow concavity of the radial head's fovea, ensuring congruent contact during motion. The articulating surfaces are both covered by a layer of articular , which varies in thickness from approximately 0.4 to 1.8 mm across the distal , providing a smooth, low-friction interface. This layer on the capitulum and radial head helps distribute compressive forces and absorb shock during joint loading. The lining the joint cavity secretes , a viscous rich in , which further reduces friction and nourishes the avascular . Stability of the humeroradial articulation is primarily provided by surrounding ligaments, including the annular ligament, which forms a strong fibrous band encircling about 80% of the radial head and attaching to the anterior and posterior margins of the ulnar radial notch, thereby holding the radial head securely against the capitulum. The radial collateral ligament, originating from the , blends with the annular ligament and reinforces the lateral aspect of the joint against varus stresses. Additionally, the envelops the articulation, being notably thin anteriorly (to allow motion) and thicker laterally where it integrates with the collateral structures for enhanced reinforcement.

Role in Elbow Joint Movements

The plays a pivotal role in facilitating pronation and supination of the by providing a spherical surface for the radial head, allowing rotation around a longitudinal axis. During these movements, the superior surface of the radial head rotates directly against the capitulum, while the ridge of the radial head glides within the groove separating the capitulum from the trochlea, enabling a total of approximately 160–180 degrees in humans (typically 80 degrees of pronation and 90 degrees of supination from neutral). This pivot-like interaction at the ensures smooth rotation without significant translation, contributing to the precision required for hand positioning in daily activities. In elbow flexion and extension, the capitulum supports the rolling and gliding arthrokinematics of the radial head, which translates along its surface to maintain . Flexion, ranging from 0 to approximately 145 degrees, involves an anterior (volar) glide and roll of the radial head on the capitulum, allowing the to approach the while distributing forces. Conversely, extension features a posterior () glide of the radial head, with rolling prominent at the extremes of motion to prevent excessive tension on surrounding soft tissues. These complement the primary action at the ulnohumeral , enhancing overall stability during movements. The capitulum also contributes to load distribution and stability in the complex, bearing a significant portion of axial compressive forces through the radiocapitellar . In extension under axial loading, the radiocapitellar transmits approximately 60% of the total load, with the remainder borne by the ulnotrochlear , which helps prevent overload on the during weight-bearing activities. Additionally, the capitulum aids in resisting varus stress by forming part of the lateral , where up to 75% of varus at 90 degrees of flexion derives from bony and compressive forces across the radiocapitellar , supplemented by the lateral ligament complex. Biomechanically, the capitulum's rounded geometry influences dynamics around the elbow's lateral , where simplified calculations (\tau = F \times d)—with \tau as , F as applied , and d as from the capitulum's rotation —determine stress thresholds for joint integrity. Excessive varus , often exceeding 50 Nm in high-impact scenarios, can overwhelm these , leading to radiocapitellar by disrupting the radial head's centered position on the capitulum. This underscores the capitulum's role in balancing rotational and compressive loads to safeguard against .

Comparative Anatomy in Reptiles

Lepidosauromorpha

In lepidosaurs, which include squamates ( and snakes) and rhynchocephalians (), the capitulum of the forms part of the distal articular surface, characterized by a distinct, rounded capitellum for radial positioned ventrally and preaxially, adjacent to a shallower trochlea for the . These structures are separated by a shallow sulcus, with the capitellum appearing relatively smaller in proportion to the overall compared to more erect-limbed vertebrates, reflecting adaptations to a sprawling . In lizards such as the (Iguana iguana), the capitulum supports the sprawling typical of lepidosaurs, facilitating lateral undulation and weight-bearing during while permitting only limited humeral at the . This configuration allows for protraction and retraction in a primarily horizontal plane, with kinematic studies in species like the (Pogona vitticeps) showing restricted long-axis to maintain during slow walking and faster gaits. Among , which represent the limbless extreme within , the and its capitulum are vestigial or entirely absent in adults due to secondary limb reduction, though embryonic remnants may persist briefly before resorption. This loss aligns with the evolution of serpentine , eliminating the need for articulations. The capitulum is notably present in basal lepidosaurs like the (Sphenodon punctatus), where it exhibits a moderately expanded form aiding in lateral flexion of the , a trait conserved from early rhynchocephalians and contributing to subtle rotational movements during and . In these taxa, the articular at the capitulum enhances durability for terrestrial stresses as described in foundational osteological analyses.

Archosauromorpha

In archosauromorphs, the capitulum of the exhibits modifications adapted to diverse locomotor strategies, particularly in crocodilians and birds as extant representatives, with fossil evidence illuminating variations in extinct forms like dinosaurs. In crocodilians, the distal features distinct capitellar (lateral) and trochlear (medial) condyles separated by a shallow intercondylar groove, which accommodates the and during flexion and extension. This configuration supports a semi-erect posture during terrestrial "high walks" and galloping, enabling greater limb elevation and stability compared to sprawling lepidosaurs, while maintaining amphibious versatility. The groove's shallow depth reflects the 's limited rotation, prioritizing robust load-bearing for quadrupedal support in predatory ambushes. In birds, the capitulum is reconfigured into a prominent condyle equivalent, which articulates with both the and to facilitate folding essential for perching and powered flight, while the ventral capitulum is notably reduced to minimize mass and drag. This emphasis allows the antebrachium to fold tightly against the during rest, enhancing aerodynamic during soaring and . In raptorial species like eagles, the condyle is elongated to provide a pivotal that supports precise adjustments for dives and maneuvers. The intercondylar incisura between dorsal and ventral portions further aids in constraining motion to protraction and retraction, integrating seamlessly with the carpus for coordinated extension in flight—contrasting the more hinge-like mammalian by permitting greater rotational freedom at the . Variations among extinct archosauromorphs, particularly dinosaurs, highlight evolutionary trends toward enhanced functionality. These adaptations underscore archosauromorph shifts from sprawling to erect postures, optimizing the for aerial and terrestrial innovations absent in mammalian lineages.

Development and Evolution

Embryological Development

The capitulum of the arises from the lateral aspect of the distal humeral chondrification center, which forms during weeks 6-7 of as part of the initial mesenchymal condensations in the developing limb bud. By week 8, this center differentiates further, with the capitellar anlage emerging as a distinct precartilaginous structure within the lateral condyle, coinciding with the recognition of the interzone. This early separation allows for the specific shaping of the capitulum as a rounded eminence for radial . Chondrogenesis of the capitulum involves the condensation of mesenchymal cells into a cartilaginous template, regulated by (Hox) genes that pattern the proximodistal axis of the limb. Specifically, Hoxa11 plays a key role in influencing distal limb patterning and promoting differentiation by acting upstream of transcription factors such as , ensuring proper formation in the distal . These genetic controls integrate with signaling pathways like and Ihh to drive the maturation of the cartilaginous anlage. During prenatal growth, the capitellar undergoes phased expansion, with the hypertrophic zone of chondrocytes enlarging and mineralizing the surrounding matrix to prepare for . This process transitions via , where the cartilage model is gradually replaced by , beginning at birth with the appearance of the capitellar around 1-12 months postnatally. The hypertrophic zone's expansion thus bridges embryonic cartilage to postnatal bony maturation. Congenital anomalies such as capitellar aplasia are rare and typically manifest as isolated absences of the structure, potentially leading to instability. These defects have been linked to disruptions in limb patterning pathways during embryogenesis.

Evolutionary Origins and Variations

The first appeared as a small eminence on the distal end of the in early tetrapods during the Late to fin-to-limb transition, facilitating the of propped postures and limited terrestrial support from aquatic fin-like structures. In and early Permian tetrapods, such as the temnospondyl megacephalus, the capitulum enlarged significantly, dominating the lateral portion of the distal and enabling greater flexion for weight-bearing on land; for example, in specimens, the capitulum width measures approximately 33.8 mm relative to a distal humeral width of 93.7 mm. This enlargement represents a key in the progressive of the among temnospondyls, contrasting with the ancestral condition of a short featuring a small capitulum and separate ulnar condyle. By the Permian period, the capitulum was fully present in synapsids, the stem group to mammals, though it remained relatively small and permitted restricted elbow mobility compared to later forms. In basal synapsids such as pelycosaurs (e.g., ), the capitulum is notably reduced, with a width of about 22.5 mm on a of distal width 102 mm, reflecting a transitional morphology adapted to sprawling gaits and partial aquatic habits. Fossil evidence from Permian pelycosaur humeri, including those of eupelycosaurs like Gordodon kraineri, shows variations in distal flattening and capitular prominence, bridging earlier designs toward more derived configurations. In therapsids, the mammalian ancestors arising in the late Permian, the capitulum underwent refinement, becoming more rounded and integrated with a twisted humeral shaft to support increasingly parasagittal postures associated with enhanced metabolic efficiency. This shift paralleled broader postcranial changes in synapsids, where variability increased dramatically by the late Permian, predating dominance and enabling diverse locomotor modes. Within diapsids, the sister to synapsids, the capitulum diverged by the into distinct forms: more pronounced in lepidosauromorphs for flexible, sprawling locomotion, and variably reduced or specialized in archosauromorphs, as seen in early forms like Prolacerta. Across Amniota, the capitulum is a conserved feature, with its morphology tracking ecological adaptations, from the sprawling limbs of basal amniotes to the specialized extremities of derived clades.

Clinical and Pathological Aspects

Fractures and Injuries

Capitellar fractures are uncommon injuries, accounting for less than 1% of all fractures and approximately 6% of distal humeral fractures. They occur in adolescents and adults, typically resulting from high-energy such as falls on an outstretched hand, though low-energy mechanisms can also contribute. In the AO/OTA classification system, these are categorized as partial articular fractures in the , specifically type 13-B3. The primary types include coronal shear fractures, often referred to as Hahn-Steinthal fractures, which involve a large osseous fragment with attached subchondral ; avulsion fractures (Kocher-Lorenz type), characterized by a smaller fragment limited to articular ; and more complex variants such as comminuted (type III) or those extending into the trochlea (type IV McKee). is defined as greater than 2 mm and typically requires surgical intervention to restore joint congruity. The mechanism of injury generally involves hyperextension of the combined with a varus force, where the radial head shears against the capitellum during axial loading from a fall on the outstretched hand. Approximately 50% of cases are associated with radial head or , which complicates stability and necessitates careful assessment. Initial management depends on displacement and fragment size. Nondisplaced fractures with less than 2 mm of separation can be treated conservatively with in a long-arm cast for 3 to 4 weeks, followed by protected range-of-motion exercises. For displaced fractures, open reduction and (ORIF) is the standard approach, using headless compression screws such as Herbert screws to secure capitellar fragments while preserving the subchondral surface and avoiding intra-articular hardware prominence.

Associated Conditions and Imaging

Panner's disease, also known as osteochondrosis of the capitellum, represents of the capitellar primarily affecting children aged 5 to 10 years, often those engaged in due to repetitive valgus stress. This self-limiting condition typically resolves spontaneously with conservative management, including rest and activity modification, leading to full recovery without residual deformity in most cases. Osteochondritis dissecans (OCD) of the capitellum is a more common non-traumatic pathology in adolescents, particularly overhead athletes aged 12 to 16 years, resulting from repetitive microtrauma and vascular compromise to the subchondral bone. Lesions most frequently involve the anterolateral aspect of the capitellum, accounting for the majority of cases and causing lateral elbow pain, swelling, and limited extension. Recent advancements as of 2025 include arthroscopic drilling and fixation techniques for stable lesions to promote revascularization. In , the capitellum is a frequent site of erosive changes due to chronic , with radiographic erosions commonly observed on the capitellar surface alongside the lateral . These erosions contribute to progressive destruction and instability, occurring in 20% to 65% of rheumatoid patients with involvement. Tumors of the capitellum are exceedingly rare, comprising less than 1% of primary neoplasms, with chondroblastoma serving as a representative example of an epiphyseal benign that can mimic OCD through lytic destruction and . Such tumors typically arise in the second decade and require histologic confirmation for diagnosis. Diagnostic imaging plays a crucial role in evaluating capitellar pathologies. Plain radiographs, particularly lateral views, assess alignment and detect joint effusion via the sail sign, where the displaced anterior fat pad appears elevated. Computed tomography (CT) provides detailed evaluation of lesion size and fragment displacement in OCD, with three-dimensional mapping revealing typical surface areas of 50 to 200 mm² in the posterolateral capitellum. Magnetic resonance imaging (MRI) excels in assessing cartilage integrity, using T2-weighted sequences to identify defects greater than 5 mm and subchondral instability through high-signal rims or cysts. Prognosis for capitellar OCD is guided by the Nelson classification, which stages lesions from I (intact with subchondral changes) to IV (displaced fragment or loose body). Stable stages often heal with nonoperative care, while stage III lesions, indicating partial detachment, typically require to stimulate and promote union.

References

  1. [1]
    Capitulum of humerus: Anatomy and function - Kenhub
    Mar 11, 2024 · The capitulum of humerus is a rounded knob-like structure on the distal humerus, which articulates with the head of radius.
  2. [2]
    Capitulum of humerus - e-Anatomy - IMAIOS
    The lateral portion of the articular surface of humerus consists of a smooth, rounded eminence, named the capitulum of the humerus.
  3. [3]
    Capitulum of Humerus | Complete Anatomy - Elsevier
    The capitulum (capitellum) is the smooth, convex projection found on the inferolateral aspect of the condyle of the humerus.
  4. [4]
    The Humerus - Proximal - Shaft - Distal - TeachMeAnatomy
    Oct 29, 2024 · Lateral to the trochlea is the capitulum, which articulates with the radius. Also located on the distal portion of the humerus are three ...
  5. [5]
    Capitellum Fractures - Trauma - Orthobullets
    Oct 18, 2023 · Capitellum Fractures are traumatic intra-articular elbow injuries involving the distal humerus at the capitellum.
  6. [6]
    Clinical Outcomes and Management Strategies for Capitellum ... - NIH
    Jun 26, 2024 · The treatment of capitellum and trochlea fractures yields satisfactory outcomes but has a considerable rate of complications and reoperations.
  7. [7]
    Capitellum - an overview | ScienceDirect Topics
    The capitellum, the first of the elbow's six centers to ossify, generally becomes radiographically visible during the first and second years of life.
  8. [8]
    Bones of the Upper Limb – Anatomy & Physiology - UH Pressbooks
    Immediately lateral to the trochlea is the capitulum (“small head”), a knob-like structure located on the anterior surface of the distal humerus. The capitulum ...Humerus · Carpal Bones · Phalanx Bones
  9. [9]
    Anatomy, Shoulder and Upper Limb, Humerus - StatPearls - NCBI
    Clinical Significance. Radial Nerve Injury. Radial nerve palsy is among the most common peripheral nerve injuries following a humeral fracture.[44] Typically ...
  10. [10]
    [PDF] Capitellum Fractures - Thieme Connect
    lum is directed in an anterior direction about 30 degrees with respect to ... Unusual fractures of the capitulum humeri and the capitulum radii. J Bone ...<|control11|><|separator|>
  11. [11]
    Etymology of Forearm, Wrist and Hand Terms
    Capitulum and its synonym Capitellum - Both are diminutives of the Latin word caput, meaning head, and so mean a little head.
  12. [12]
    Capitellum | Radiology Reference Article - Radiopaedia.org
    Jan 25, 2020 · The capitellum, also referred to as the capitulum, is the lateral part of the humeral condyle that articulates with the radial head.<|control11|><|separator|>
  13. [13]
    Capitulum - Etymology, Origin & Meaning
    Originating from Latin capitulum, meaning "little head," this word denotes a "leader," "summit," "division," or "principal sum," reflecting its diverse ...
  14. [14]
    Capitellum - an overview | ScienceDirect Topics
    Capitellum is defined as the original anatomic name for the capitulum, which ... The capitellum is directed 30 degrees anteriorly and distally with ...
  15. [15]
    capitulum humeri - TA2 Viewer
    FMA, 23373. UBERON, 0010853. UMLS, C0223688. Gray's Anatomy (20th ed.) page 212. Wikipedia, Capitulum of the humerus. Wikidata, Q718149. Wikipedia sites (7).
  16. [16]
    6a. 3. The Humerus - Collection at Bartleby.com
    The humerus (Figs. 207, 208) is the longest and largest bone of the upper extremity; it is divisible into a body and two extremities.
  17. [17]
    The rich heritage of anatomical texts during Renaissance and ... - NIH
    The practice of modern human anatomy was started by Vesalius in sixteenth century Europe during the Renaissance. His exploits are documented in his legendary ...
  18. [18]
    CT-based measurement and analysis of distal humerus morphology ...
    This study aims to investigate the morphological characteristics of the distal humerus in healthy adults from northern China using computed tomography and ...
  19. [19]
    Elbow ossification | Radiology Reference Article | Radiopaedia.org
    Aug 31, 2025 · Age ; capitellum: 1 year ; radial head: 3 years ; internal epicondyle: 5 years ; trochlea: 7 years ; olecranon: 9 years.Mnemonic · Elbow radiograph - age eleven · Question 40
  20. [20]
    The extraosseous and intraosseous arterial anatomy of ... - PubMed
    The capitellum and the lateral aspect of the trochlea were supplied by posterior perforating vessels arising from the radial recurrent, radial collateral, and ...<|control11|><|separator|>
  21. [21]
    CT-based measurement and analysis of distal humerus morphology ...
    Oct 1, 2024 · Regarding gender differences, 21 parameters were significantly more prominent in males than females (P < 0.05*). The distal humerus of males was ...
  22. [22]
    Pitfalls in Elbow Imaging: Osseous Anatomic Variants
    Jan 3, 2023 · The radial head has a central concavity that allows smooth articulation with the rounded, anteriorly directed capitellum or capitulum of the ...
  23. [23]
    Elbow Imaging with an Emphasis on MRI - NCBI - NIH
    Apr 13, 2021 · This article will discuss the normal anatomy of the elbow and the most common types of elbow pathology as well as their appearances on various imaging studies.Missing: capitulum | Show results with:capitulum
  24. [24]
    Canine Anatomy - Veterian Key
    Jul 8, 2016 · The proximal surface of the radius articulates with the humeral capitulum, which is not as prominent as in the human. The canine distal radius ...
  25. [25]
    [PDF] Animal Anatomy
    COMPARATIVE ANATOMY OF THE HUMERUS. Cattle. The humerus of cattle is smoother than that of equines. The musculospiral groove is shallower and the deltoid ...Missing: chimpanzees whales squirrels rodents
  26. [26]
    [PDF] OSTEOLOGY OF THE MAMMALIA - Mike Taylor
    WILLIAM HENRY FLOWER, LL.D., F.R.S.. DIRECTOR OF THE NATL'RAL HISTORY DEPARTMENTS OF THE BRITISH MUSEUM. LATE HUNTERIAN PROFESSOR OF COMPARATIVE ANATOMY ...
  27. [27]
    Capitular morphology in primates and fossils: 3-D measurements of ...
    Using a sample of 43 great apes, 59 cercopithecids, and 31 humans, 3D images were used to take measurements of the area, length, and angle of the capitulum to ...
  28. [28]
    Phylogeny - Musculoskeletal Key
    Aug 28, 2016 · Only in the hominoid primates, which include humans, chimpanzees, gorillas, orangutans, and gibbons, is the medial aspect of the distal ...
  29. [29]
    Comparative Anatomy - New Bedford Whaling Museum
    Cetacean flippers have the same bones as the forelimbs of other mammals. The humerus, radius, ulna, carpals, metacarpals and phalanges are all present.Missing: capitulum chimpanzees squirrels rodents
  30. [30]
    Cetaceans Humerus Radiodensity by CT: A Useful Technique ... - NIH
    Jul 13, 2022 · In cetaceans, the humerus is a relatively short bone that lacks the medullar cavity. However, its bone compartments, cortical and trabecular, ...
  31. [31]
    Size And Locomotor Ecology Have Differing Effects on the External ...
    Humeral external shape​​ PC1 accounted for 36.8% of the humeral shape variation and largely separates the morphospaces among ground squirrels, chipmunks and tree ...
  32. [32]
    Integrative Approach Uncovers New Patterns of Ecomorphological ...
    Dec 15, 2021 · In this work, we first partitioned humeral and femoral anatomy into single traits and identified those that correlate with the slow arboreal ...<|separator|>
  33. [33]
    Locomotory Adaptations in 3D Humerus Geometry of Xenarthra
    The humerus is a load-bearing postcranial element in quadrupedal mammals (Bertram and Biewener, 1992) and correlates quite strongly with body mass, locomotory, ...
  34. [34]
    Humerus - an overview | ScienceDirect Topics
    The humerus reaches 2.2 and 3.3 mm at 49 and 53 days, respectively, and at the end of the embryonic period it is about 4.8 mm in length (Streeter, 1949; O' ...
  35. [35]
    Appendicular Morphology
    Humerus (Figure 2, Figure 3). The overall form of the humerus of ... fusion in rodents is opposite the distal fusion of Protypotherium). Both ...
  36. [36]
    Elbow joint: Anatomy, ligaments, movements, blood supply | Kenhub
    Did you know that the elbow is a synovial hinge joint? Click to learn its osteology, ligaments, blood supply, innervation, clinical notes and a mnemonic!Missing: capitellum | Show results with:capitellum
  37. [37]
    Elbow - Physiopedia
    ... capitulum of the humerus. The head rotates within the annular ligament to ... 30 degrees Hyper-extension 10-15 degrees Radial or medial deviation 15-20 ...
  38. [38]
    Humeroradial Joint - an overview | ScienceDirect Topics
    The humeroradial joint is a pivot joint resembling a ball-and-socket joint. The proximal and distal radioulnar joints are mirror images, allowing the radius to ...
  39. [39]
    Cartilage thickness of distal humerus and its relationships with bone ...
    Results: Cartilage thickness showed a significant variation (range, 0.4-1.8 mm) independent of sex and side. It appeared thinner at the medial and lateral edges ...Missing: humeroradial | Show results with:humeroradial
  40. [40]
    Anatomy, Shoulder and Upper Limb, Elbow Annular Ligament - NCBI
    The annular ligament stabilizes the radial head within the radial notch of the ulna as it rotates during supination and pronation of the forearm. The proximal, ...Introduction · Structure and Function · Muscles · Physiologic VariantsMissing: capitulum | Show results with:capitulum
  41. [41]
    Annular Ligament - Physiopedia
    It encircles 80% of the radial head and functions to maintain the relationship between the head of the radius and the humerus and ulna.Attachment · Clinical significance · Assessment · Treatment
  42. [42]
    The Elbow Joint - Structure - Movement - TeachMeAnatomy
    ### Summary of Elbow Joint Movements Involving the Capitulum, Flexion, Extension, Pronation, Supination
  43. [43]
    Anatomy of Selected Synovial Joints - Lumen Learning
    The articular capsule of the elbow is thin on its anterior and posterior ... The lateral side of the elbow is supported by the radial collateral ligament.
  44. [44]
    Proximal radioulnar joint: Anatomy, movements - Kenhub
    The superior surface of the radial head rotates against the capitulum of humerus · The ridge of the radial head glides against the groove between the capitulum ...
  45. [45]
    Range of Motion Normative Values - Physiopedia
    Range of Motion Normal Values[edit | edit source] ; Forearm, Pronation, 80 ; Forearm · Supination, 80 ; Elbow, Flexion, 140 ; Elbow · Extension, 0.
  46. [46]
    Arthrokinematics - Physiopedia
    Dorsal or posterior glide of the head of radius increases elbow extension. Volar or anterior glide of the head of the radius increases elbow flexion. Hip ...
  47. [47]
    Elbow and Forearm - Clinical Gate
    Mar 18, 2015 · The arthrokinematics at the humeroradial joint involve a spin of the fovea of the radial head against the rounded capitulum of the humerus.
  48. [48]
    Elbow Anatomy & Biomechanics - Shoulder & Elbow - Orthobullets
    Mar 24, 2024 · The radiohumeral articulation is a pivot joint. Radial head is covered by cartilage for approximately 240 degrees.Missing: humeroradial capitulum
  49. [49]
    Quantitative Assessment of Radiocapitellar Joint Stability - MDPI
    At 90 degrees of elbow flexion, approximately 75% of the elbow joint stability against varus stress is attributed to the joint articulation [5]. The radial head ...
  50. [50]
    The Osteology of the Reptiles/Chapter 5 - Wikisource
    Sep 2, 2024 · The two bones of the forearm or antibrachium are always complete in reptiles and movable upon each other, freely in most terrestrial reptiles, ...
  51. [51]
  52. [52]
    Musculoskeletal modeling of sprawling and parasagittal forelimbs ...
    Jan 21, 2022 · Median MMA values for supination (external rotation of the humerus) are greatest in the echidna but peak values are highest in the tegu (Figure ...
  53. [53]
    From Lizard to Snake; Behind the Evolution of an Extreme Body Plan
    Primitive snakes such as Boas and Pythons do possess vestigial hindlimbs at the position of the cloaca [46], however no indications have ever been ...<|control11|><|separator|>
  54. [54]
    An exceptionally preserved Sphenodon-like sphenodontian reveals ...
    Mar 3, 2022 · Their proximal ends have a shallow concavity for articulation with the humeral capitulum. The ulnae have a moderately developed and ossified ...
  55. [55]
    Three-dimensional skeletal kinematics of the shoulder girdle and ...
    The elbow is formed by the articulation of the ulna and radius with the distal humerus. ... Crocodilian galloping: an unique gait for reptiles. Copeia ...
  56. [56]
    Ossification Pattern in Forelimbs of the Siamese Crocodile ...
    Jan 30, 2018 · While they are on land, crocodylians have a wide variety of gaits, thanks to their specific semierect posture for better terrestrial mobility ( ...
  57. [57]
    [PDF] AVIAN ANATOMY:
    On the ventral surface of the distal ulna the trochlea is markedly deepened in some birds (e.g., Phoenicopterus) fonning the Sulcus intercondylaris (Fig.
  58. [58]
    Distal humeri of extant Australian eagles; upper, caudal view; lower,...
    A large accipitrine bird of prey is described from the Mid Miocene Camfield Beds, Bullock Creek, Northern Territory, Australia, based on a distal humeral ...
  59. [59]
    Osteology of the Late Triassic Bipedal Archosaur Poposaurus ...
    Dec 8, 2019 · The capitulum is larger and has a subcircular articular facet, whereas the tuberculum is smaller and has a craniocaudally oriented elliptical ...
  60. [60]
    [PDF] The Osteology of Compsognathus longipes W AGNER - Zobodat
    the classic or archtypical. “coelurosaurian“ theropod ...
  61. [61]
    Embryology and Developmental Anatomy of the Elbow | Clinical Gate
    Apr 17, 2015 · Chondrification of the humeral, radial and ulnar shaft is present by the 6th intrauterine week in embryos of 12 mm and by this stage there has ...Missing: capitulum center
  62. [62]
    Multiple roles of Hoxa11 and Hoxd11 in the formation of the ...
    Jan 15, 2004 · Chondrogenesis appears to initiate normally in Hoxa11/d11double mutants. The reduced zeugopod condensation size in double mutant embryos may ...
  63. [63]
    Hoxa11 and Hoxd11 Regulate Chondrocyte Differentiation ... - NIH
    Aug 20, 2012 · We demonstrate that Hoxa11 and Hoxd11 act upstream of Runx2 and Shox2, two key regulators of chondrocyte differentiation.
  64. [64]
    Congenital Abnormalities of the Elbow - Clinical Gate
    Apr 11, 2015 · Absence of the capitellum is probably an example of congenital aplasia, but hypoplasia of the capitellum may occur after dislocation from ...
  65. [65]
    Sonic Hedgehog Signaling in Limb Development - Frontiers
    Altered Shh signaling is implicated in clinical conditions with congenital limb defects and in the evolution of the morphological diversity of vertebrate limbs.Missing: capitellar hypoplasia
  66. [66]
    The early evolution of the tetrapod humerus - PubMed
    A tetrapod humerus from the Late Devonian of Pennsylvania has a novel mix of primitive and derived characters.
  67. [67]
    [PDF] The lissamphibian humerus and elbow joint, and the origins of ...
    Jun 23, 2009 · Temnospondyli is the only group of early tetrapods that shows a progressive evolution of lissamphibian traits in the humerus and elbow joint.
  68. [68]
    [PDF] A new eupelycosaur from the Permian of New Mexico, USA
    Gordodon kraineri is a new genus and species of edaphosaurid eupelycosaur known from an associated skull, lower jaw and incomplete postcranium found in the.
  69. [69]
    Digital restoration of the pectoral girdles of two Early Cretaceous ...
    Mar 31, 2022 · New digital restorations indicate that the structure and function of the shoulder joint were highly variable among Early Cretaceous birds, ...Missing: capitulum | Show results with:capitulum
  70. [70]
    Trabecular architecture in the forelimb epiphyses of extant ...
    Nov 29, 2017 · Indeed, only the BV/TV of the glenoid cavity, humeral head and capitulum, and radial trochlea are not correlated to size (the radial head stands ...
  71. [71]
    Analysis of the humeral data. (A) The relationship between the distal...
    This study examines the bone architecture of the forelimb bones (humerus and radioulna) and the hindlimb bones (femur, tibiafibula, tibiale, and fibulare) of ...
  72. [72]
    Coronal Shear Fractures of the Distal Humerus - NIH
    Fractures of the capitellum and trochlea are uncommon fractures of the elbow and can be challenging to treat due to their size, location, and articular nature.
  73. [73]
    Type 4 capitellum fractures: Diagnosis and treatment strategies - PMC
    Mechanism of injury is usually a fall on the out-stretched hand, the radius imparting a shearing force on the capitellum., Maximum force transmission through ...Case 2 · Figure 2 · Case 3
  74. [74]
    Panner disease | Radiology Reference Article - Radiopaedia.org
    Jan 16, 2023 · Panner disease is typically seen in children (5-10 years of age), although it is also seen in throwers due to repeated trauma. Panner disease ...
  75. [75]
    Panner's disease: literature review and treatment recommendations
    Panner's disease is a self-limiting disease and the majority of patients heal without clinical impairment. Conclusions. Based on the results of this review, ...Missing: resolves | Show results with:resolves
  76. [76]
    Osteochondritis dissecans of the capitellum in adolescents - PMC
    The prevalence of OCD of the humeral capitellum was 3.4% among more than 2000 adolescent baseball players[2]. Not all of these patients had symptoms[2].
  77. [77]
    Incidence of elbow involvement in rheumatoid arthritis. A 15 year ...
    Erosions were most often located on the capitellum and the lateral epicondyle of the humerus or the olecranon of the ulna. Severe destruction was most often ...Missing: capitulum | Show results with:capitulum
  78. [78]
    Chondroblastoma - StatPearls - NCBI Bookshelf - NIH
    Aug 14, 2023 · Chondroblastoma is a rare benign neoplasm, accounting for less than 1% of all primary bone tumors. Most chondroblastomas are diagnosed in the ...
  79. [79]
    "Primary" aggressive chondroblastoma of the humerus: a case report
    Chondroblastoma is a rare epiphyseal benign bone tumor, occurring usually in the second decade and constituting less than 1% of bone tumors.Missing: capitellum | Show results with:capitellum
  80. [80]
    Sail sign (elbow) | Radiology Reference Article - Radiopaedia.org
    Feb 11, 2025 · It indicates the presence of an elbow joint effusion. The anterior fat pad is usually concealed within the coronoid fossa or seen paralleling ...Missing: capitulum size cartilage >5mm
  81. [81]
    Osteochondritis dissecans of the capitellum: lesion size and pattern ...
    The median OCD surface area was 101 mm2 (range, 49-217 mm2). On the basis of OCD heat mapping, the posterolateral zone of the capitellum was most frequently ...
  82. [82]
    MRI Findings of Osteochondritis Dissecans of the Capitellum with ...
    Unstable osteochondritis dissecans lesions are surrounded by a rim of high signal intensity or a fluid-filled cyst on T2-weighted images. Stable osteochondritis ...
  83. [83]
    Radiographic evaluation of osteochondritis dissecans of the ... - NIH
    ICRS Stage I indicates a stable lesion with continuous, softened area covered by intact cartilage; ICRS Stage II indicates a stable lesion with partial ...
  84. [84]
    Radial Head Changes in Osteochondritis Dissecans of the Humeral ...
    Apr 18, 2018 · ... Nelson grade 4 capitellar ... Classification, treatment, and outcome of osteochondritis dissecans of the humeral capitellum.