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Carpal bones

The carpal bones are a set of eight small, irregularly shaped bones located in the wrist that connect the distal ends of the radius and ulna bones of the forearm to the bases of the five metacarpal bones of the hand. These bones are organized into two transverse rows of four each, forming a flexible yet stable structure essential for wrist mobility. The proximal row, positioned closest to the forearm and arranged from lateral (thumb side) to medial (pinky side), includes the scaphoid, lunate, triquetrum, and pisiform bones. The distal row, adjacent to the hand, consists of the trapezium, trapezoid, capitate, and hamate bones. Each carpal bone has unique articular surfaces that interlock with neighboring bones and ligaments, creating a concave anterior arch roofed by the flexor retinaculum to form the carpal tunnel, through which flexor tendons and the median nerve travel. Functionally, the carpal bones contribute to the radiocarpal and midcarpal , enabling a wide range of movements including flexion, extension, , adduction, and circumduction while providing structural support for load transmission during gripping and weight-bearing activities. They serve as attachment sites for numerous ligaments and tendons, enhancing stability and facilitating precise hand coordination. supply to the carpals primarily arises from branches of the radial and ulnar arteries, with the scaphoid being particularly vulnerable to due to its retrograde vascularization. The carpal bones develop through from mesenchymal precursors during fetal hand , which occurs primarily between 6 and 14 weeks of , though full centers appear postnatally in a sequential manner starting with the capitate and hamate around 1 to 3 months of age. Clinically, the carpals are prone to injury, with scaphoid fractures being the most common carpal injury due to falls on an outstretched hand, often requiring or surgical fixation to prevent complications like nonunion. Disorders such as arise from compression within the tunnel, affecting function and leading to pain, numbness, and weakness in the hand.

Anatomy

Bones

The carpal bones comprise eight short bones that form the , arranged in two transverse rows between the distal and the metacarpal bases. The proximal row, situated closest to the , consists of four bones from radial (lateral) to ulnar (medial): the scaphoid, lunate, triquetrum, and pisiform. The distal row, adjacent to the hand, similarly includes four bones in radial-to-ulnar order: the , , capitate, and hamate. This arrangement allows for the complex mobility and stability of the . A widely used mnemonic for recalling the sequence of carpal bones from proximal radial to distal ulnar is "She Looks Too Pretty; Try To Catch Her," where each word's initial letter corresponds to scaphoid (She), lunate (Looks), triquetrum (Too), pisiform (Pretty), (Try), (To), capitate (Catch), and hamate (Her). The following table summarizes the morphology, position, key features, and primary articulations of each carpal bone:
BoneRowShape and MorphologyKey FeaturesPrimary Articulations
ScaphoidProximalBoat-shaped (navicular) with a peanut-like body, elongated waist, and concave distal surface; largest in the proximal row.Prominent tubercle on the anterior (palmar) surface, forming part of the ; nutrient foramina typically on dorsal ridge.Proximal: radius; Distal: , , capitate; Interosseous: lunate.
LunateProximalCrescent- or moon-shaped (luna) with a concave proximal surface and convex distal; central position in proximal row.Concavo-convex articular facets; variable including type I (single facet) or type II (medial facet for hamate).Proximal: radius; Distal: capitate; Interosseous: scaphoid, triquetrum.
TriquetrumProximalor irregular-shaped with a flattened ulnar surface and three articular facets.Small facet on palmar surface for pisiform; vascular supply via dorsal and palmar branches.Proximal: complex (ulnar side); Distal: hamate; Interosseous: lunate, pisiform.
PisiformProximalSmall, nodular, and pea-like sesamoid ; ovoid with irregular surfaces.Embedded within the flexor carpi ulnaris ; unique secondary ossification center among carpals.Interosseous: triquetrum (single facet).
TrapeziumDistalor irregularly shaped with a ridged palmar surface and four articular facets. on anterior ridge for transverse carpal attachment; supports thumb opposition.Proximal: scaphoid; Distal: first metacarpal; Interosseous: , second metacarpal.
TrapezoidDistalWedge- or quadrangular-shaped; smallest carpal overall.Concave proximal surface; minimal surface projections.Proximal: scaphoid; Distal: second metacarpal; Interosseous: , capitate.
CapitateDistalHead-, body-, and neck-shaped with a rounded proximal head and elongated shaft; largest carpal overall.Rounded head for scaphoid articulation; waist-like constriction at neck.Proximal: scaphoid, lunate; Distal: third metacarpal; Interosseous: hamate, .
HamateDistalWedge-shaped with a triangular base and prominent hook.Hamulus (hook) process on palmar surface for flexor and attachment; three articular facets.Proximal: triquetrum, lunate (via capitate); Distal: fourth and fifth metacarpals; Interosseous: capitate, pisiform (via coalition variant).

Joints

The carpal bones articulate at several synovial joints that facilitate wrist mobility, primarily classified into the radiocarpal, midcarpal, and carpometacarpal joints. The radiocarpal joint, also known as the wrist joint, is an ellipsoid synovial joint formed between the distal radius and the proximal row of carpal bones, including the scaphoid, lunate, and triquetrum. The midcarpal joint represents a complex synovial articulation, consisting of multiple gliding interfaces between the proximal and distal rows of carpal bones. The carpometacarpal joints connect the distal row of carpal bones to the metacarpals; those for digits 2 through 5 are plane synovial joints, while the thumb's carpometacarpal joint is a saddle synovial joint. Within the midcarpal complex, specific include those of the proximal row, such as the scapholunate and lunotriquetral joints, which are synovial articulations between adjacent bones in that row. The distal row features synovial joints between the and , the trapezoid and capitate, and the capitate and hamate. These contribute to the overall segmented nature of the wrist's synovial architecture. Each carpal joint is enclosed by a capsule comprising an outer fibrous layer, which provides structural integrity and attaches to the of the articulating bones, and an inner synovial layer that lines the cavity and secretes for lubrication. The capsules of the radiocarpal and midcarpal joints exhibit palmar and reinforcements, where the fibrous layer is thickened to enhance stability without direct attachments. Synovial sheaths and compartments associated with the carpal joints include the common flexor sheath, which extends through the and envelops multiple flexor tendons, and separate extensor sheaths that also traverse this region to minimize during motion. These sheaths form distinct synovial compartments that communicate with certain cavities, such as the ulnar linking to the radiocarpal in some individuals.

Ligaments

The ligaments of the carpus are classified into extrinsic and intrinsic types based on their connections. Extrinsic ligaments originate from the , , or metacarpals and insert onto the carpal bones, providing broad support to the . Intrinsic ligaments, in contrast, interconnect adjacent carpal bones within the carpus itself. Additionally, carpal ligaments are categorized by location as volar (palmar) or , and by association with the proximal or distal carpal rows. They can also be distinguished as membranous (thin, intra-articular sheets) or capsular (thicker, reinforcing the capsules).

Extrinsic Ligaments

The palmar radiocarpal arises from the anterior rim of the distal and attaches to the palmar surfaces of the lunate and capitate bones, forming part of the volar . The radioscaphocapitate , a distinct volar extrinsic band, originates from the styloid process of the and courses distally to insert on the scaphoid tuberosity and capitate, traversing the space of Poirier between the palmar radiocarpal and radioscaphocapitate ligaments. The dorsal radiocarpal extends from the posterior distal to the dorsal aspects of the lunate and triquetrum. The radial collateral ligament originates from the tip of the radial styloid process and inserts onto the radial aspect of the scaphoid, blending with the dorsal and volar radiocarpal ligaments. The ulnar collateral ligament arises from the ulnar styloid process and attaches to the ulnar side of the triquetrum and pisiform, contributing to the ulnocarpal complex. The triangular fibrocartilage complex (TFCC), located ulnarly, is a key component of the ulnocarpal complex and incorporates the ulnar collateral ligament along with dorsal and palmar radioulnar ligaments and an articular disc, attaching from the ulnar styloid and distal radioulnar joint to the triquetrum, lunate, and pisiform. The pisohamate ligament, a volar extrinsic structure, connects the palmar aspect of the pisiform bone to the hook of the hamate.

Intrinsic Ligaments

Intrinsic ligaments primarily link bones within the proximal and distal carpal rows. The interconnects the scaphoid and lunate, consisting of three parts: a thick band, a membranous central portion, and a volar band. The lunotriquetral ligament similarly joins the lunate and triquetrum, with comparable , membranous, and volar components, forming part of the proximal row's interosseous stabilizers. The dorsal intercarpal ligament forms an arcuate structure connecting the dorsal surfaces of the scaphoid, lunate, triquetrum, and to the distal row bones, including the capitate and hamate. On the palmar side, the intrinsic ligaments include the palmar scaphotriquetral ligament and other capsular bands reinforcing the proximal-distal row interfaces.

Accessory bones

Accessory ossicles of the carpal region are supernumerary bones that develop independently from the standard eight carpal bones, arising either from separate secondary centers that fail to fuse during development or from avulsion that become rounded and corticated over time. These ossicles are typically asymptomatic and discovered incidentally on , but they hold as they may mimic acute , leading to unnecessary interventions, or cause pain through impingement, , or in symptomatic cases. Their overall prevalence in the is approximately 1.6% to 9.7%, varying by and imaging modality, with higher rates observed in radiographic surveys of patients. Among the more common accessory ossicles, the os centrale carpi is a small, ovoid bone located in the distal row of the carpus, adjacent to the scaphoid and capitate, often fusing with the scaphoid in early development but persisting separately in about 1% of individuals. It may contribute to symptoms such as intermittent pain, clicking, or due to abnormal , and can be mistaken for a or lead to osteonecrosis. The os styloideum, a variant near the , appears as a prominence at the base of the second or third metacarpal and has a prevalence of approximately 0.3% to 2% in radiographic studies of the general population, with cadaveric studies reporting up to 19%; notably higher prevalence (e.g., 81%) observed in athletes like players, likely due to . Clinically, it is associated with carpal boss , presenting as a painful exacerbated by repetitive motion. Pisiform variants, such as the pisiforme secundarium, involve irregular or multipartite of the , typically at its proximal pole, and occur during the late phase between ages 8 and 12; these are often incidental but can cause ulnar-sided discomfort if associated with impingement. The os triangulare, sometimes referred to as an accessory near the lunate due to its proximity, is positioned distally to the ulnar fovea between the ulnar styloid, lunate, and triquetrum, with a prevalence of about 2.4% in radiographic studies. It is usually asymptomatic but may simulate an ulnar styloid . Rarer accessory ossicles include the os epilunatum, a small bone on the dorsal surface of the lunocapitate adjacent to the lunate, with an estimated of 0.3% to 0.5%. This ossicle is exceptionally uncommon, absent in many large radiographic reviews, and can cause bilateral pain if symptomatic, often requiring surgical excision for relief. The os hamuli proprium represents a separate hook of the , resulting from failure of the hamulus to fuse with the hamate body between ages 12 and 15, and is rare with below 1%. It may be confused with a hamate hook on imaging and, in isolated cases, contribute to carpal tunnel-like symptoms necessitating resection. The os triquetrum accessorium (or secundarium), located near the aspect of the triquetrum, has a prevalence of approximately 1.3% and forms via segmentation of the triquetral . Though generally , it can become symptomatic following , presenting with due to fracture or ligamentous attachment stress, and must be differentiated from avulsion injuries using cortical margins on advanced . Overall, recognition of these ossicles prevents misdiagnosis, with CT or MRI aiding in confirmation of their benign, developmental nature.

Development

Embryonic formation

The carpal bones begin their embryonic development as mesenchymal condensations within the somatic layer of the , emerging during weeks 4 to 6 of as the buds form. These precursors arise from the activation and proliferation of mesenchymal cells in the , which contribute to the foundational skeletal elements of the . This initial phase establishes the basic framework for the eight carpal bones, aligning with the proximodistal outgrowth of the limb bud. The proximal-distal identity of these carpal precursors is regulated by specific expression patterns of , particularly Hoxa-11 and Hoxd-13, which are essential for patterning the . Hoxa-11 is predominantly expressed in the proximal regions, influencing the zeugopod () and adjacent carpal elements, while Hoxd-13 drives distal autopod (hand) development, including the carpal row formation. Disruptions in these genes can alter the segmental identity, leading to malformations in carpal . Chondrogenesis of the carpal anlagen commences around week 6 and is largely complete by week 8, transforming the mesenchymal condensations into cartilaginous models. This process first manifests in the central carpal elements, such as the capitate and hamate, and progresses to form distinct pre-axial (radial, or thumb-side) structures like the scaphoid and , alongside post-axial (ulnar-side) elements including the triquetrum and pisiform. These cartilaginous precursors provide the template for subsequent skeletal maturation. Congenital anomalies of the carpal bones, such as synostosis, originate from failures in the segmentation of these early cartilaginous anlagen during embryogenesis. For instance, scapholunate fusion results from incomplete separation of the interzones between adjacent precursors, often linked to genetic or developmental disruptions in the chondrogenic phase. These conditions highlight the precision required in embryonic segmentation for normal formation.

Ossification

The ossification of the carpal bones occurs through primary endochondral ossification centers from the cartilaginous precursors for most bones. Unlike long bones, carpal bones lack secondary ossification centers and epiphyseal plates, resulting in growth primarily through surface apposition rather than endochondral elongation or fusion events. The sequence of ossification is predictable and begins shortly after birth, providing a reliable marker for skeletal maturity. The capitate ossifies first, typically between 1 and 3 months of age, followed closely by the hamate at 2 to 4 months. The triquetrum and lunate appear next, around 2 to 3 years and 2 to 4 years, respectively. The trapezium and trapezoid ossify between 4 and 5 years, the scaphoid at 4 to 6 years, and the pisiform last, between 8 and 12 years. This timeline exhibits variability, with ossification generally occurring 6 to 12 months earlier in females than in males due to hormonal influences on skeletal development. The pisiform is unique among the carpals as a that ossifies intramembranously within the of the flexor carpi ulnaris, rather than in a preformed cartilaginous model. Its delayed reflects its functional role in , and it shows greater variability in compared to other carpals. In , the radiographic and sequence of carpal centers on hand and X-rays serve as a standard for assessing and overall skeletal maturity, aiding in the diagnosis of growth disorders. These images reveal the progressive mineralization, with the number and size of visible centers correlating to chronological age within typical ranges.
Carpal BoneTypical Ossification Age Range
Capitate1–3 months
Hamate2–4 months
Triquetrum2–3 years
Lunate2–4 years
4–5 years
4–5 years
Scaphoid4–6 years
Pisiform8–12 years

Function

Stability and load transmission

The carpal bones ensure wrist stability through a combination of bony and constraints that maintain alignment under load. The proximal row transmits the majority (~80-85%) of axial loads from the to the distal carpal row, with the scaphoid handling ~50% via the radioscaphoid and the lunate ~35% via the radiolunate ; the transmits ~20% via the TFCC to the ulnar carpals. This balanced transmission prevents excessive on individual bones and supports overall integrity. Ligamentous structures, such as the triangular fibrocartilage complex (TFCC), play a critical role in stabilizing the ulnar side by absorbing and distributing ulnar loads, typically handling about 20% of the total axial force in neutral ulnar variance. Bone geometry further enhances stability through interlocking row linkages, where the concave-convex articulations between proximal and distal rows resist shear forces and promote efficient load transfer. The biomechanical column theory conceptualizes the carpus as three interconnected columns—radial (scaphoid, , ), central (lunate, capitate, hamate), and medial (triquetrum, pisiform)—each optimized for force distribution along specific pathways. This model, originally proposed by in 1921, underscores how these columns collectively transmit compressive forces while minimizing instability. Dynamic stability is augmented by from flexors and extensors, which generate compressive forces across the carpal arch to counteract potential displacement during loading. These muscular contributions complement the static stabilizers, ensuring adaptive responses to varying biomechanical demands.

Role in kinematics

The carpal bones facilitate wrist motion through a kinematic model in which the proximal row (scaphoid, lunate, and triquetrum) functions as an adaptive intercalated segment that rotates relative to the more rigid distal row (, , capitate, and hamate) primarily at the . This inter-row rotation allows the proximal row to adjust its position in response to extrinsic forces and muscle activity, optimizing overall alignment and range. For instance, during radial deviation, the scaphoid flexes while the distal row extends, enabling smooth translation of the hand relative to the . The wrist's multi-axial arise from the combined articulations of the radiocarpal and midcarpal , with motion contributions varying by : approximately 67% of extension occurs at the radiocarpal and 33% at the midcarpal , whereas radial deviation derives about 60% of its motion from the midcarpal and 40% from the radiocarpal . This distribution ensures efficient energy transfer and minimizes stress on individual carpal elements during dynamic activities. Intra-row further enhance adaptability, as bones within each row exhibit subtle translations and rotations to maintain . Muscle force couples, involving the flexor carpi radialis and flexor carpi ulnaris paired with the extensor carpi radialis longus and brevis, coordinate carpal alignment by counterbalancing radial and ulnar forces, thereby stabilizing the proximal row during multi-planar excursions. Fluoroscopic imaging demonstrates this in the dart-thrower's motion, an oblique plane integrating extension with radial deviation and flexion with ulnar deviation, where the proximal row remains relatively stationary to provide a stable base for distal row rotation.

Movements

Flexion and extension

Flexion, or palmar flexion, of the wrist involves bending the hand toward the palmar surface of the forearm, achieving a normal range of motion of 70-80 degrees. Extension, also termed dorsiflexion, moves the hand dorsally toward the posterior forearm, with a typical range of 70-90 degrees. These sagittal plane movements occur through coordinated action at the radiocarpal and midcarpal joints, with each joint contributing approximately 50% to the total excursion in both directions. Mechanically, palmar flexion advances the distal carpal row volarly relative to the proximal row, facilitated by the concave-convex articulations allowing glide of the proximal carpals on the during this motion. In contrast, extension involves volar glide of the proximal carpal row on the distal and , promoting posterior displacement of the hand. These glides maintain congruence and distribute loads across the carpal bones while preventing excessive shear forces. The primary muscles driving flexion are the flexor carpi radialis, flexor carpi ulnaris, and palmaris longus, which originate from the and insert on the metacarpal bases, generating palmar pull via their tendons passing through the flexor retinaculum. Extension is powered by the extensor carpi radialis longus, extensor carpi radialis brevis, and extensor carpi ulnaris, which arise from the lateral epicondyle and forearm , inserting on the metacarpal bases to produce elevation of the hand. These muscles act synergistically to stabilize the during load-bearing activities. Specific carpal bone shifts accompany these motions to optimize kinematics: during extension, the lunate and scaphoid undergo extension relative to the radius, while in flexion, the scaphoid and lunate exhibit flexion, ensuring balanced proximal row alignment and preventing dissociation under tensile loads from surrounding ligaments.

Radial and ulnar deviation

Radial and ulnar deviation refer to the abduction and adduction movements of the wrist in the frontal plane, primarily involving the radiocarpal and midcarpal joints, with contributions from the carpal bones to facilitate hand positioning relative to the forearm. These motions allow the hand to deviate toward the radius (radial deviation) or ulna (ulnar deviation), enabling activities such as pouring from a container or hammering. The carpal bones, particularly in the proximal row (scaphoid, lunate, triquetrum, and pisiform), undergo coordinated translations and rotations during these movements to maintain joint congruence and load distribution. The normal for radial deviation is approximately 15-20°, while ulnar deviation typically measures 30-35°. These ranges occur predominantly at the midcarpal joint, which contributes about 60% to radial deviation and up to 86% to ulnar deviation, with the radiocarpal joint accounting for the remainder. Greater midcarpal involvement in ulnar deviation allows for enhanced mobility on the ulnar side of the . Mechanically, radial deviation elevates the scaphoid while depressing the triquetrum, resulting in supination of the proximal carpal row as the approximates the . In contrast, ulnar deviation involves rotation of the lunate, with extension of the scaphoid and flexion of the triquetrum, promoting pronation of the proximal row. The distal carpal row (, , capitate, and hamate) largely rotates synchronously with the overall motion during deviation. The primary muscles driving radial deviation are the extensor carpi radialis longus and brevis, which originate from the lateral and insert onto the bases of the second and third metacarpals, respectively. Ulnar deviation is primarily powered by the flexor carpi ulnaris, originating from the medial and inserting onto the pisiform, hamate, and fifth metacarpal base. Constraints on these motions include bony impingements from the styloid processes: the radial styloid limits ulnar deviation by contacting the triquetrum, while the ulnar styloid restricts radial deviation through abutment with the triquetrum. Additionally, tension in the triangular fibrocartilage complex (TFCC) increases during ulnar deviation, stabilizing the ulnar carpus and preventing excessive translation, whereas collateral ligaments (radial for ulnar deviation, ulnar for radial deviation) provide soft-tissue checks at the extremes.

Combined and accessory motions

Combined and accessory motions of the carpal bones encompass multiplanar movements that integrate primary flexion-extension and radial-ulnar deviation, enabling complex hand positioning beyond isolated axes. Circumduction involves a conical path traced by the hand, combining these primary motions to produce a circular or elliptical in an oblique plane, which supports tasks requiring broad excursion such as pouring liquids or reaching. This motion arises primarily from coordinated radiocarpal and midcarpal joint contributions, with the proximal carpal row (scaphoid, lunate, triquetrum) rotating and translating relative to the distal row (, , capitate, hamate). Accessory motions refer to subtle translations and rotations that facilitate overall without constituting primary movements. The proximal row undergoes volar (palmar) and glide, allowing the carpal surfaces to slide over the concave distal during flexion and extension, respectively; volar glide supports extension, while glide aids flexion. Additionally, the scaphoid exhibits axial rotation of approximately 38° relative to the lunate during flexion, contributing to the bone's adaptive positioning and load distribution across the carpus. These accessory actions are essential for and are often targeted in techniques to restore mobility. Clinical assessment of these motions includes the piano key sign, performed by stabilizing the in pronation and applying dorsal-volar to the ulnar head; excessive translation or a "bouncing" return indicates axial involving the distal radioulnar and adjacent carpal structures. For thumb opposition, primarily driven by the , the carpal bones provide facilitatory support through the trapezium's saddle articulation, enabling metacarpal rotation and palmar abduction while the scaphoid stabilizes the proximal foundation. The dart-thrower's motion represents a key combined pattern, an oblique arc from radial extension (approximately 30°-40° radial deviation with extension) to ulnar flexion (approximately 40°-50° ulnar deviation with flexion), predominantly occurring at the midcarpal joint with minimal radiocarpal contribution. This motion, spanning up to 120° in healthy wrists, is critical for daily activities like writing, eating, and throwing, as it maintains carpal stability and reduces stress on intrinsic ligaments.

Clinical significance

Fractures and dislocations

Fractures of the carpal bones are relatively uncommon, representing about 18% of all hand fractures, with the scaphoid being the most frequently affected due to its position and vulnerability to hyperextension and axial loading injuries, such as falls on an outstretched hand. Dislocations, often involving ligamentous disruptions around the lunate, typically result from high-energy trauma and can lead to severe instability if not promptly addressed. Acute management emphasizes , confirmation, and surgical intervention for unstable patterns to prevent complications like or . The accounts for 60-70% of all carpal bone fractures, most commonly occurring at the waist (midportion) in over 70% of cases, followed by the proximal pole (20-30%) and distal pole (less than 10%). Waist fractures are typically stable if nondisplaced and managed with thumb spica casting for 6-12 weeks, but proximal pole fractures carry a high risk of due to retrograde blood supply disruption, with rates up to 30% overall and approaching 100% in the most proximal fragments, often necessitating vascularized or salvage procedures like proximal row carpectomy. Other carpal fractures are less common but clinically significant; for instance, hook of hamate fractures, comprising 2-4% of carpal injuries, often arise from direct palmar impact during sports involving gripping, such as or , and may mimic boxer's fractures of the fifth metacarpal but require for diagnosis due to their occult nature on plain radiographs, with treatment ranging from to excision of the hook fragment. Triquetral fractures, the second most frequent at 15-18% of carpal fractures, frequently present as chip avulsions from ligamentous tension during hyperextension falls, managed conservatively with casting unless associated with instability. Perilunate dislocations, often combined with fractures like transscaphoid-transcapitate patterns, represent high-energy injuries disrupting the dorsal radiocarpal ligaments and are classified by the Mayfield staging system, which outlines progressive perilunate instability: Stage I involves tear; Stage II adds capitolunate disruption; Stage III includes lunotriquetral dissociation; and Stage IV culminates in volar lunate dislocation, the most unstable form requiring urgent closed or open reduction and fixation to restore alignment. Isolated volar lunate dislocations, comprising about 50% of all carpal dislocations, result from similar mechanisms and demand immediate surgical intervention to avoid compression and long-term . Diagnosis of carpal fractures and dislocations relies on clinical findings and advanced ; tenderness in the anatomic snuffbox has a of over 90% for scaphoid fractures, prompting immobilization even if initial radiographs are negative. For occult fractures, MRI is the modality of choice, detecting up to 100% of cases with edema signaling injury, while aids in precise classification of orientation and for surgical planning.

Disorders and conditions

The carpal bones are susceptible to various non-traumatic disorders that can lead to pain, instability, and functional impairment of the . These conditions often involve , cystic formations, or inflammatory processes affecting the bone structure or surrounding tissues. Early diagnosis through imaging and clinical evaluation is crucial for managing progression and preserving function. , also known as lunate malacia, is a progressive form of specifically affecting the lunate bone, typically occurring in young adults aged 20 to 40 years. It results from disrupted blood supply to the lunate, leading to bone ischemia, sclerosis, fragmentation, and eventual collapse. Risk factors include ulnar negative variance, present in approximately 75% of cases, which increases load on the lunate due to uneven distribution of force across the radiocarpal joint. The condition is classified using the Lichtman staging system, which guides treatment based on radiographic and MRI findings: Stage I shows no visible changes on X-rays but abnormal signal intensity on MRI indicating early necrosis; Stage II features lunate sclerosis without architectural disruption; Stage IIIA involves lunate fragmentation without carpal collapse; Stage IIIB includes fragmentation with collapse and proximal migration of the capitate; and Stage IV demonstrates secondary degenerative in the radiocarpal and midcarpal joints. Preiser's disease represents a rare idiopathic avascular necrosis of the scaphoid bone, occurring without associated fracture or significant trauma history. It primarily affects the proximal pole of the scaphoid due to its retrograde blood supply vulnerability, resulting in progressive sclerosis, fragmentation, and potential carpal if untreated. The condition is more common in males and often presents with insidious wrist pain, swelling, and reduced , diagnosed via MRI showing and in early stages. Ganglion cysts are benign, fluid-filled sacs arising from the or sheaths around the carpal bones, representing the most common mass. Dorsal ganglia, comprising 60-70% of cases, typically originate near the and present as painless, fluctuant masses on the back of the , potentially causing discomfort with extension. Volar ganglia, less frequent, arise from the radioscaphoid or adjacent to the , risking compression of nearby neurovascular structures and leading to symptoms like or ischemia if large. Carpal tunnel syndrome involves compression of the within the , a fibro-osseous canal formed by the carpal bones and roofed by the transverse carpal . Although primarily a , the carpal bones contribute to the tunnel's rigid boundaries, exacerbating pressure on the nerve from synovial proliferation or anatomical narrowing. Symptoms include nocturnal pain, in the thumb, , and fingers, and thenar muscle weakness, with carpal involvement confirmed by electrodiagnostic studies showing conduction delays. Rheumatoid arthritis frequently targets the through chronic , leading to erosion of the carpal bones and . In advanced cases, at the distal radioulnar progresses to caput ulnae syndrome, characterized by dorsal of the ulnar head, prominent , and extensor due to . This results in ulnar deviation, painful pronation-supination, and potential carpal translocation, with radiographic evidence of space narrowing and erosions.

Comparative anatomy

In mammals

In mammals, the number and structure of carpal bones exhibit significant variation across species, reflecting locomotor specializations. Primates, including non-human forms such as monkeys and apes, typically possess eight carpal bones arranged in two rows of four, mirroring the human configuration with distinct scaphoid, lunate, triquetrum, pisiform in the proximal row, and trapezium, trapezoid, capitate, hamate in the distal row. In contrast, horses (Equus caballus) have eight carpal bones: a proximal row consisting of the radial, intermediate (central), ulnar, and accessory bones, and a distal row of four numbered bones (I–IV), where fusions such as between the radial and intermediate carpal can occur congenitally or post-traumatically. Dogs (Canis lupus familiaris) generally feature seven carpal bones due to the fusion of the radial (scaphoid equivalent) and intermediate (lunate equivalent) into a single scapholunate bone, with the pisiform (accessory carpal) being particularly prominent and elongated to support tendon redirection during locomotion. These structural variations underpin key adaptations for diverse mammalian lifestyles. In (even-toed ungulates such as and deer), the proximal carpal row is often reduced—e.g., to six bones in bovines through fusions—facilitating a more rigid, columnar limb configuration that minimizes lateral deviation and enhances speed and stability during galloping to evade predators. Bats (Chiroptera), the only mammals capable of powered flight, exhibit elongated carpal bones integrated into an extended , where the proximal carpals (including scaphoid and lunate homologs) provide flexible yet robust anchorage for the wing membrane (), enabling precise aerodynamic control during hovering and maneuvering. Homologically, the scaphoid and lunate in mammals derive from a composite embryonic cartilage representing remnants of the reptilian radiale and intermedium, respectively, with developmental fusion often blurring strict boundaries and contributing to the reduced count in derived species like carnivores. In veterinary medicine, conditions affecting these bones highlight clinical parallels to human pathologies.

Evolutionary aspects

The carpal bones trace their origins to the Late Devonian period, when early tetrapods such as Acanthostega and Ichthyostega possessed a primitive wrist composed of 7-9 radials, which were segmented elements homologous to the distal radials of sarcopterygian fish fins and served as precursors to the ossified carpals of later forms. These radials supported polydactylous manus with up to eight digits, enabling initial weight-bearing during the fin-to-limb transition on terrestrial substrates. From amphibians to reptiles, evolutionary reduction occurred, streamlining the carpus to 6-8 elements, including proximal radials, intermedium, ulnare, and distal series, to optimize stability and propulsion in more fully terrestrial locomotion. In the lineage leading to mammals, the proximal carpal row evolved into a tripartite structure—comprising radiale (scaphoid), intermedium (lunate), and ulnare (triquetrum)—inherited from non-mammalian ancestors, with progressive and articulation refinements enhancing joint . A notable innovation was the pisiform, which originated as a within the of the m. flexor carpi ulnaris, becoming incorporated into the carpal row to stabilize the ulnar side and support enhanced grip strength during early mammalian foraging. Fossil evidence from therapsids, such as cynodonts from the Permian (~250 million years ago), reveals the separation of the lunate and scaphoid as distinct ossifications, diverging from the fused centrale-intermedium of earlier synapsids and allowing greater midcarpal flexibility. This separation is evident in specimens like , marking a phylogenetic shift toward more versatile mechanics in proto-mammalian forms. Functional adaptations in carpal transitioned from load distribution in quadrupedal among early tetrapods and reptiles to specialized in , where increased intercarpal mobility—driven by elongated scaphoid and triquetrum—facilitated precise grasping and arboreal suspension. This primate-specific diversification, evident from Eocene fossils like Adapis, prioritized dexterity over rigidity, underscoring the carpals' role in behavioral innovation.

Nomenclature

Etymology

The term "carpal" originates from the karpos (καρπός), meaning "," which entered Latin as carpus and later formed the Modern Latin carpalis to describe structures pertaining to the wrist region. The eight carpal bones received their names primarily during the , reflecting their distinctive shapes in Latin or Greek terms. In the proximal row, the scaphoid derives from the Greek skaphē (σκάφη), meaning "," due to its elongated, boat-like form; the lunate takes its name from the Latin luna, "," for its crescent or moon-shaped appearance; the triquetrum comes from the Latin triquetrus, combining tri- ("three") with a reference to its pyramidal or three-cornered structure; and the pisiform is from the Latin pisum (via Greek pison), meaning "," alluding to its small, pea-like nodule. In the distal row, the trapezium stems from the trapeza (τράπεζα), "table," describing its table-like base; the trapezoid combines trapez- (from the same root as trapezium) with Greek eidos ("form" or "shape"), indicating its resemblance to a or irregular ; the capitate arises from the Latin caput, "head," for its rounded, head-like prominence; and the hamate derives from the Latin hamatus, "hooked," referring to the prominent hook-like process on its palmar surface. The carpal bones were recognized in ancient texts, with Aristotle referencing the wrist as a joint in his History of Animals, connecting the hand to the forearm. Their detailed nomenclature and anatomy were formalized during the Renaissance by Andreas Vesalius in his seminal 1543 work De humani corporis fabrica, which provided precise illustrations and descriptions of the wrist's osseous components.

Historical naming

The nomenclature of the carpal bones has evolved significantly over centuries, reflecting advances in anatomical observation and standardization efforts. In antiquity, the Greek physician (c. 129–c. 216 ) was among the first to systematically describe the , recognizing eight distinct carpal bones without a formal numbering or naming system but using descriptive terms based on shape, observed in dissections primarily of animal models. During the , advanced carpal nomenclature in his seminal work De humani corporis fabrica (1543), where he provided detailed illustrations of the eight carpal bones and introduced early Latin designations, numbering them from 1 to 8 starting proximally from the radial side. Vesalius named the scaphoid as os naviculare (boat-shaped bone) and the capitate as os capitatum (head-shaped bone), drawing on morphological resemblances to establish a more precise terminological foundation that influenced subsequent anatomists. The marked a push toward uniformity with the adoption of the Nomina Anatomica (BNA) in 1895 by the German Anatomical Society, which standardized Latin terms for the carpal bones to resolve proliferating synonyms across . Under the BNA, bones received official designations such as os triquetrum for the triquetrum (three-cornered bone), emphasizing geometric features while aiming for international consistency in and texts. In the , the Federative Committee on Anatomical Terminology (FCAT) introduced the () in 1998, revised and updated in 2019, shifting toward a blend of Latin and English hybrids for broader while retaining classical roots. The classifies all eight bones as carpals, including the pisiform as os pisiforme, though ongoing debates persist regarding its status as a true carpal versus a embedded in the flexor carpi ulnaris tendon, due to its developmental and positional uniqueness. Historical variations in naming persisted in older texts, with the capitate often termed os magnum (great bone) to highlight its size as the largest carpal, and the hamate referred to as os cuneiform (wedge-shaped) or os unciforme (hooked), reflecting alternative emphases on form that predate standardized systems.

References

  1. [1]
    Anatomy, Shoulder and Upper Limb, Hand Carpal Bones - NCBI - NIH
    The carpal bones are bones of the wrist that connect the distal aspects of the radial and ulnar bones of the forearm to the bases of the 5 metacarpal bones ...
  2. [2]
    Bones of the Upper Limb – Anatomy & Physiology - UH Pressbooks
    The eight carpal bones form the base of the hand. These are arranged into proximal and distal rows of four bones each. The metacarpal bones form the palm of the ...
  3. [3]
    Embryology, Hand - StatPearls - NCBI Bookshelf - NIH
    Aug 14, 2023 · Fetal hand morphogenesis, which occurs between 6 and 14 weeks of gestation, can be described in three distinct phases of development.
  4. [4]
    Time of appearance of ossification centers in carpal bones
    The main finding in this study is that carpal bones start to ossify in the first year of life, starting approximately at the age of 6 months. This ossification ...
  5. [5]
    Carpal tunnel syndrome - Symptoms and causes - Mayo Clinic
    Feb 6, 2024 · Carpal tunnel syndrome is one of the most common hand conditions. It is caused by pressure on the median nerve in the carpal tunnel of the wrist.Carpal tunnel release · Carpal tunnel exercises: Can... · Diagnosis and treatment
  6. [6]
    Characterization of a Pre-Clinical Mini-Pig Model of Scaphoid Non ...
    Jun 16, 2015 · Mapping Scaphoid Bone Morphology and Vascular Supply. Radiologic images and 3D representations (Figure 1) of the adult human hand (Figure 1A ...<|separator|>
  7. [7]
    The association of lunate morphology with the development of wrist ...
    Apr 30, 2025 · Vranješ M, Bjelobrk M, Lošonci V, Lažetić F. Lunate bone morphology and wrist arthrosis: A radiographic study of the type 2 lunate. SN Compr ...
  8. [8]
    The morphologic and morphometric features of the triquetrum
    Nov 15, 2022 · The aim of this study was to describe the morphology of triquetrum using anatomical landmarks, evaluate its morphometric features, and determine its vascular ...
  9. [9]
    Triquetral Fractures Overview - PMC - NIH
    Jan 23, 2021 · Anatomy. The triquetrum has three facets for articulation with adjacent carpal bones. The flat facet on the ulnar surface of the triquetrum ...
  10. [10]
    [PDF] VARIATION IN PISIFORM MORPHOLOGY - PSU-ETD
    This chapter provides a review of bone and growth plate development, as well as crucial background information about wrist anatomy for subsequent chapters. Page ...
  11. [11]
    Bone morphological changes of the trapezium and first metacarpal ...
    1. Trapezium Morphology Measures: Trapezium height and width were defined using the dimensions of a radial-ulnar section which spanned the bone radial to ulnar ...
  12. [12]
    Bilateral bipartite trapezoid: a rare anatomical variant - PMC - NIH
    Six new examples of the bipartite trapezoid bone: morphology, significant population variation, and an examination of pre-existing criteria to identify ...
  13. [13]
    [PDF] The Chevron Capitate-Trapezoid Joint: A Case Report
    Posteroanterior radiographs (Figures 1A and 1B) and a T2 coronal MRI (Figure 2) showed a chevron-shaped, capitate-trapezoid joint and capitate diameter of 22 mm.
  14. [14]
    Hamate-pisiform coalition: morphology, clinical significance, and a ...
    Hamate-pisiform coalition is characterized by the abnormal union of the pisiform bone and hamulus of the hamate.
  15. [15]
    Joints and Ligaments Tables - Medical Gross Anatomy
    the joint which connects the proximal and distal rows of carpal bones, a series of synovial plane joints; small ranges of motion are permitted between carpal ...
  16. [16]
    Joints and Ligaments of the Upper Limb - UAMS College of Medicine
    Joints and Ligaments of the Upper Limb ; midcarpal joint, synovial, gliding, connects proximal & distal rows of carpal bones ; oblique cord, syndesmosis, connects ...
  17. [17]
    9.4 Synovial Joints - Anatomy and Physiology 2e | OpenStax
    Apr 20, 2022 · Saddle joints are functionally classified as biaxial joints. The primary example is the first carpometacarpal joint, between the trapezium (a ...Types Of Synovial Joints · Saddle Joint · Aging And The
  18. [18]
    Anatomy, Shoulder and Upper Limb, Wrist Joint - StatPearls - NCBI
    The wrist joint also referred to as the radiocarpal joint is a condyloid synovial joint of the distal upper limb that connects and serves as a transition point ...Structure And Function · Surgical Considerations · Clinical Significance
  19. [19]
    Midcarpal joint: anatomy, function, movements - Kenhub
    The midcarpal joint is a series of synovial gliding joints between the proximal and distal rows of carpal bones. Like any other synovial joint, the adjacent ...
  20. [20]
    Radiocarpal (wrist) joint: Bones, ligaments, movements - Kenhub
    The outer portion of the capsule is composed of fibrous connective tissue which provides structural support to the joint, while the inner layer is composed of ...
  21. [21]
    Carpal tendinous sheaths: Anatomy and clinical relevance - Kenhub
    The common flexor synovial sheath contains the tendons of flexor digitorum superficialis and flexor digitorum profundus. It is also known as the ulnar bursa, as ...
  22. [22]
    The Carpal Tunnel - Borders - Contents - TeachMeAnatomy
    Nov 10, 2020 · The tendon of flexor pollicis longus is surrounded by its own synovial sheath. These sheaths allow free movement of the tendons. Sometimes ...
  23. [23]
    Wrist Joint Anatomy - Medscape Reference
    Mar 11, 2025 · The joints of the wrist are surrounded by a fibrous capsule ... joint capsule and provides dorsal stabilization to the wrist throughout its range ...Missing: synovial layers
  24. [24]
    Wrist and Hand - Physiopedia
    ... ligaments of the wrist can be divided into extrinsic and intrinsic ligaments. The extrinsic ligaments connect the carpal bones to the radius or metacarpals.Wrist and Hand Examination · Hand Osteoarthritis · Edit
  25. [25]
    Wrist ligaments | Radiology Reference Article - Radiopaedia.org
    Jan 5, 2024 · The intrinsic and extrinsic wrist ligaments play a vital role in the stability of the wrist joint. There are numerous ligaments but included below are the most ...
  26. [26]
    Wrist Ligaments & Biomechanics - Hand - Orthobullets
    Mar 7, 2024 · The ligaments of the wrist include extrinsic ligaments, intrinsic ligaments, characteristics, space of Poirier between the volar radioscaphocapitate ligament ...
  27. [27]
    The Wrist Joint - TeachMeAnatomy
    The wrist joint (also known as the radiocarpal joint) is an articulation between the radius and the carpal bones of the hand.
  28. [28]
    Anatomy, Biomechanics, and Loads of the Wrist Joint - PMC
    Jan 27, 2022 · The wrist is composed of two rows of carpal bones: the proximal carpal row (PCR) includes from radial to ulnar the scaphoid, lunate, triquetrum ...
  29. [29]
    Anatomy of the Hand & Wrist: Bones, Muscles & Ligaments
    Jun 12, 2023 · Volar carpal ligaments: Ligaments that support and stabilize the bottom (palmar side) of your wrist. Dorsal radiocarpal ligaments: Ligaments ...
  30. [30]
    Pisohamate ligament - e-Anatomy - IMAIOS
    The pisohamate ligament is a volar ligament in the hand connecting the pisiform to the hamate, and is a prolongation of the Flexor carpi ulnaris tendon.
  31. [31]
    Narrative Review of Ligamentous Wrist Injuries - PMC
    The extrinsic carpal ligaments connect the forearm to the carpus and are contained inside the joint. They can be divided into palmar, dorsal, radiocarpal, and ...
  32. [32]
    High-resolution ultrasound of the extrinsic carpal ligaments - PMC
    The extrinsic ligaments originate from the carpal bones and pass out of the wrist to insert onto the radius or ulna. These features make them important for ...
  33. [33]
    The Incidence of Accessory Ossicles of the Wrist: A Radiographic ...
    The clinical significance of os styloideum stems from its relationship with carpal boss, which is a painful lump in the dorsum of the wrist. While os styloideum ...
  34. [34]
    Small but troublesome: accessory ossicles with clinical significance
    Accessory ossicles are inconstant, independent, and considered well formed bones, not arising from fractures or other diseases.Missing: prevalence | Show results with:prevalence
  35. [35]
    Bilateral Symptomatic Os Epilunatum: A Case Report - PMC - NIH
    The overall incidence of accessory ossicles in the wrist is estimated to be around 1.6%. In a review of 1,452 radiographs, Bogart did not find any case ...Missing: prevalence significance
  36. [36]
    Os centrale carpi | Radiology Reference Article | Radiopaedia.org
    Jul 25, 2017 · Os centrale carpi have been observed to cause symptoms such as intermittent pain, clicking and crepitus due to its mobility in the carpus.Missing: significance | Show results with:significance
  37. [37]
    Prevalence of Os Styloideum in National Hockey League Players
    Previous clinical and cadaveric studies estimated a general prevalence of up to 19% ( P < 0.001). For the 10 players who had their leading wrist scanned, 9 had ...
  38. [38]
    Ultrasonographic Diagnosis of an Os Styloideum in an Adolescent ...
    Nov 16, 2022 · The os styloideum is an accessory ossicle of the wrist located at the carpometacarpal junction dorsally in approximately 0.3%-2.3% of wrists [1] ...
  39. [39]
    [PDF] Hand and wrist : a kaleidoscopic view of accessory ossicles, variants ...
    Definition: irregular ossification of the pisiform bone at the age of 8 to 12. • Prevalence: often occurs in the carpal bones during development. The pisiform ...
  40. [40]
    The wizard of os: accessory ossicles from the spine and ... - EPOS™
    Os triangulare. Prevalence: 1% on right hand, 0.9% on left hand; one of the six most common wrist ossicles. Location: Just distal to the ulnar fovea, between ...
  41. [41]
    Os triangulare | Radiology Reference Article | Radiopaedia.org
    Oct 8, 2023 · The os triangulare (also known as the os intermedium antebrachii or os triquetrum secundarium) is an accessory ossicle located between the ulnar styloid, ...Missing: prevalence | Show results with:prevalence
  42. [42]
    Os epilunate | Radiology Reference Article | Radiopaedia.org
    Jul 25, 2017 · The os epilunate is an accessory ossicle of the wrist that lies at the dorsal surface of the lunocapitate articulation.
  43. [43]
    Prevalence of accessory carpal ossicles – a CT-based survey
    ... os hamuli proprium (n:18, 0.813%), os ulnostyloideum (n:16, 0.723%), os epilunatum (n:12, 0.542%) and os styloideum (n:11, 0.497%) respectively. The study ...
  44. [44]
  45. [45]
    Hamate Bone - an overview | ScienceDirect Topics
    The hook typically fuses to the hamate body between the ages of twelve and fifteen; failure to fuse leads to a separate ossicle termed the os hamuli proprium.18 ...
  46. [46]
    Hook of the Hamate: The Spectrum of Often Missed Pathologic ...
    Most cases of os hamuli proprium are discovered incidentally and are often in the differential diagnosis of hamate fractures (Fig. 7A). Differentiating an ...
  47. [47]
    Imaging Report With Radiologic Criteria for Differentiation: Avulsion ...
    Oct 6, 2025 · Accessory ossicles of the wrist are uncommon bone variants that can be mistaken for avulsion fractures, potentially leading to unnecessary ...
  48. [48]
  49. [49]
    Homeobox genes d11–d13 and a13 control mouse autopod cortical ...
    ... Hoxd13 showed a severely disturbed differentiation process of the distal skeletal elements. ... identity as long bones and are transformed into carpal-like bones.
  50. [50]
    Development and morphogenesis of human wrist joint during ...
    Mar 19, 2012 · The development of the human wrist joint has been studied widely, with the main focus on carpal chondrogenesis, ligaments and triangular fibrocartilage.
  51. [51]
    Musculoskeletal System - Bone Development Timeline - Embryology
    Feb 9, 2020 · Carpal Bones. Data from a wrist study of 57 human embryonic (stages 17-23) and fetal (9-14 weeks). stage 17 - undifferentiated ...Introduction · Historic Limb Data · Carpal Bones · References
  52. [52]
    Carpal Coalitions and Metacarpal Synostoses: A Review - PMC
    Sep 1, 2016 · Embryologically, carpal coalitions arise from a failure of separation of the cartilaginous interzone of adjacent carpal bones. The interzone is ...
  53. [53]
    Carpal Synostosis (coalition) - SpringerLink
    Aug 21, 2014 · Congenital carpal fusion is due to failure of segmentation during embryonic limb development with subsequent failure of joint formation.
  54. [54]
    Pisiform Bone - an overview | ScienceDirect Topics
    Supernumerary, or accessory, carpal bones occasionally form in the wrist. Several of these variants occur frequently enough to warrant specific names: the ...
  55. [55]
    Ossification centers of the wrist | Radiology Reference Article
    Oct 20, 2020 · Ossification of the carpal bones · capitate: 1-3 months · hamate: 2-4 months · triquetrum: 2-3 years · lunate: 2-4 years · scaphoid: 4-6 years ...
  56. [56]
    Pisiform Bone - an overview | ScienceDirect Topics
    The pisiform bone is a sesamoid bone that is enveloped within the flexor carpi ulnaris tendon and that is superficial to the triquetrum on the volar surface. It ...
  57. [57]
    Pisiform | Radiology Reference Article - Radiopaedia.org
    Jul 4, 2025 · Often the last carpal bone to ossify, the pisiform has one ossification center that ossifies in the ninth to twelfth year, later in males.
  58. [58]
    Bone age assessment | Radiology Reference Article
    Apr 10, 2013 · Bone age assessment is used to radiologically assess the biological and structural maturity of immature patients from their hand and wrist x-ray appearances.
  59. [59]
    Evaluation of Bone Age in Children: A Mini-Review - Frontiers
    Mar 11, 2021 · Hand and wrist X-rays are considered as an important indicator of children's biological age. Nowadays, many methods are available to evaluate ...
  60. [60]
    Scapholunate Ligament - Physiopedia
    The carpal bones in the distal row distribute axial loads relatively evenly, 50% of the axial load is transmitted through the radioscaphoid joint and 35 ...<|control11|><|separator|>
  61. [61]
    Triangular Fibrocartilage Complex (TFCC) Tear - DynaMed
    Oct 16, 2023 · TFCC functions to stabilize the distal radioulnar joint, and provide shock absorption between the ulna and the carpus · disruption of the complex ...
  62. [62]
    Carpal dynamic stability mechanisms. Experimental study - Elsevier
    The motor muscles of the wrist exert a dynamic effect on the carpal bones, increasing the stability of the joint.
  63. [63]
    The application of computer-based quantitative analysis to explain ...
    The stable central column theory of carpal biomechanics provides a unified concept for explaining carpal mechanics and understanding ways to address ...Figure 1 · Wrist Biomechanics · Adequate Flexion And...
  64. [64]
    Wrist Bone Motion during Flexion-Extension and Radial-Ulnar ... - NIH
    Sep 20, 2022 · During the RUD, carpal bones in the distal row rotated along with wrist movement while the scaphoid, lunate, and triquetrum primarily flexed and extended.
  65. [65]
    Study of wrist motion in flexion and extension - PubMed
    During maximum flexion, the motion is 40% radio--carpal and 60% mid carpal. During maximum extension, the motion is 66.5% radiocarpal and 33.5% mid carpal.
  66. [66]
    Kinematics of the midcarpal and radiocarpal joints in radioulnar ...
    Radial and ulnar deviation occurred primarily in the midcarpal joint. Midcarpal motion accounted for 60% of radial deviation and 86% of ulnar deviation.
  67. [67]
    Non-invasive 3D fluoroscopic/CT imaging of the Wrist
    In particular, the dart throwing motion (DTM) is important because it is characterized by proximal row kinematics unique to humans and displays multiple planes ...
  68. [68]
    A study on the measurement of wrist motion range using ... - PubMed
    Results: Normal values for wrist ROM are 73 degrees of flexion, 71 degrees of extension, 19 degrees of radial deviation, 33 degrees of ulnar deviation, 140 ...
  69. [69]
    Kinematics of the midcarpal and radiocarpal joint in flexion and ...
    To apply carpal kinematic analysis using noninvasive medical imaging to investigate the midcarpal and radiocarpal contributions to wrist flexion and extension.
  70. [70]
    WRIST JOINT COMPLEX - OUHSC.edu
    Most wrist extension occurs around the midcarpal joint's lateral axis. Most wrist flexion occurs around the radiocarpal joint's lateral axis.
  71. [71]
    Anatomy, Shoulder and Upper Limb, Forearm Flexor Carpi Ulnaris ...
    Jan 30, 2024 · The FCU primarily flexes the wrist and is the most powerful wrist flexor. The muscle also assists in wrist adduction and elbow and forearm flexion.
  72. [72]
    Anatomy, Shoulder and Upper Limb, Wrist Extensor Muscles - NCBI
    The wrist extensor muscles comprise a significant component of the posterior forearm musculature. These muscles generally originate on or near the lateral ...
  73. [73]
  74. [74]
    Mobilization With Movement as an Adjunct Intervention in a Patient ...
    During each intervention session, MWM involving a sustained radial glide of the proximal row of carpal bones resulted in in- creased range of pain-free ...<|separator|>
  75. [75]
    A four-dimensional-CT study of in vivo scapholunate rotation axes
    Feb 27, 2019 · During flexion–extension motions, the scaphoid rotates 38° (SD 0.6°) relative to the lunate; the rotation axis intersects the dorsal ridge of ...<|separator|>
  76. [76]
    [PDF] Intra-Rater Reliability of the Mobil-Aider® Device
    Oct 18, 2021 · Similar to the GHJ, the. RCJ requires accessory glide to achieve full range of motion. As the convex proximal row of carpal bones moves upon the ...
  77. [77]
    Wrist Instability - StatPearls - NCBI Bookshelf - NIH
    May 22, 2023 · The scapholunate ligamentous complex and dorsal intercarpal ligaments stabilize this joint from distraction, torsion, and translation. Wrist ...
  78. [78]
    Proprioceptive Neuromuscular Facilitation Protocol for Thumb ...
    The unique biomechanics of the CMC joint allows opposition to the digits and performance of resistive pinch as needed for daily tasks such as buttoning ...
  79. [79]
    [PDF] dart-throwers-consensus-2007.pdf - Dr Scott Wolfe
    The dart-throwing motion (DTM) plane can be defined as a plane in which wrist functional oblique motion occurs, specifically from radial extension to ulnar ...
  80. [80]
    The Dart-Throwing Motion of the Wrist: Is It Unique to Humans? - PMC
    Kinematic analysis has shown a near-stationary proximal carpal row during the dart-thrower's motion, which is believed to provide a stable platform for the ...Missing: fluoroscopy | Show results with:fluoroscopy
  81. [81]
    Scaphoid Fracture - Hand - Orthobullets
    Jul 25, 2025 · most common mechanism of injury is axial load across a hyper-dorsiflexed, pronated and ulnarly-deviated wrist ... link between proximal and distal ...Missing: transmission | Show results with:transmission
  82. [82]
    Lunate Dislocation (Perilunate dissociation) - Hand - Orthobullets
    Oct 3, 2025 · Mayfield Classification ; Stage I. Scapholunate dissociation ; Stage II. + lunocapitate disruption ; Stage III. + lunotriquetral disruption, " ...
  83. [83]
    Diagnosis and Management of Scaphoid Fractures - AAFP
    Sep 1, 2004 · Anatomic snuffbox tenderness is a highly sensitive test for scaphoid fracture, whereas scaphoid compression pain and tenderness of the scaphoid tubercle tend ...
  84. [84]
    Scaphoid Fracture: Symptoms, Causes & Treatment - Cleveland Clinic
    More than 70% of scaphoid fractures occur in the waist. Proximal pole: The end of your scaphoid closest to your forearm (pointing in toward your body).
  85. [85]
    Scaphoid Fractures - TeachMeSurgery
    Jun 3, 2021 · Avascular necrosis is common complication of a scaphoid fracture (in around 30% of cases), with its risk increasing the more proximal the ...
  86. [86]
    Hook of Hamate Fracture - Hand - Orthobullets
    Apr 13, 2025 · Hook of hamate fractures are rare, often missed, injuries generally as a result of a direct blow to the hamate bone most commonly seen in ...
  87. [87]
    Triquetral fracture | Radiology Reference Article - Radiopaedia.org
    Aug 18, 2025 · Triquetral fractures are carpal bone fractures generally occurring on the dorsal surface of the triquetrum.Missing: elevation | Show results with:elevation
  88. [88]
    Mayfield classification of carpal instability (perilunate instability)
    Oct 16, 2025 · Mayfield classification of carpal instability, also known as perilunate instability classification (carpal dislocations), describes carpal ligament injuries.
  89. [89]
    Carpal dislocations: pathomechanics and progressive perilunar ...
    Lunate dislocations (stage IV PLI) had the highest degree of carpal instability. Radial styloid fractures were produced in seven as a result of avulsion.
  90. [90]
    Scaphoid fracture | Radiology Reference Article | Radiopaedia.org
    Oct 5, 2025 · Epidemiology. Scaphoid fractures account for 70-80% of all carpal bone fractures 1. Although they occur essentially at any age, adolescents ...
  91. [91]
    Different evolutionary pathways underlie the morphology of wrist ...
    Oct 23, 2013 · The evolutionary morphological changes in some wrist bones are consistent with similarities in locomotor behaviour shared across taxa (scaphoid, ...
  92. [92]
    The Equine Carpus | Veterian Key
    May 27, 2016 · The equine carpus is composed of three main articulations: the antebrachiocarpal joint; the middle carpal joint, and the carpometacarpal joint.
  93. [93]
    Arthrographic description of the canine carpal joint cavities and its ...
    Mar 10, 2024 · The canine carpus with its seven carpal bones, arranged in two rows and held together by a multitude of carpal ligaments, is a complex joint ...
  94. [94]
    The evolution of the unguligrade manus in artiodactyls
    Artiodactyls show a step-wise morphological transition from groups with a primitive and pentadactyl limb configuration to groups with more derived and didactyl ...Missing: carpal | Show results with:carpal
  95. [95]
    Bat flight: aerodynamics, kinematics and flight morphology
    Mar 1, 2015 · The bat forelimb has elongated bones relative to non-flying mammals, resulting in a large wing area (e.g. Simmons, 1994; Swartz, 1997 ...Missing: carpal | Show results with:carpal
  96. [96]
    New Developmental Evidence Clarifies the Evolution of Wrist Bones ...
    Sep 30, 2014 · We support the use of the term “scapholunare” for the bone that develops from the embryonic cartilage that is composite of radiale+intermedium, ...Missing: lunate | Show results with:lunate
  97. [97]
    Navicular Disease: Possible New Treatment Option from Human ...
    Apr 9, 2011 · Surgeons perform this core decompression surgery on humans who are experiencing pain caused by increased pressure within a dying bone; the ...Missing: Kienböck's | Show results with:Kienböck's
  98. [98]
    The phantoms of a high-seven - or - why do our thumbs stick out?
    Sep 15, 2015 · The earliest tetrapods had hands and feet with up to eight digits but this number was subsequently reduced during evolution.
  99. [99]
    [PDF] Evolution and identity of synapsid carpal bones
    Nov 6, 2020 · Homology of the reptilian and mammalian carpals after Ge- genbaur (1864). Canonical nomenclature. Mammalian nomenclature radiale scaphoideum.
  100. [100]
    Evolution and identity of synapsid carpal bones - ResearchGate
    Nov 6, 2020 · PDF | On Nov 6, 2020, Susanna Kümmell and others published Evolution and identity of synapsid carpal bones | Find, read and cite all the ...<|control11|><|separator|>
  101. [101]
    Carpal - Etymology, Origin & Meaning
    Originating from Modern Latin carpalis, meaning "of or pertaining to the wrist," derived from carpus "wrist," the term relates to the wrist area and its ...
  102. [102]
    Musculoskeletal etymology: What's in a name? - PMC
    The carpal (carpus means wrist) bones are named more imaginatively based on their shapes- 'scaphoid' (boat-shaped, Fig. 6a), 'lunate' (moon-shaped, Fig. 6b), ' ...
  103. [103]
    The Bones of the Wrist (Ossa Carpalia) - Anatomy Standard
    Oct 4, 2020 · Os scaphoideum is the greek-origin word meaning the boat-like bone (gr. skáphē – the boat). The shape of the bone corresponds well to the name – ...
  104. [104]
    8 Handy Tips to Remember the Bones of the Wrist | Anatomy Slices
    it means three corners; “tri” meaning three, “quetra” meaning corner. ...
  105. [105]
    Carpal Bones Mnemonic - Picmonic
    The etymology for this bone is derived from latin, meaning "3-cornered." It is also the second most commonly fractured carpal bone. Pisiform Pizza-farm. This ...
  106. [106]
    Aristotle, History of Animals - ToposText
    Then come the shoulder-bones, or blade-bones, and the arm-bones connected with these, and the bones in the hands connected with the bones of the arms. With ...
  107. [107]
    The carpus and bones of the hand and the eight bones of the wrist ...
    Download this stock image: The carpus and bones of the hand and the eight bones of the wrist shown from various aspects from De humani corporis fabrica ...
  108. [108]
    [History of the carpus and carpal instability] - PubMed
    At first, the carpal bones were simply numbered from one to eight. Only in the 17th century did Lyser of Leipzig suggest naming the individual bones, something ...
  109. [109]
    1 The Story of Hand Anatomy | Musculoskeletal Key
    Jan 25, 2021 · In discussing the upper limbs in the Fabrica, Vesalius uses the terms “radius” and “ulna” for the two forearm bones. The teleology of the ...<|separator|>
  110. [110]
    Anatomical Terminology, Then and Now1 - Karger Publishers
    It became common practice to number the carpal bones from lateral to medial, first in the proximal and then in the distal row. This procedure was adopted by ...
  111. [111]
    [The pisiform bone: sesamoid or carpal bone?] - PubMed
    The pisiform bone is unusual, located in front of other carpal bones. Theories suggest it could be a sesamoid or a carpal bone, with the latter being the most ...
  112. [112]
    Capitate | Radiology Reference Article | Radiopaedia.org
    Mar 30, 2023 · The capitate, also known as the os magnum, is the largest of the carpal bones and sits at the center of the distal carpal row.Missing: cuneiform | Show results with:cuneiform