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Dragonfly

Dragonflies are aerial predatory insects in the order Odonata, distinguished by their large compound eyes, elongated bodies, and two pairs of transparent, veined wings held outstretched at rest, setting them apart from the related damselflies which fold their wings together.^[1] Belonging to the suborder Anisoptera within Odonata, which encompasses approximately 3,120 species worldwide (as of 2025) alongside about 3,300 damselfly species in Zygoptera,^[2] dragonflies exhibit remarkable flight agility, capable of speeds up to 100 body lengths per second and precise aerial maneuvers for hunting.^[1] Their life cycle is hemimetabolous, featuring aquatic nymphs (naiads) that undergo 9 to 17 molts over months to years in freshwater habitats before emerging as winged adults that live for weeks to months, during which they mate, lay eggs, and continue predation.^[3] Physically, adult dragonflies possess a robust thorax supporting powerful flight muscles, six jointed legs adapted for perching and capturing prey mid-air, and a prehensile labium for grasping insects, with eyes containing up to 28,000 ommatidia for near-360-degree vision.^[1] Nymphs, in contrast, are bulky, aquatic predators equipped with internal tracheal gills and a specialized lower lip (labium) that shoots out to capture small fish, tadpoles, and invertebrates, highlighting their role as top consumers in lentic and lotic freshwater ecosystems.^[3] Evolutionarily ancient, Odonata fossils date back to the Carboniferous period around 325 million years ago, with ancestral forms like Protodonata exhibiting wingspans up to 75 cm, though modern dragonflies range from 2 to 13 cm in body length and are found on every continent except Antarctica.^[1] Behaviorally, male dragonflies are often territorial, patrolling and defending oviposition sites near water bodies with displays of speed and acrobatics, while mating involves the formation of a "wheel" where the male grasps the female behind the head with abdominal appendages.^[3] Ecologically, they serve as vital bioindicators of environmental health, with larvae controlling mosquito populations and adults preying on flying insects like flies and moths, thus facilitating nutrient transfer between aquatic and terrestrial food webs.^[4] Their cultural significance spans folklore and art across societies, often symbolizing agility and transformation, while recent research explores their flight mechanics for biomimetic applications in robotics and aerospace.^[4]

Overview

Etymology

The English common name "dragonfly" originated in the 1620s as a compound of "dragon" and "fly," with "dragon" deriving from Old English draca (from Latin draco, meaning "serpent" or "dragon") and "fly" from Old English flēoge (a winged insect). This nomenclature reflects the insect's elongated body, large eyes, and predatory flight, which evoked associations with mythical dragons in European folklore.^[5] In scientific classification, dragonflies belong to the order Odonata, a term coined by Danish entomologist Johan Christian Fabricius in 1793 from the Ancient Greek odous (Ionic form of odōn), meaning "tooth," in reference to the strong, toothed mandibles used for capturing prey. Key genera include Aeshna (encompassing many hawker species), derived from Greek aischma or aeschne, connoting "shame," "disgrace," or "ugly/misshapen" to describe the insect's robust form, and Libellula (skimmers), from Latin libella, a diminutive of libra ("balance" or "level"), alluding to the horizontal posture of the resting wings that resembles a carpenter's level.^[6]^[7]^[8] Across cultures, dragonfly nomenclature often highlights observed behaviors like rapid flight. In French, the standard term is libellule, sharing the Latin root with Libellula and emphasizing the balanced wing position, though regional folklore sometimes links the insect's speed to lightning, inspiring poetic associations with swift, electric movement. In Japanese, tombo (an ancient term possibly onomatopoeic for the sound of wings or descriptive of tumbling flight) symbolizes agility and victory, with dragonflies revered in samurai culture for their evasive maneuvers; ancient texts even named Japan Akitsushima ("dragonfly island") due to the abundance of these insects.^[7]^[9]

Taxonomy

Dragonflies belong to the class Insecta within the phylum Arthropoda, and they are classified in the order Odonata, which encompasses both dragonflies and damselflies.^[10] The order Odonata comprises approximately 6,400 extant species worldwide as of 2025, divided into three suborders: Anisozygoptera (a small group of relict species), Zygoptera (damselflies), and Anisoptera (true dragonflies).^[10]^[2] The suborder Anisoptera includes approximately 3,120 species, distinguished from Zygoptera primarily by morphological traits such as the position of the wings at rest—dragonflies hold their wings outstretched and perpendicular to the body, whereas damselflies fold their wings together over the abdomen.^[11] Additional distinguishing features in Anisoptera include a more robust body build, eyes that meet at the top of the head, and hind wings broader at the base than the forewings.^[12] The suborder Anisoptera is further classified into 11 families, with major groups including Aeshnidae (darners), Libellulidae (skimmers), and Gomphidae (clubtails).^[13] Aeshnidae, known for large, hawklike species often seen patrolling territories, contains over 500 species globally as of 2023.^[10]^[14] Libellulidae is the largest family, with over 1,000 species, characterized by perching behaviors and diverse color patterns, and it dominates in tropical and temperate regions near standing waters.^[10] Gomphidae, featuring club-tailed nymphs and adults with separated eyes, includes approximately 1,000 species and is particularly diverse in flowing waters across the Holarctic and Oriental realms.^[10] Other notable families include Corduliidae (emeralds) and Macromiidae (cruisers), which together contribute to the suborder's ecological variety in lotic and lentic habitats.^[13] Recent taxonomic revisions in Odonata have been driven by molecular phylogenetics, incorporating multi-gene datasets such as nuclear rRNA, mitochondrial genes, and transcriptomics to resolve family-level relationships; ongoing updates, such as the World Odonata List as of 2025, continue to refine species counts while maintaining the 11-family structure for Anisoptera.^[10]^[2] A comprehensive analysis of 510 Anisoptera representatives confirmed the monophyly of most families, including Aeshnoidea (Austropetaliidae + Aeshnidae) as sister to all other dragonflies, and Libelluloidea (encompassing Synthemistidae, Macromiidae, Corduliidae, and Libellulidae).^[13] However, debates persist regarding the monophyly and placement of smaller families like Chlorogomphidae, Neopetaliidae, and Petaluridae, with weak bootstrap support in some phylogenies suggesting potential paraphyly or alternative sister-group relationships.^[13] These revisions have led to taxonomic adjustments, such as the expansion of Synthemistidae to include genera previously in Corduliidae and the recognition of new subfamilies within Libellulidae, like Dythemistinae.^[13]

Evolutionary History

Fossil Record

The fossil record of Odonata and their stem-group relatives, the Odonatoptera, begins in the Late Carboniferous period, with the earliest known fossils dating to approximately 323 million years ago during the Pennsylvanian subperiod.^[15] These early forms were large predatory insects with winged adults, representing the initial diversification of flying odonatopterans in swampy, forested environments rich in oxygen.^[15] During the subsequent Permian period (298–251 million years ago), stem-odonates reached their peak in size, exemplified by the griffenfly Meganeuropsis permiana, which had a wingspan of up to 71 cm and likely preyed on smaller arthropods in aerial hawking strategies similar to modern dragonflies.^[16] This gigantism is attributed to high atmospheric oxygen levels, allowing for greater body sizes without respiratory limitations, though these forms went extinct by the end of the Permian.^[17] The transition to crown-group Odonata, encompassing modern dragonflies and damselflies, occurred by the Late Triassic, with clear evidence of anisopteran (dragonfly) morphology appearing in the Middle Jurassic approximately 168 million years ago.^[18] Key fossil sites, such as the Solnhofen limestone in southern Germany, have yielded exceptionally preserved Jurassic specimens that reveal detailed wing structures and body plans closely resembling those of extant species, including genera like Aeschnidium.^[19] Following the Cretaceous-Paleogene extinction event approximately 66 million years ago, which affected many insect lineages, Odonata underwent renewed diversification in the Cenozoic, leading to the radiation of modern families.^[20] A notable feature of the odonate fossil record is the morphological stasis in wing venation, where the intricate network of veins—critical for flight stability and identification—has remained largely unchanged from Carboniferous stem forms to present-day species over more than 300 million years, underscoring the evolutionary success of this design.^[18]

Phylogenetic Origins

Odonata, the order encompassing dragonflies and damselflies, occupies a basal position within the Paleoptera clade of winged insects (Pterygota), characterized by the retention of certain primitive wing articulation traits. Molecular clock analyses, calibrated with fossil constraints, estimate that the divergence of Odonata from other pterygote lineages occurred approximately 400 million years ago during the Devonian period, marking one of the earliest radiations of flying insects. This ancient origin aligns with the broader emergence of Hexapoda, where Odonata represents a foundational branch diverging before the diversification of Neoptera. Cladistic analyses based on morphological characters indicate that the suborder Anisoptera (true dragonflies) evolved from ancestors shared with Zygoptera (damselflies), with the common odonate ancestor likely resembling modern Zygoptera in overall body plan. Key synapomorphies defining Anisoptera include the complete fusion of the compound eyes into a single seamless structure, enhancing visual acuity for aerial predation, and modifications to leg morphology, such as elongated and spinose tibiae adapted for grasping prey in flight.^[21] These traits distinguish Anisoptera from the more plesiomorphic, separated eyes and less specialized legs of Zygoptera, supporting a phylogeny where Epiprocta (including Anisoptera) is sister to Zygoptera within Odonata.^[22] Recent mitogenomic studies since 2020 have refined family-level phylogenies across Odonata, utilizing complete mitochondrial genomes to resolve deep divergences and intrasubordinal relationships with high resolution. For instance, analyses of over 100 mitogenomes have confirmed the monophyly of major families like Libellulidae and Aeshnidae, while revealing discrepancies in divergence estimates between mitochondrial and nuclear data, with crown-group radiations often placed in the Jurassic to Cretaceous.^[23] These investigations also highlight instances of convergent evolution, such as independent refinements in wing venation patterns across Anisoptera families that optimize high-speed flight, and parallel adaptations in predatory behaviors, including aerial interception strategies that mimic those in distantly related insect predators.^[24] Such convergences underscore the selective pressures of aerial hunting lifestyles in shaping odonate diversity.

Distribution and Diversity

Global Distribution

Dragonflies (Odonata) exhibit a cosmopolitan distribution, occurring on every continent except Antarctica and showing near-absence in polar regions of the Arctic, where persistently low temperatures constrain larval development and adult flight activity, as minimum temperatures for growth typically exceed 8–12°C.^[25]^[26] Their presence is predominantly in tropical and temperate zones, with maximum densities concentrated in Southeast Asia and Africa, where tropical rainforests and diverse freshwater habitats support exceptional richness in flowing waters.^[27]^[28] Continental patterns reveal regional hotspots influenced by climate and habitat availability; in North America, species richness peaks in the southeast United States, driven by warm, humid conditions and varied wetlands that foster high endemism.^[29] Similarly, island endemism is pronounced in Madagascar, where over 190 species occur, with nearly 95% of damselflies (Zygoptera) unique to the island due to its isolation and diverse aquatic ecosystems.^[30] Climate change is altering these ranges, particularly in the Northern Hemisphere, with observed poleward shifts in Europe since the late 20th century; for instance, southern lentic dragonfly species have expanded northward at an average rate of 115 km per decade, reflecting improved thermal suitability at higher latitudes.^[31] These shifts underscore temperature as a primary limiter of dragonfly distributions, enabling colonization of previously unsuitable cooler areas while potentially stressing southern populations through habitat loss.^[32]

Species Diversity

The order Odonata, encompassing both dragonflies and damselflies, includes approximately 6,442 recognized extant species worldwide as of 2025.^[2] Dragonflies, belonging to the suborder Anisoptera, comprise about 3,120 of these species, distributed across 11 families and 348 genera.^[33] Ongoing taxonomic research continues to uncover new species, with estimates suggesting 1,000 to 1,500 additional Odonata species remain to be described, leading to dozens of annual additions through field surveys and genetic analyses.^[34] Biodiversity hotspots for dragonflies are concentrated in tropical regions, particularly the Indo-Malayan realm, which harbors a significant portion of global Odonata diversity—nearly 30% of all species, exceeding 1,900 in total, with dragonflies forming a substantial share.^[35] This region supports high endemism due to diverse wetland habitats, though many species face habitat fragmentation. In contrast, isolated archipelagos like Hawaii host only two endemic dragonfly species—Anax strenuus and Nesogonia blackburni—both vulnerable to extinction from invasive predators, altered water flows, and development, rendering their populations critically imperiled.^[36]^[37] Globally, conservation assessments indicate that about 16% of the 6,016 evaluated Odonata species are threatened with extinction, based on the 2022 IUCN Red List update, highlighting risks from wetland loss and climate change, with tropical endemics disproportionately affected.^[38]

Morphology

Anatomy

Dragonflies exhibit a segmented body plan typical of insects, consisting of three primary tagmata: the head, thorax, and abdomen. The head is dominated by a pair of large compound eyes, each comprising up to 30,000 ommatidia that provide panoramic vision and high acuity for detecting motion.^[39] These eyes nearly cover the head surface, with three simple ocelli positioned dorsally for additional light detection. The head also features a robust mandible-equipped mouthpart adapted for biting and consuming prey, along with short antennae for mechanosensory input.^[40] The thorax is specialized for locomotion, comprising three segments—a small prothorax and the fused mesothorax and metathorax (pterothorax)—that house powerful direct flight muscles. Unlike the indirect muscles of many insects, these direct muscles attach to the wing bases via tendons, enabling precise control of wing movement and high maneuverability.^[41] The legs, arising from the thorax, are elongated and equipped with spines; the forelegs are particularly modified to form a "basket" structure using the femur, tibia, and tarsus for capturing airborne prey.^[42] Emerging from the thorax are four wings, each independently controllable and supported by a complex network of veins that form a reticulate pattern for structural reinforcement. Key features include the nodus, a cross-vein hinge midway along the wing that enhances flexibility and stability during flapping, and the pterostigma, a thickened, pigmented cell at the leading edge near the tip that shifts the wing's center of mass to dampen oscillations and prevent flutter.^[43] The abdomen is elongated and cylindrical, typically comprising 10 segments that elongate the body for streamlined flight and house reproductive organs. In males, the posterior segments include claspers for grasping females during mating, while in females, they form an ovipositor for egg-laying.^[40] Internally, dragonflies possess an open circulatory system where hemolymph is pumped by a dorsal vessel through the hemocoel to bathe tissues directly, without enclosed vessels.^[44] Excretion occurs via Malpighian tubules that filter hemolymph to remove nitrogenous wastes, primarily as uric acid, before reabsorption in the hindgut.^[45] Respiration relies on a tracheal system, with oxygen delivered through a network of tubes branching from 10 pairs of spiracles—two thoracic and eight abdominal—that open to allow passive diffusion to cells.^[46]

Coloration

Dragonfly coloration results from the interplay of pigmentary and structural mechanisms, producing a diverse array of hues that serve ecological functions. Pigmentary colors arise primarily from melanins, which generate black and brown tones by absorbing light in the exocuticle, and pterins, which contribute yellows, reds, and whites through selective absorption in the epidermis.^[47] Structural coloration, often iridescent, stems from nanoscale features such as multilayered cuticles and wing scales that cause thin-film interference, yielding metallic blues and greens visible at specific angles.^[47] These mechanisms combine in the integument, where the transparent cuticle overlays pigmented epidermal layers to enhance color intensity.^[47] Although less common than in damselflies, female color polymorphism occurs in some dragonfly species, such as certain Libellulidae.^[47] Sexual dimorphism is prevalent, with males typically exhibiting brighter, pruinose blues or reds—such as in Crocothemis species where males are vivid red compared to yellow females—for territorial display and mate attraction.^[47] These colors fulfill adaptive roles in survival and reproduction. Nymphs employ cryptic melanin-based patterns in dull browns and greens to camouflage against aquatic vegetation and substrates, reducing predation risk as seen in Anax junius.^[48] In adults, conspicuous patterns act as warning signals to deter predators, signaling unpalatability through chemical defenses in some species.^[47] Ultraviolet reflectance, often iridescent from wing structures in genera like Rhyothemis, enhances mate attraction by providing visual cues during courtship, with pterin pigments amplifying UV signals for species recognition.^[49]

Sensory and Physiological Adaptations

Eyesight

Dragonflies possess large compound eyes composed of thousands of ommatidia, providing a nearly panoramic field of view spanning approximately 360 degrees, which allows them to detect movement from virtually any direction without turning their heads.^[50] This visual coverage is facilitated by the eyes' positioning on the sides of the head, with each eye containing up to 30,000 individual facets that capture light independently.^[51] Within these compound eyes, specialized regions known as acute zones or fovea-like areas offer higher resolution for forward and dorsal vision, where ommatidial packing is denser to enhance acuity during prey pursuit.^[52] The visual system supports trichromatic color vision augmented by ultraviolet (UV) sensitivity, enabled by a diverse array of opsin genes—up to 33 in some species—that encode photoreceptors tuned to UV, short-wavelength (blue), and long-wavelength (green to red) spectra.^[53] This UV capability, stemming from a conserved UV-sensitive opsin expressed across eye regions, aids in detecting patterns invisible to humans, such as UV reflections on prey or mates, and contributes to habitat orientation.^[53] Electrophysiological studies confirm peak sensitivities around 350 nm (UV), 440 nm (blue), and 530 nm (green), allowing dragonflies to discriminate colors in complex environments.^[54] Neural processing in the dragonfly brain involves wide-field motion-sensitive neurons, such as the centrifugal small target motion detector 1 (CSTMD1), which selectively respond to small moving targets against cluttered backgrounds, facilitating target-selective descent onto prey.^[55] These neurons exhibit spatiotemporal tuning that prioritizes targets moving at velocities matching the dragonfly's flight speed (around 90°/s angular velocity), suppressing responses to background motion through competitive mechanisms.^[55] Response latencies for CSTMD1 onset are approximately 40-50 ms, enabling rapid behavioral reactions with takeoff times around 75 ms during interception. This swift processing underpins the dragonfly's high prey capture success rate of over 95%.^[56] Dragonflies also utilize polarized light sensitivity for navigation, particularly during long-distance migration, where atmospheric polarization patterns provide compass cues independent of the sun's position. A 2025 study highlighted polarization vision in odonates like dragonflies for water detection and likely navigation, though direct evidence for its role in long-distance migration and integration with wind compensation remains a research gap.^[39] This adaptation, detected via specialized photoreceptors in the compound eyes, enhances orientation accuracy in varying light conditions.^[39]

Temperature Regulation

Dragonflies, as ectothermic insects, rely primarily on behavioral strategies to regulate body temperature, given their limited physiological capacity for endothermy. To avoid overheating, many species, particularly perchers, adopt the obelisk posture, in which the abdomen is raised vertically toward the sun, thereby minimizing the body surface area exposed to direct solar radiation and reducing heat gain. This behavior is especially prevalent in hot environments, where it helps maintain thoracic temperatures below critical thresholds. Conversely, for warming up in cooler conditions, dragonflies employ wing-whirring, a rapid vibration of the wings that generates metabolic heat through muscular activity, elevating thoracic temperatures above ambient levels to enable flight. Additionally, convective cooling via wing movements or fanning can dissipate excess heat when thoracic temperatures approach 40°C, preventing thermal stress during prolonged activity.^[57] Behavioral thermoregulation in dragonflies varies with latitude and habitat, reflecting adaptations to local climates. In temperate regions, species often bask with spread wings to absorb solar heat, achieving optimal thoracic temperatures of 30–40°C for activity, with physiological limits allowing flight from approximately 10°C to 45°C ambient temperatures. Tropical species, such as those in the genus Micrathyria, exhibit more reliance on shade-seeking to mitigate intense midday heat, as their higher minimum flight temperatures (often >25°C) restrict early and late-day activity in smaller individuals; larger tropical perchers, however, extend activity periods through effective postural adjustments and microhabitat selection, maintaining body temperatures within 25–42°C despite ambient fluctuations up to 45°C. These strategies ensure activity within the broad physiological envelope of 10–45°C, beyond which torpor or reduced locomotion occurs.^[57] Climate change exacerbates heat stress for dragonflies, leading to shortened activity periods and physiological strain in recent decades. Rising temperatures in disturbed habitats, such as deforested tropical dry forests, expose individuals to maximum air temperatures exceeding 49°C, prompting increased shade-seeking and reduced foraging or mating time to conserve energy reserves like lipids and proteins. Studies show that in disturbed habitats with high temperatures exceeding 49°C, dragonflies exhibit increased thermal stress, leading to reduced energy reserves.^[58]

Behavior and Ecology

Flight

Dragonflies utilize asynchronous flight muscles, which operate via stretch-activation rather than synchronous neural impulses, enabling wingbeat frequencies of 20 to 50 beats per second for sustained powered flight.^[59] These muscles allow rapid oscillations decoupled from direct motor neuron firing, providing the high power output necessary for the insect's agile aerial performance.^[60] The direct musculature attached to each of the four wing bases grants dragonflies independent control over individual wings, facilitating extraordinary maneuverability such as precise hovering, abrupt directional changes, and even backward flight.^[61] This capability arises from the ability to vary flapping amplitude, phase, and stroke plane orientation across wings, enabling complex aerodynamic interactions for stability and propulsion.^[62] Such control supports accelerations up to 20 m/s² during maneuvers, far exceeding those of many other insects.^[63] Aerodynamically, dragonfly wings generate lift through the stabilization of leading-edge vortices (LEVs) on their upper surfaces during both downstroke and upstroke, creating low-pressure regions that enhance force production beyond traditional steady-state models.^[64] These vortices, promoted by the wings' corrugated leading edges and flexible structures, contribute to a lift-to-drag ratio improvement of up to 4% compared to smooth equivalents.^[65] In bursts, dragonflies achieve maximum speeds of 30-60 km/h, with the Australian dragonfly (Austrophlebia costalis) recorded at up to 58 km/h, showcasing their burst capabilities for pursuit.^[66] Recent biomimicry research has leveraged these principles for micro-drone development; for instance, the 2024 HiFly-Dragon project incorporates dragonfly-inspired tandem flapping wings with independent control to achieve efficient hovering and forward flight in a 33 g vehicle flapping at 28 Hz.^[67] Complementary studies on insect flapping mechanisms, including the clap-and-fling motion where wings briefly clap together before separating to amplify LEV formation, have informed designs for enhanced lift in such small-scale drones, drawing from Odonata aerodynamics.^[68]

Feeding

Dragonflies are exclusively carnivorous throughout their life cycle, with adults primarily engaging in aerial interception to capture flying insects, which form the bulk of their diet. These agile predators use their exceptional flight capabilities to pursue and seize prey mid-air, often targeting smaller insects such as mosquitoes, flies, and midges. The capture process involves rapid maneuvers and precise targeting, enabling dragonflies to achieve high success rates in interception.^[69]^[70] In contrast, dragonfly nymphs are ambush predators in aquatic environments, remaining motionless until suitable prey ventures within striking distance. They deploy a specialized, extensible labium—a modified lower lip forming a basket-like mask armed with movable hooks—to rapidly extend and grasp aquatic invertebrates, tadpoles, or small fish. This mechanism allows for a powerful bite and secure hold, facilitating efficient predation without prolonged pursuit.^[71]^[72] Adult dragonflies exhibit two primary foraging modes: perchers, which wait on vegetation and make short sallying flights to intercept passing prey, and fliers (or hawkers), which remain airborne continuously to patrol and pursue targets. These strategies reflect adaptations to energy efficiency, with perchers conserving resources for brief bursts of activity and fliers investing in sustained flight for broader coverage. Daily energy budgets are substantial, as adults may consume up to 20% of their body weight in prey to fuel their high metabolic demands.^[73]^[74]^[75] As keystone predators, dragonflies play a critical role in regulating insect populations, particularly by preying on mosquito larvae and adults, thereby reducing vector-borne disease transmission. Prey selection is often size-based, with larger dragonfly species capable of targeting bigger prey items that smaller conspecifics avoid, optimizing capture success and nutritional intake. This selective predation helps maintain ecological balance in both aquatic and terrestrial habitats.^[76]^[77]^[78]^[79]

Habitat and Migration

Dragonfly larvae, or nymphs, primarily inhabit freshwater environments characterized by still or slow-moving waters, including ponds, lakes, marshes, and the margins of rivers and streams. These aquatic habitats provide the oxygen-rich conditions and prey necessary for their predatory lifestyle during the often multi-year larval stage. Species diversity in larval assemblages is highest in wetlands with varied submerged and emergent vegetation, which offers shelter from predators and supports periphyton growth for food chains.^[80]^[81]^[82] Adult dragonflies exhibit a strong preference for vegetated wetlands and riparian zones, where dense emergent plants like sedges, cattails, and bulrushes provide perching sites, oviposition substrates, and hunting grounds. These areas facilitate thermoregulation and mate location while minimizing exposure to terrestrial predators. In landscapes with fragmented wetlands, adults may travel considerable distances to locate suitable breeding sites, underscoring their dependence on connected aquatic-terrestrial interfaces.^[83]^[84]^[85] Many dragonfly species undertake migrations, with the common green darner (Anax junius) serving as a prominent example of a long-distance traveler capable of covering up to 700 km in seasonal movements across North America. These migrations often involve crossing large bodies of water, including coastal and oceanic stretches, facilitated by favorable tailwinds that can boost flight speeds beyond 30 km/h. Migration is typically cued by environmental factors such as wind direction, temperature increases, and shortening photoperiods in late summer, prompting southward journeys from breeding grounds in northern latitudes to overwintering sites in the tropics.^[48]^[86]^[87] In Europe, vagrant patterns are exemplified by the vagrant emperor (Anax ephippiger), which exhibits irregular, wind-assisted dispersals from African breeding populations into northern regions, sometimes forming swarms that appear sporadically in countries like the UK and Germany. These movements are not strictly seasonal but respond to episodic weather events, leading to temporary colonizations of temporary ponds before populations wane without sustained reproduction. Stable isotope analysis has revealed that such vagrants originate from distant Saharan oases, highlighting the role of atmospheric currents in continental-scale connectivity.^[88]^[89] Habitat fragmentation, particularly in urbanizing landscapes, severely impacts dragonfly populations by isolating breeding sites and reducing connectivity between aquatic and terrestrial habitats. Loss of wetlands since 2000 has contributed to declines in many species, with global assessments indicating that habitat degradation threatens 16% of all dragonfly and damselfly species. In urban areas, where impervious surfaces and drainage alter hydrology, populations of wetland-dependent odonates have decreased in fragmented regions, as breeding success drops due to diminished water quality and vegetation cover. Conservation efforts emphasize restoring wetland corridors to mitigate these effects and support migratory pathways.^[90]^[91]^[92]

Reproduction and Life Cycle

Mating and Reproduction

Dragonfly mating begins with male territorial behaviors, where individuals patrol specific areas near water bodies to attract receptive females and repel rival males. These patrols often involve rapid flights along linear routes, such as pond edges or river margins, allowing males to intercept and court passing females while monitoring for intruders.^[93] In many species, such as those in the Libellulidae family, territorial males achieve higher mating success compared to non-territorial "satellite" males, as they gain priority access to females arriving at reproductive sites.^[94] Once a female is encountered, the male initiates contact by grasping her behind the head or prothorax with abdominal claspers, forming a tandem linkage that may involve synchronized flight to assess receptivity or deter rivals. If the female is willing, the pair transitions to the "wheel" position, where the female curls her abdomen to receive sperm from the male's secondary genitalia located on the underside of his abdomen. This indirect insemination allows for sperm competition, as the male's aedeagus (penis) is equipped with spines and lobes that physically remove or displace rival sperm from the female's spermathecae, ensuring higher paternity for the last mating partner.^[95] Such mechanisms are widespread in Anisoptera, promoting last-male sperm precedence rates often exceeding 80% in species like Leucorrhinia dubia.^[94] Following insemination, the male may remain in tandem with the female during oviposition to guard against further matings, a behavior known as mate-guarding. Females then deposit eggs either endophytically, by slicing into plant tissues with their ovipositor (common in families like Aeshnidae), or exophytically, by scattering them on the water surface or vegetation (typical in Libellulidae). Site selection by females is influenced by water quality, with species favoring clear, well-oxygenated habitats that support larval survival, as indicated by correlations between adult assemblages and in-stream conditions like pH and nutrient levels.^[96]^[97] Sexual conflict arises during these processes, as females often resist unwanted advances to control mating frequency and avoid energy costs or injury. In species exhibiting strong male harassment, females employ behaviors such as feigned death or erratic flight to evade claspers, while persistent male grasping can lead to physical damage, including wing tears or abdominal wounds. For instance, in the damselfly Calopteryx (closely related to dragonflies within Odonata), multiple copulations increase the incidence of copulatory wounding on female genitalia, highlighting the antagonistic coevolution between sexes.^[98]^[99]

Developmental Stages

Dragonflies undergo incomplete metamorphosis, characterized by three primary life stages: egg, nymph, and adult, without a distinct pupal phase.^[100] The egg stage typically lasts 1 to 4 weeks, depending on species and environmental conditions, during which the embryo develops within a protective shell laid in or near water.^[101] Upon hatching, the pronymph—a short-lived initial form—quickly molts into the first nymphal instar.^[48] The nymphal stage, which is aquatic and represents the longest phase of the life cycle, can endure for up to 2 years in many temperate species, though durations vary from several months in tropical forms to over 5 years in some larger or colder-climate species.^[102] Nymphs undergo 9 to 17 molts, passing through 10 to 18 instars, progressively increasing in size and refining predatory features like extendable labium for capturing prey.^[103] In temperate regions, many species exhibit semivoltine cycles, requiring two years to complete nymphal development due to overwintering diapause. Adaptations for underwater life include internal gills located in a rectal chamber, which facilitate oxygen extraction from water drawn in and expelled through muscular contractions; this mechanism also enables jet propulsion for rapid escape or pursuit, achieved by forcefully pumping water from the anus.^[104] Emergence marks the transition to adulthood, where the final instar nymph crawls out of the water—often at dawn—to a vertical perch, splits its exoskeleton, and expands its wings in a process lasting 1 to 2 hours.^[102] The resulting teneral adult is pale, soft-bodied, and highly vulnerable to predation and dehydration, remaining flightless and grounded until its exoskeleton hardens and colors develop over several hours to days.^[105] Developmental rates across stages are strongly temperature-dependent, with warmer water accelerating growth and shortening overall duration; for instance, a 5°C increase can advance adult emergence by about 30 days compared to ambient conditions, effectively speeding larval development by 20-30% in controlled studies.^[106] This thermal sensitivity influences voltinism, allowing some populations to complete cycles more rapidly in heated environments while risking higher mortality from metabolic stress.^[107]

Population Dynamics

Dragonfly populations frequently display male-biased sex ratios, often around 1.5:1 in many species, often arising from differential mortality, where the larger sex (frequently females) experiences higher rates during the larval stage, combined with protandry (males emerging earlier than females).^[108]^[109] This bias is further exacerbated by protandry, a common pattern where males emerge earlier than females, allowing males to establish territories but potentially increasing their exposure to risks before females arrive at breeding sites.^[110] Although sex ratios at hatching are typically 1:1, these demographic shifts contribute to uneven adult distributions, influencing mating opportunities and overall population structure.^[111] Density-dependent factors play a key role in regulating dragonfly population sizes, particularly through cannibalism among nymphs, which intensifies at higher densities and can reduce cohort survival by up to 50% in asynchronous hatching scenarios.^[112] Cannibalism rates escalate with size disparities between individuals—reaching 53% for one-instar differences and 100% for two-instar differences—primarily affecting smaller nymphs and thereby stabilizing population levels by limiting overcrowding.^[112] Carrying capacity in larval habitats is closely tied to prey availability, as abundant food supports larger body sizes and potentially mitigates some density effects, though survival remains predominantly governed by intraspecific predation rather than direct resource competition.^[112]^[113] Odonata, including dragonflies, are widely recognized as bioindicators of aquatic ecosystem health due to their sensitivity to habitat alterations, pollution, and climate shifts, enabling effective monitoring of environmental changes.^[114] A 2009 global assessment found approximately 10% of species threatened with extinction; a 2021 IUCN update raised this to 16%. In Europe, a 2024 report documented a 29% population decline in recent decades, attributed to habitat loss and degradation, with more severe reductions observed in vulnerable species through coordinated trend analyses and Red List evaluations. Recent studies highlight climate change as a growing threat, altering emergence timing and habitat suitability, exacerbating declines (as of 2024).^[115]^[114]^[91] These monitoring efforts underscore the need for ongoing surveillance to track demographic patterns like sex ratios and cohort reductions in response to anthropogenic pressures.

Biotic Interactions

Predators and Defenses

Dragonfly nymphs are vulnerable to predation by fish, frogs, and other aquatic organisms in their habitats. Fish, in particular, consume large numbers of nymphs, with studies showing that larvae exposed to predatory fish exhibit survival rates 2.5 to 4.3 times lower than those in predator-free environments. Frogs also prey on nymphs, especially during vulnerable stages near water surfaces. Adult dragonflies face threats primarily from birds and spiders; birds such as hawks (e.g., Mississippi Kites and Swallow-tailed Kites) and kingfishers make dragonflies a significant portion of their diet, while spiders ambush perched individuals. Lizards occasionally prey on adults as well. Predation contributes to high mortality rates among adults, with one study documenting up to 68% mortality from bird predation alone during emergence periods. The introduction of invasive fish species to ponds has exacerbated declines in dragonfly nymph populations by intensifying predation pressure. In fish-invaded ponds, odonate community composition shifts significantly, with lower nymph abundance and selection for traits like increased burst speeds in survivors to evade detection. For instance, goldfish introductions lead to shifts in morphological traits, such as larger body sizes and elongated spines in some surviving nymph species, along with reduced overall species richness compared to fishless ponds.^[116] Dragonflies employ several anti-predator defenses to mitigate these threats. Thanatosis, or feigning death by becoming motionless, serves as a passive defense, particularly in nymphs and some adults when disturbed, allowing them to avoid further attention until the threat passes. Evasive flight maneuvers, including rapid takeoffs at 45-degree angles and sinuous paths, enable quick escapes from aerial predators, leveraging their agile flight capabilities.

Parasites and Pathogens

Dragonflies are susceptible to a range of parasites and pathogens that can impair their physiology, behavior, and survival. Ectoparasites, such as larval water mites (Hydracarina, particularly Arrenurus spp.), commonly attach to the bodies of dragonfly nymphs during their aquatic phase, feeding on host fluids and potentially reducing host mobility and overall fitness.^[117] These mites alter host activity levels and survivorship, with attachments often occurring on the legs and abdomen, which may hinder swimming efficiency and escape responses in nymphs. Prevalence of water mite parasitism varies by species and habitat but can reach significant levels in lentic waters, contributing to decreased reproductive success in emerging adults.^[118] Endoparasites, including gregarine protozoans (Apicomplexa: Eugregarinorida), infect the gut of both larval and adult dragonflies, where they attach to the intestinal lining and absorb nutrients.^[119] These parasites disrupt muscle function, leading to reduced flight muscle power output—approximately 21% lower in infected individuals (117 W kg⁻¹ versus 148 W kg⁻¹ in uninfected ones)—due to altered protein degradation and lipid metabolism in the flight muscles.^[120] Gregarine prevalence can reach 41% across dragonfly species, with intensities aggregating up to a median of 5 parasites per host, and higher rates (18–52%) in common species like those in Central Texas reservoirs.^[119] Factors such as host density, habitat type, and seasonality influence infection levels, with prevalence increasing over the flight season.^[119] Fungal pathogens, notably Ophiocordyceps odonatae (Ophiocordycipitaceae), infect adult dragonflies by invading body tissues with hyphae, ultimately killing the host and producing fruiting bodies for spore dispersal.^[121] This entomopathogenic fungus has been documented in tropical regions, such as Singapore and Taiwan, where it emerges post-mortem from the host's abdomen.^[121] Although behavioral manipulation akin to that in ant hosts is not fully confirmed, some observations suggest potential influences on host positioning for optimal spore release, similar to other Ophiocordyceps species.^[122] Bacterial pathogens are less frequently reported but can occur as opportunistic infections in the gut microbiome, potentially exacerbated by environmental stressors; dominant phyla like Proteobacteria include genera with pathogenic potential in compromised individuals.^[123] Viral pathogens, including diverse circular single-stranded DNA viruses (CRESS-DNA) such as cycloviruses, have been identified in dragonfly populations worldwide, often through metagenomic surveys of wild-caught specimens.^[124] These viruses may circulate asymptomatically but could contribute to mortality in dense aggregations, as seen in larval samples from European ponds.^[125] Ecologically, parasite loads impose significant burdens on dragonfly populations, particularly migrants, by compromising flight performance and energy allocation, which can lower migration success in species like Pantala flavescens.^[120] Infection rates approach 50% in some temperate and tropical species, with higher diversity and prevalence in warmer, humid habitats that facilitate transmission.^[119]^[126] These impacts may regulate population dynamics, as heavily parasitized individuals exhibit reduced dispersal and survival during long-distance movements.^[127]

Dragonflies and Humans

Conservation

Dragonfly populations worldwide face significant threats, primarily from habitat loss due to agricultural expansion and wetland drainage, which has resulted in approximately 50% of global wetlands being lost since 1900.^[128] This drainage, often for farmland conversion, directly impacts breeding sites such as ponds, streams, and marshes essential for larval development.^[91] Pollution from pesticides, nutrient runoff, and water contamination further exacerbates declines by degrading water quality and introducing toxins that affect both aquatic larvae and adult dragonflies.^[114] Climate change compounds these pressures by altering precipitation patterns, raising temperatures, and shifting seasonal timings, which disrupt breeding habitats and migration routes for many species.^[129] A 2021 IUCN assessment found that 16% of the world's approximately 6,016 dragonfly and damselfly species—totaling 962 taxa—are threatened with extinction, a figure that highlights the urgency of addressing these interconnected threats.^[91]^[129] However, only about 25% of Odonata species have been assessed for the IUCN Red List as of 2025, with 29.4% of assessed species classified as Data Deficient, emphasizing the need for expanded monitoring. In Europe, for instance, habitat destruction and pollution have led to critical declines in odonate populations, with 29 species assessed as threatened (Vulnerable, Endangered, or Critically Endangered), including 11 Endangered or Critically Endangered, in the 2024 European Red List assessment.^[130] Conservation efforts focus on protecting and restoring wetland habitats through international frameworks like the Ramsar Convention, which designates Wetlands of International Importance to safeguard critical areas for dragonfly reproduction and survival.^[131] These protected sites help mitigate habitat loss by promoting sustainable water management and limiting agricultural encroachment, with many serving as refuges for vulnerable species.^[132] Reintroduction programs for endangered odonates, such as those targeting habitat specialists in fragmented European landscapes, aim to bolster populations by relocating individuals to restored sites, though success depends on addressing ongoing pollution and climate stressors.^[130] Citizen science initiatives play a vital role in monitoring dragonfly populations and informing conservation strategies, with apps and programs enabling volunteers to record sightings and collect data on species distribution and health.^[133] Projects like the Dragonfly Pond Watch and the Dragonfly Mercury Project engage participants in tracking migratory patterns and assessing pollution impacts through larval sampling, contributing to broader IUCN assessments and local restoration efforts.^[134] These tools enhance data collection in understudied regions, supporting targeted interventions to prevent further declines.^[135]

Cultural Representations

In Japanese folklore, dragonflies are revered as symbols of courage, happiness, and victory, often incorporated into samurai helmets and armor due to their forward-flying nature, which represents focused determination and vigilance.^[136] Known as kachimushi or "victory insect," they embody prosperity and good fortune, appearing in motifs on weapons and crests to invoke success in battle.^[137] Among Native American cultures, such as the Navajo, dragonflies symbolize water and renewal, frequently depicted in sacred sandpaintings to represent this vital element.^[138] In Zuni tradition, they serve as harbingers of rain and messengers from the spirit world, arriving alongside storms to signal life-giving moisture and good health.^[139] Dragonflies have inspired literary and poetic works across eras, capturing their ephemeral grace and predatory essence. In Matsuo Bashō's haiku, the insect's futile attempt to perch on a grass blade evokes themes of transience and the beauty of imperfection in nature.^[140] Similarly, Ted Hughes' poem "To Paint a Water Lily" portrays the dragonfly as a swift, meat-eating predator that "bullets by" or hovers to strike, highlighting the raw, violent harmony beneath serene landscapes.^[141] Artistically, dragonflies appear in ancient Egyptian motifs, such as amulets from the Middle Kingdom (ca. 1981–1640 B.C.) and tomb paintings of river scenes, where they symbolize the vitality of wetlands and seasonal renewal.^[142] In contemporary culture, dragonfly tattoos often represent personal transformation, drawing from the insect's dramatic life cycle—from aquatic nymph to airborne adult—to signify growth, adaptability, and overcoming adversity.^[143]

Technological Inspirations

Dragonfly morphology and locomotion have significantly influenced advancements in unmanned aerial vehicle (UAV) design, particularly through biomimicry of their agile flight capabilities. Engineers have drawn from the dragonfly's four-winged structure and rapid wingbeats, which enable precise maneuvers and hovering, to develop flapping-wing micro-drones for surveillance and reconnaissance. A prominent example is the Skeeter micro-drone developed by Animal Dynamics, which replicates dragonfly aerodynamics with independently flapping wings, achieving speeds up to 45 km/h and exceptional maneuverability in complex environments like urban settings.^[144] This design enhances stability and efficiency over traditional rotary-wing UAVs, as demonstrated in military-funded prototypes that prioritize low detectability and high agility.^[145] The dragonfly's compound eyes, comprising thousands of ommatidia for panoramic vision and motion detection, serve as a model for innovative imaging systems in robotics and autonomous vehicles. These eyes provide nearly 360-degree coverage with minimal blind spots, inspiring artificial compound eye cameras that eliminate the need for bulky lenses and fisheye distortions. In 2024, researchers at the Hong Kong University of Science and Technology created a curved array of 302 microlenses mimicking insect compound eyes, offering a 160-degree field of view with uniform resolution for enhanced obstacle detection in robots and self-driving cars.^[146] Such systems improve real-time environmental awareness, reducing computational demands compared to multi-camera setups in conventional autonomous navigation.^[147] Dragonfly nymphs' underwater jet propulsion, achieved by expelling water through rectal contractions for rapid bursts and directional control, has inspired soft robotic designs for aquatic exploration. This mechanism allows nymphs to achieve vectoring by asymmetrically opening their anal valves, enabling precise steering without rigid components. Advancements in soft robotics have incorporated compliant materials and fluidic actuators to replicate nymph-like jetting, resulting in lightweight underwater bots capable of efficient propulsion in confined or murky environments. These bioinspired systems offer advantages in stealth and adaptability for applications like ocean monitoring, where traditional propellers are inefficient.^[148]

References

[1]
Odonata: Dragonflies and Damselflies
Odonata, including dragonflies and damselflies, are ancient insects with large eyes, two pairs of wings, and a long abdomen. Dragonflies have stout bodies, ...
[2]
Odonata: Who They Are and What They Have Done for Us Lately
Feb 28, 2019 · Odonata (dragonflies and damselflies) are well-known but often poorly understood insects. Their phylogeny and classification have proved difficult to ...
[3]
Dragonflies and Damselflies (Insecta: Odonata) - University of Florida
Dragonflies are unmatched as fliers and have a very agile, deliberate flight. Males are often territorial, defending oviposition sites from other males. Dorsal ...
[4]
Dragonfly - Etymology, Origin & Meaning
[winged insect] Middle English flie (2), from Old English fleoge, fleogan "a fly, winged insect," from Proto-Germanic *fleugon "the flying (insect)" (compare ...
[5]
ODONATA Definition & Meaning - Merriam-Webster
The meaning of ODONATA is an order of insects containing the dragonflies and ... Word History. Etymology. New Latin, irregular from Greek odōn tooth + ...
[6]
Etymology - Dragonflies of Ireland - Habitas
Etymology of latin and greek names ; Aeshna, Gr. ugly, mishapen ; Aeshnidae, see Aeshna ; Agriidae ; Anax, Gr. lord, master, king ; arctica, from the polar regions; ...
[7]
LIBELLULA Definition & Meaning - Merriam-Webster
Word History. Etymology. New Latin, diminutive of libella dragonfly, from Latin, level (instrument); from the horizontal position of the wings ...
[8]
Cultural Odonatology References
The dragonfly, now called "tombo" in Japanese, used to be called "akitsu". Japan was once called Akitsushima meaning "The Island of the Dragon-fly". Songs They ...
[9]
http://casswww.ucsd.edu/archive/personal/ron/CVNC/odonata/cultural_odonatology.html
[10]
Dragonflies and damselflies - Order Odonata - The Australian Museum
The Order Odonata is divided into two suborders, the Anisoptera containing the dragonflies, and the Zygoptera containing the damselflies.
[11]
Order Odonata – ENT 425 – General Entomology - NC State University
Greek Origins of Name: Odonata, derived from the Greek “odonto-“, meaning tooth, refers to the strong teeth found on the mandibles of most adults.
[12]
[PDF] A molecular phylogeny and classification of Anisoptera (Odonata)
Aug 20, 2015 · A phylogeny of Anisoptera employing 510 representatives of 184 genera (of ca. 380) in 11 families is presented based on an analysis.
[13]
Evolutionary history and divergence times of Odonata (dragonflies ...
Nov 19, 2021 · Dragonflies and damselflies are among the earliest flying insects with extant representatives. However, unraveling details of their long ...Missing: etymology credible
[14]
Palaeozoic giant dragonflies were hawker predators - Nature
Aug 14, 2018 · The largest insects to have ever lived were the giant meganeurids of the Late Palaeozoic, ancient stem relatives of our modern dragonflies.
[15]
Dragonfly fossil calibrations - Palaeontologia Electronica
This paper reviews crown group fossils for Odonata, using them to calibrate divergence time analysis for nine nodes, providing the oldest fossils for each.
[16]
New fossil Odonata from the Upper Jurassic of Bavaria ... - Biotaxa
Dec 20, 2019 · Abstract. Three new taxa of odonates are described from the Upper Jurassic Solnhofen limestone from Eichstätt and Painten in Bavaria (Germany), ...
[17]
Evolutionary history and divergence times of Odonata (dragonflies ...
Nov 19, 2021 · A possible exception is a putative mid-Cretaceous diversification of Zygoptera, suggested by both our analysis and the content of Myanmar ...
[18]
Phylogenetic analysis of higher‐level relationships of Odonata
Apr 7, 2003 · Wing vein homologies. Wing venation has long provided students of Odonata with a rich source of diagnostic characters at all taxonomic levels.
[19]
Phylogeny and classification of Odonata using targeted genomics
The molecular revision of the family by Sanchez Herrera et al. (2018) showed that the family is monophyletic. The lateral abdominal gills of the nymphs are ...
[20]
Article Phylogenetic relationships and divergence times of Odonata ...
Feb 21, 2025 · Our results revealed significant differences in divergence time estimates inferred solely from mitochondrial or nuclear data within Odonata.Missing: post | Show results with:post
[21]
Comparative analysis of mitochondrial genomes among the family ...
Aug 14, 2022 · A well-constructed molecular phylogeny could substantially benefit the understanding of evolutionary relationships of major lineages and ...
[22]
(PDF) Temperature response of growth of larval dragonflies
Jan 11, 2016 · All information indicates that larval growth is generally warm adapted, with species-specific variation of Topt of 21–31°C, Tmin of 8–12°C, and ...
[23]
[PDF] Introduction to Dragonfly and Damselfly Watching
Dragonflies and damselflies are insects with two pairs of net-veined wings and large compound eyes. Damselflies have separated eyes, while dragonflies do not. ...
[24]
Global diversity of dragonflies (Odonata) in freshwater - ResearchGate
Aug 6, 2025 · The highest diversity is found in flowing waters in rain forests of the tropics, the Oriental and Neotropical regions being the most speciose.
[25]
Focus on African freshwaters: Hotspots of dragonfly diversity and ...
Feb 24, 2012 · Major centers of dragonfly diversity in Africa are tropical forest areas that include highlands. Most threatened species – as classified by the ...
[26]
[PDF] Diversity of Nearctic Dragonflies and Damselflies (Odonata)
Jul 18, 2022 · ... absence distribution ... Some of the areas with the lowest sampling effort may indeed be areas where dragonflies are effectively absent; however, ...
[27]
Dragonflies and damselflies of Madagascar and the ... - Vahatra
This is the first book on the spectacular dragonflies and damselflies of the Malagasy Region, covering over 190 species known from Madagascar, as well as 36 ...
[28]
[PDF] Recent range shifts of European dragonflies provide support for an ...
Results Lentic species of the southern group expanded their range boundaries on average 115 km northwards per decade, whereas lotic species of the southern ...
[29]
Distribution trends of European dragonflies under climate change
Mar 12, 2019 · In this paper, we investigate how the occurrence of dragonflies in Europe has changed in recent decades, and if these changes are in parallel with climate ...
[30]
World Odonata List (NEW) - University of Puget Sound
Apr 5, 2025 · This is an ongoing attempt to list all of the valid species of Odonata. It is based on past compilations by the authors listed below and constant additional ...
[31]
Zygoptera systematics: past, present, and future - Oxford Academic
Jul 7, 2025 · Odonata is a midsized insect order (~6,420 species) comprised of 3 suborders: Anisoptera (dragonflies), Zygoptera (damselflies), and the ...
[32]
A Bibliometric Analysis of the Global Research in Odonata - MDPI
Dec 5, 2022 · The countries with the most Odonata publications were the USA, Brazil and China. Most studies were conducted on streams, ponds and rivers.
[33]
Odonata enter the biodiversity crisis debate: The first global ...
Odonate species richness is clustered in the Neotropical and Indo-Malayan realms, which contain almost 60% of the world's Odonata diversity (Fig. 2) ...
[34]
[PDF] Damselflies and dragonflies
Oct 1, 2015 · Endangered and threatened wildlife and plants; endangered status for 23 species on Oahu and designation of critical habitat for 124 species ...
[35]
The native dragonflies and damselflies of Hawaiʻi
May 18, 2018 · Evolving over 20 million years of isolation, these native damselflies and dragonflies are threatened by habitat loss and introduced predators like mosquito ...
[36]
New assessment finds dragonflies and damselflies in trouble ...
Feb 9, 2022 · In total, the assessment found that about 16% of 6,016 species of dragonflies and damselflies, making up the taxonomic order Odonata, are at ...Missing: discoveries | Show results with:discoveries
[37]
Polarized vision in the eyes of the most effective predators
Jan 21, 2025 · Their compound eyes, composed of up to 30,000 individual ommatidia ... Sherk TE (1977) Development of the compound eyes of dragonflies (Odonata).
[38]
Dragonfly Biology | MDS
Dragonflies have three main body parts: head, thorax, and abdomen. The head has a complex mouth, the thorax controls movement, and the abdomen has ten segments.Missing: scientific | Show results with:scientific
[39]
Evolution of Flight Muscle Contractility and Energetic Efficiency
Synchronous direct flight muscles are found in lower species of insects like locusts (Orthoptera) and dragonfly (Odonata), which have wing beats at lower ...
[40]
Ontogenetic shifts in functional morphology of dragonfly legs (Odonata
In contrast, adult Anisoptera use their legs as devices for prey catching and handling by forming a “basket” during flight (St. Quentin, 1953), while locomotion ...
[41]
Beyond the wing planform: morphological differentiation between ...
Jan 18, 2016 · ... nodus provides stability during the stroke, whereas the pterostigma reduces the loss of muscular energy. Additionally, wing surface ...
[42]
Chapter 4 Classes of Arthropod Pests of the Urban Community
Aug 23, 2002 · The abdomen is 10- or 1l-segmented. Paired spiracles are located between the second and third coxae and in the third and fourth abdominal ...
[43]
THE PERMEABILITY OF DRAGONFLY MALPIGHIAN TUBULE ... - NIH
THE PERMEABILITY OF DRAGONFLY MALPIGHIAN TUBULE CELLS TO PROTEIN USING HORSERADISH PEROXIDASE AS A TRACER ... Hormonal control of excretion. J Exp Biol. 1966 Jun; ...
[44]
Spiracle Control in Adult Dragonflies (Odonata)
All the spiracles of the adult dragonfly possess valves which are closed by a short triangular muscle (the closer) and opened by a cuticular spring on ...INTRODUCTION · SPIRACLE BEHAVIOUR IN... · PERIPHERAL CONTROL OF...
[45]
Dragon colors: the nature and function of Odonata (dragonfly and ...
Mar 10, 2022 · Odonates present three of four mechanisms of producing colors recorded in animals: pigmentary absorption, structural reflectance, and ...
[46]
Pterin-pigmented nanospheres create the colours of the ...
Apr 17, 2019 · While the coloration primarily arises from a disordered assembly of nanospheres in the epidermis, morph-dependent changes result from ...
[47]
The genomics and evolution of inter-sexual mimicry and female ...
Nov 6, 2023 · The common bluetail damselfly Ischnura elegans (Odonata) has three female-limited morphs (namely O, A and I) that differ in colouration, ...
[48]
Anax junius | INFORMATION - Animal Diversity Web
Feb 26, 2014 · Additionally, their brown or green coloration serves as camouflage. More green nymphs are found in the spring and summer, when there is more ...
[49]
Diversity of UV Reflection Patterns in Odonata - Frontiers
Jul 8, 2020 · Dragonflies and damselflies (Odonata) are large-eyed diurnal insects that exhibit a variety of color patterns on their wings and/or bodies.
[50]
Dragonfly – Nursery IPM - Sites@Rutgers
Aug 26, 2025 · Large compound eyes with a nearly 360-degree field of view and ultraviolet vision allows them to catch prey in midair with a 95% success rate ...
[51]
Species Spotlight - Dragonflies (U.S. National Park Service)
Jan 15, 2025 · This is combined with an almost complete 360 degree field of vision processed at 200 images per second. To humans, this would appear as HD ...
[52]
The separation of visual axes in apposition compound eyes - Journals
The acute zone is usually forward looking, but is upward looking in some insects which catch prey or mate in flight. In addition, some dragonflies have a ...
[53]
Extraordinary diversity of visual opsin genes in dragonflies - NIH
Human color vision is tri-chromatic, with three opsins expressed in cone photoreceptors that are sensitive in the red, green, and blue region of the spectrum.
[54]
The price of looking sexy: visual ecology of a three‐level predator ...
Sep 21, 2016 · Most dragonflies and damselflies are at least trichromatic, with sensitivity from the UV to the red part of the spectrum (reviewed in Bybee et ...
[55]
Dragonfly Neurons Selectively Attend to Targets Within Natural ...
Apr 4, 2022 · Here, we describe intracellular responses of CSTMD1 (an identified STMD) to the visual presentation of targets embedded within cluttered, natural scenes.
[56]
Heuristic Rules Underlying Dragonfly Prey Selection and Interception
Apr 24, 2017 · Because the takeoff reaction time is ∼75 ms, this should be the ... prey interception time (348 ± 110 ms). If the dragonfly can ...
[57]
https://doi.org/10.2307/1936214
[58]
https://doi.org/10.3390/biology14080956
[59]
Systems-Level Analysis of Resonant Mechanisms for Flapping-Wing ...
The Anax imperator dragonfly has a wing-beat frequency range of 29.2 to 47.5 Hz [8], corresponding to a deviation from the center frequency of ± 2 4 % ± 24 % .
[60]
Structure, function and evolution of insect flight muscle - PMC
In fruitfly (Drosophila), it is reported that the wing-beat frequency is increased from 185 to 195 Hz when the frequency of nerve impulses is increased from 3 ...
[61]
Phasing of dragonfly wings can improve aerodynamic efficiency by ...
In contrast to most other insects, direct musculature acts at each wing base, enabling dragonflies to control each wing independently.Missing: maneuverability | Show results with:maneuverability
[62]
The secret of dragonflies' flight - Phys.org
Nov 25, 2014 · Dragonflies can easily right themselves and maneuver tight turns while flying. Each of their four wings is controlled by separate muscles.
[63]
Experimental and Numerical Investigation on Dragonfly Wing and ...
Jan 17, 2018 · During this process, the maximum vertical acceleration could reach 20 m/s2. Evidence also shows that acceleration is generated only by the ...Missing: per | Show results with:per
[64]
The role of the leading edge vortex in lift augmentation of steadily ...
Jul 26, 2017 · The presence of a stable leading edge vortex (LEV) on steadily revolving wings increases the maximum lift coefficient that can be generated from the wing.
[65]
Research on the aerodynamic characteristics of dragonfly leading ...
Sep 10, 2024 · In these studies, the wavy leading edge significantly enhances fluid attachment, effectively reducing drag, increasing lift, and raising the ...INTRODUCTION · OBSERVING, MODELING... · RESULTS DISCUSSION AND...
[66]
Fastest flying insect | Guinness World Records
The maximum airspeed attained by this dragonfly is 58 km/h (36 mph), making it one of the swiftest fliers in the insect world.
[67]
HiFly-Dragon: A Dragonfly Inspired Flapping Flying Robot ... - MDPI
Mar 28, 2024 · We propose a compact microrobot dragonfly with four tandem independently controllable wings, which is directly driven by four modified resonant flapping ...
[68]
Effects of wing kinematics, corrugation, and clap-and-fling on ...
The study showed that the clap-and-fling effect improved the vertical lift ... Kim et al. Effects of corrugation of the dragonfly wing on gliding performance ...
[69]
Heuristic Rules Underlying Dragonfly Prey Selection and Interception
Aug 6, 2025 · Although they do not use gravity to reduce their energy costs, dragonflies are capable of intercepting prey, using highly efficient flight paths ...
[70]
Flight of the dragonflies and damselflies - PMC - PubMed Central
This work is a synthesis of our current understanding of the mechanics, aerodynamics and visually mediated control of dragonfly and damselfly flight.
[71]
Prey capture by dragonfly larvae (Odonata; Anisoptera)
Aug 6, 2025 · Dragonfly larvae are aquatic ambush predators, capturing their prey with a strongly modified extensible mouthpart.
[72]
Intersection of motor volumes predicts the outcome of ambush ... - NIH
Dragonfly nymphs hunt by remaining motionless and waiting for prey to come within ambush distance (Robert Evans Snodgrass, 1954; Pritchard, 1965) (Fig. 1).
[73]
(PDF) Fliers and perchers among Odonata: Dichotomy or ...
Aug 7, 2025 · Some departures from expectation based on the percher/flier dichotomy reflect the increased flight activity that occurs at the rendezvous under ...
[74]
[PDF] INTRODUCTION - University of California Press
The two general types of aerial feeding used by dragonflies are hawk- ing (the constant pursuit of flying insects) and sallying (darting out from a perch to ...
[75]
Dragonflies | The Real Dirt - UC Agriculture and Natural Resources
Apr 21, 2023 · They have almost 360-degree vision due to their two huge compound eyes, which comprise nearly the entire bulk of the dragonfly head; each ...Missing: field view
[76]
(PDF) Dragonflies as an Important Aquatic Predator Insect and Their ...
Aug 9, 2025 · ... keystone species, exerting a ... The dragonfly nymphs as predators could be used in biological control of Aedes mosquitoes.
[77]
Predation potential of odonates on mosquito larvae - ResearchGate
Aug 6, 2025 · Many of the potential predator taxa such as dragonfly and damselfly larvae are frequently observed as effective larval mosquito predators in ...
[78]
(PDF) Prey size selection and distance estimation in foraging adult ...
Aug 7, 2025 · Our results indicated that dragonflies were highly selective for bead size. Furthermore, the smaller Sympetrum preferred beads of smaller size ...
[79]
[PDF] Integrative and Comparative Biology - The Combes Lab
Jun 19, 2013 · Because larger insects generally fly faster than smaller ones, we expected that larger prey would be more difficult for dragonflies to catch, ...
[80]
Dragonfly Larvae (U.S. National Park Service)
Jan 30, 2023 · Dragonflies live in streams, wetlands, lakes, and other water bodies. We usually see adult dragonflies' shimmering wings and ...
[81]
Lakefront property owners can protect dragonflies as they reduce ...
Apr 25, 2012 · Dragonflies spend their lifecycles in and near aquatic environments, and are most abundant in standing waters such as lakes, ponds and wetlands ...
[82]
[PDF] Guidelines for Creating & Managing Habitat for Dragonflies ...
A diversity of aquatic vegetation is vital for a good dragonfly pond; plants take up nutri- ents, oxygenate and cool the water, create shelter and habitat for ...
[83]
Relative Influence of Prior Life Stages and Habitat Variables ... - MDPI
Abundance and diversity of adult Odonata increase in riparian areas with tall wetland vegetation such as sedges and bulrushes, although the underlying reasons ...Missing: preferences | Show results with:preferences
[84]
Aquatic and terrestrial drivers of dragonfly (Odonata) assemblages ...
Vegetation provides adults with perching structures for thermoregulation, foraging ... Foraging modes in an assemblage of odonate larvae—effects of prey and ...
[85]
Riparian and in-channel habitat properties linked to dragonfly ...
Oct 19, 2020 · Trees provide habitat structures to support the successful emergence of dragonflies, and are likely favored over other emergence structures ...
[86]
Tracking dragons: stable isotopes reveal the annual cycle of a long ...
Dec 19, 2018 · Previous estimates of darner daily migration rates indicate this journey would take 55.42 ± 4.13 days to complete [12]. Therefore, the first ...
[87]
Simple rules guide dragonfly migration - PMC - PubMed Central
Green darners exhibited distinct stopover and migration days. On average, they migrated every 2.9±0.3 days, and their average net advance was 58±11 km in 6.1±0 ...
[88]
European Dragonfly Migratory Isoscape - Ecosystems in the Sky
Jul 24, 2020 · Using stable isotopes, the large scale pattern of Europe's migratory dragonflies is to be revealed ... Hemianax ephippiger, Vagrant emperor, Brun ...
[89]
Migrant Dragonflies Project
Vagrant Darter · Vagrant Emperor · Red-veined Darter · Scarlet Darter · Yellow-winged Darter. Typical resident migrant species to Britain include: Four-spotted ...
[90]
Landscape-scale terrestrial factors are also vital in shaping Odonata ...
Dec 3, 2019 · Habitat loss and fragmentation causes a decline in insect populations. Odonata (both dragonflies and damselflies) are especially threatened ...
[91]
Dragonflies threatened as wetlands around the world disappear
Dec 9, 2021 · The assessment of the world's dragonflies and damselflies reveals that 16% out of 6,016 species are at risk of extinction, as their freshwater ...
[92]
Connectivity of stormwater ponds impacts Odonata abundance ... - NIH
Feb 28, 2024 · Our results suggest that dragonflies are positively affected by increased connectivity in an urban landscape, with no benefit of connectivity to damselflies.Missing: percentage | Show results with:percentage
[93]
CHAPTER 16 Territoriality in odonates - Oxford Academic
This chapter discusses causes and consequences of territorial behaviour in odonates. In territorial species, males may use two mating tactics or strategies.
[94]
Reproductive Tactics of Sexes and Fitness in the Dragonfly ...
Apr 16, 2007 · The prediction is that females that copulate with satellite males will more often accept second copulations, while females that mate with ...
[95]
Sperm competition in Odonata (Insecta): the evolution of female ...
Feb 28, 2006 · Odonates (dragonflies) are well known for the ability of the males to displace sperm stored in the female's sperm-storage organs during ...
[96]
Aerial adult dragonflies are highly sensitive to in-water conditions ...
This makes dragonfly assemblages excellent indicators of water quality ... exophytic oviposition, and a positive correlation with endophytic oviposition.
[97]
Different Oviposition Strategies of Closely Related Damselfly ...
Jan 10, 2019 · The insertion of eggs into living or decaying plant tissues, referred to as endophytic oviposition, reduces water loss, minimizes winter ...
[98]
Two matings lead to more copulatory wounding than a ... - PubMed
Jun 18, 2025 · Copulation can result in males inflicting wounds to the female genitalia, so-called traumatic mating. Such wounds are potentially costly as ...
[99]
Why Female Dragonflies Go to Extreme Lengths to Avoid Sex
May 1, 2017 · A scientist has observed that female moorland hawker dragonflies freeze mid-air, crash to the ground, and lie motionless when faced with aggressive males.<|control11|><|separator|>
[100]
Macroinvertebrates - Kenilworth Park & Aquatic Gardens (U.S. ...
Dec 6, 2023 · Dragonflies have a three-part life cycle: egg, larva, adult. A dragonfly's life cycle is an example of incomplete metamorphosis. Dragonfly eggs ...
[101]
Dragonflies and Damselflies | Yard and Garden - Iowa State University
The eggs hatch after three to five weeks into nymphs called naiads. The naiads live more or less buried in the mud or attached to submerged plants for several ...
[102]
Dragonfly Life Cycle | MDS
The migratory Wandering Glider will take as few as four weeks to complete its development from egg to adult, whereas there are Asian species that take as long ...Missing: instars | Show results with:instars
[103]
Dragonfly and Damselfly | San Diego Zoo Animals & Plants
The nymph feeds on other insect larvae, tadpoles, and small fish. It is ... Nymphs molt 10 to 20 times, and the time between molts is called an instar.<|control11|><|separator|>
[104]
Larva - Bedfordshire Odonata
Dragonfly larvae do not have caudal lamellae, and instead use jet propulsion to move rapidly through the water, the only insects to do so. Muscular contractions ...
[105]
Odonates (Dragonflies & Damselflies) - Mass Audubon
The newly emerged adult, termed a teneral, is fragile and unable to fly at first. If you look carefully on vegetation along the edges of weedy ponds or on ...Missing: vulnerable | Show results with:vulnerable
[106]
Simulated climate change increases larval mortality, alters ...
Mar 13, 2018 · There was a trend toward reduced larval survival with increasing temperature. Warming strongly affected the phenology of adult emergence, ...
[107]
The Combined Effects of Warming and Body Size on the Stability of ...
We experimentally tested how water temperature alters the functional response and population stability of dragonfly nymphs (Cordulegaster boltonii) feeding on ...
[108]
Sexual size dimorphism and sex ratios in dragonflies (Odonata)
Aug 7, 2025 · As dragonflies have a 50/50 sex ratio at hatching [21], this hypothesis about sex-specific mortality can be tested by examining population sex ...
[109]
The Effects of Dispersal and Recruitment Limitation on ... - jstor
These results and a male biased sex-ratio in adults observed ... dragonfly communities at small artificial ponds. ... would be male biased (1.5:1). This assumption ...
[110]
Sexual dimorphism in Odonata: Age, size, and sex ratio at emergence
Sep 16, 2025 · Protandry, here defined as the earlier emergence of males, is a common feature in life histories and could be the result of sexual selection ...
[111]
Ornamented species incur higher male mortality in the larval stage
May 17, 2023 · As dragonflies have a 50/50 sex ratio at hatching [21], this hypothesis about sex-specific mortality can be tested by examining population sex ...Missing: protandry differential
[112]
Density Dependence, Hatching Synchrony, and within‐Cohort ...
Jan 1, 1996 · The rate at which cannibalism increased with size differences between larvae was greater when food was absent than it was when food was provided ...Missing: nymph | Show results with:nymph
[113]
Density-Dependent Cannibalism in Dragonfly Nymphs (Odonata
Dec 17, 2021 · ... nymphs consumed) we experimentally manipulated nymph density in aquaria. ... density-dependent cannibalism among dragonfly nymphs. Keywords ...Missing: regulation population cohorts
[114]
Critical declines in European Odonata populations highlighted in ...
Aug 8, 2024 · Over the past decade, it has experienced a decline of over 50%. Of the 146 species assessed, 21% are currently threatened with extinction, ...
[115]
(PDF) Odonata enter the biodiversity crisis debate: The first global ...
Aug 6, 2025 · We have found that one in 10 species of dragonflies and damselflies is threatened with extinction. This threat level is among the lowest of ...
[116]
REVIEW OF PARASITIC ASSOCIATIONS OF LARVAL WATER ...
May 31, 2012 · Associations of parasitic larvae with insect hosts are confirmed for species of water mites representing 70 of the nearly 300 described genera, ...
[117]
Escape of parasitic water mites from dragonfly predators attacking ...
Aug 6, 2025 · Freshwater ectoparasitic mites negatively alter host population dynamics by reducing survivorship, mating success, fitness and altering activity ...Missing: mobility | Show results with:mobility
[118]
Patterns of Gregarine Parasitism in Dragonflies: Host, Habitat, and ...
Gregarines are ubiquitous protozoan parasites that infect arthropods worldwide. More than 1600 gregarine species have been described, but only a small ...
[119]
Parasites, proteomics and performance: effects of gregarine gut ...
Dec 15, 2007 · In previous work, we found that dragonflies infected with gregarine gut parasites have reduced muscle power output, loss of lipid oxidation ...
[120]
Two rare ophiocordycipitaceous fungi newly recorded in Taiwan - NIH
Oct 24, 2015 · Ophiocordycipitaceous fungi are parasites of more than ten orders of Arthropoda and one fungal genus Elaphomyces (Kobayasi 1941, 1981, 1982 ...
[121]
Dragonfly mind control? - Katatrepsis
Mar 14, 2015 · The most famous of these kinds of fungi is perhaps Ophiocordyceps unilateralis, which infects ants and influences their behaviour. The fungus ...
[122]
Effects of Life Stage, Site, and Species on the Dragonfly Gut ... - NIH
Jan 28, 2020 · In this study, we surveyed the gut microbiomes of 13 species of dragonfly collected from five different locations subject to different levels of human impact.
[123]
Diverse circular ssDNA viruses discovered in dragonflies (Odonata
This study explored the diversity of CRESS-DNA viruses circulating among insect populations by targeting dragonflies (Epiprocta), top insect predators.
[124]
(PDF) Identification of viruses associated with larvae of the dragonfly ...
Aug 7, 2025 · Dragonflies and damselflies are hosts to a variety of parasites and pathogens. However, very few stud- ies have investigated which viruses ...Missing: outbreaks | Show results with:outbreaks
[125]
Do latitudinal and bioclimatic gradients drive parasitism in Odonata?
Recent advances in the study of host–parasite interactions have unravelled spatial and environmental gradients (Stephens et al., 2016). Parasites are notable ...
[126]
A critical overview of progress in studies of migration of dragonflies ...
Nov 20, 2012 · In general, migrant Odonata are not currently under threat from human activity, largely because they are adapted to more or less unstable ...
[127]
Agricultural use of wetlands: opportunities and limitations - PMC
Worldwide, wetlands have been drained to convert them into agricultural land or industrial and urban areas. A realistic estimate is that 50 % of the world's ...Wetland Ecosystem Services... · Agriculture In Dutch... · Literature Cited
[128]
Scientists' warning on the need for greater inclusion of dragonflies in ...
Mar 15, 2025 · In terms of threat, of those assessed on the IUCN Red List of Threatened Species (www.redlist.org), 29.4% are currently Data Deficient, while ...<|control11|><|separator|>
[129]
[PDF] Dragonflies & Damselflies - IUCN
Aug 8, 2024 · The EU's Member States stretch from the Arctic. Circle in the north to the Mediterranean in the south, and from the Atlantic coast and several.Missing: absence | Show results with:absence
[130]
[PDF] Wetlands and agriculture: impacts of farming practices ... - Ramsar.org
For example, more than 20% of the world's Wetlands of International Importance are affected by dams, and 20% by drainage.<|separator|>
[131]
[PDF] European Dragonflies: - Conservation Planning Specialist Group
Of the 142 species assessed, 29 (21.0%) extant species for which sufficient ... Numbers indicate the number of species in each Red List Category. Not ...
[132]
Mobile application for biological species recognition in citizen science
Jan 9, 2019 · We describe a mobile application that allows the public to map and report occurrences of the odonata species (dragonflies and damselflies) found in the Czech ...
[133]
Dragonfly Mercury Project - Citizen Science - National Park Service
May 27, 2025 · The Dragonfly Mercury Project (DMP) is a nationwide study using citizen scientists to collect dragonfly larvae for mercury analysis, helping to ...
[134]
Using Technology and Citizen Science to Understand Dragonfly ...
Apr 27, 2016 · If you are an iPhone user, you can download the Dragonfly ID app to help you identify the dragonflies you see. Authors. Michele Blackburn.
[135]
Helmet (Kawari-kabuto) Surmounted by a Dragonfly - Japanese
$$12.95 deliveryA favorite emblem of the samurai, the dragonfly was known as kachi-mushi ("victorious insect") and was respected for its hunting technique.Missing: symbolism folklore<|control11|><|separator|>
[136]
Dragonfly 蜻蜓 Tonbo - Planet Dizer - WordPress.com
Jul 18, 2021 · For that they have been used as a symbol of bravery and victory by the samurai and were called kachimushi 勝ち虫 (victory insect). In the 17th ...
[137]
Native American Dragonfly Mythology
To the Navajo tribe, the dragonfly is a symbol of water, and dragonfly images frequently appear in sacred sandpaintings to represent the element of water.
[138]
The Origin of the Dragonfly: A Zuni Tale
Aug 15, 2011 · He is followed by his companion, the green dragonfly. Together they arrive with the rains, harbingers of life and good health for the spring ...Missing: dream catchers bringers
[139]
The dragonfly by Matsuo Basho - Famous poems - All Poetry
This haiku by Matsuo Bashō captures a brief moment in nature, where a dragonfly attempts but fails to land on a blade of grass.
[140]
How To Paint A Water Lily by Ted Hughes - Famous poems - All Poetry
The speaker observes the vibrant dragonflies and the dangerous creatures in the water, highlighting the juxtaposition between beauty and violence. Hughes's ...
[141]
Dragonfly amulet - Middle Kingdom - The Metropolitan Museum of Art
Artwork Details ; Title: Dragonfly amulet ; Period: Middle Kingdom ; Dynasty: Dynasty 12–13 ; Date: ca. 1981–1640 B.C. ; Geography: From Egypt, Memphite Region, ...Missing: motifs | Show results with:motifs
[142]
Unraveling the Symbolic Tapestry of Dragonfly Tattoos: Meanings ...
Mar 11, 2025 · The dragonfly represents change, growth, and self-realization, much like its metamorphosis from a nymph to a winged beauty.Dragonfly Tattoos and... · Dragonfly Tattoos and Cultural...
[143]
The drone inspired by a dragonfly | Department of Biology
Skeeter will have four independently flapping wings, a camera and microphone, and will fly at up to 45 kph. Yet it will be no more than 12cm long and 30g in ...Missing: biomimicry micro clap
[144]
Flap Like a Dragonfly
Jun 27, 2017 · The resulting aircraft, named Skeeter, is inspired by a dragonfly. And like the insect, Skeeter employs two sets of wings, and flight results ...Missing: project | Show results with:project
[145]
Cameras inspired by insect eyes could give robots a wider view
May 15, 2024 · Cameras inspired by the compound eyes of insects enable an extremely wide field of view without expensive lenses, potentially offering cheap, simple and ...
[146]
How the natural world is inspiring the robot eyes of the future - Nature
May 29, 2025 · The design of a compound eye holds certain advantages over conventional cameras. To sense movement, for example, flying insects register ...
[147]
A controllable dual-catapult system inspired by the biomechanics of ...
26; the dragonfly-larvae-inspired compliant robot that can emulate a ... A similar pressure pulse will produce jet propulsion if the dragonfly nymph opens its ...