Fact-checked by Grok 2 weeks ago

Dewlap

A dewlap is a pendulous fold of loose that hangs beneath the or of certain vertebrates, often serving as a specialized anatomical feature adapted to specific physiological or behavioral needs. In mammals, dewlaps are prominent in large ungulates such as , , and elands, where they typically occur in males of species exceeding 400 kg in body mass and are associated with through heat dissipation, as evidenced by infrared thermography showing elevated surface temperatures in these structures. Unlike hypotheses of or predator deterrence, which lack strong empirical support in these taxa, the thermoregulatory role aligns with the dewlaps' vascularization and exposure in hot environments. In reptiles, particularly lizards, the dewlap is an extensible flap supported by the hyoid apparatus, displaying vibrant colors and patterns for , including mate attraction, territorial defense, and species recognition during dynamic head-bobbing displays. These lizard dewlaps exhibit remarkable diversity, with over 65 unique combinations of six patterns and 13 colors observed across species, often converging independently rather than reflecting phylogenetic relationships. Dewlaps also appear in birds, such as the wattled structures in turkeys and roosters, where they may aid in or signaling, though less extensively studied than in mammals and reptiles. In veterinary contexts, dewlaps in animals like rabbits can become prone to under moist conditions, highlighting their role in everyday animal . Overall, while dewlaps vary in form and function across taxa, they represent an evolutionary emphasizing display, cooling, or structural utility in diverse ecological niches.

Overview

Definition

A dewlap is a longitudinal flap or pendulous fold of loose that hangs beneath the lower , , or of certain vertebrates. This is typically composed of layers supported by underlying , which provides flexibility and support. In some species, additional components such as subcutaneous fat deposits contribute to its bulk and pendulous nature, while minimal muscle fibers may enable limited movement or extension. Variations in dewlap and form occur across vertebrates, with the often being extensible or inflatable due to properties of and vascular elements within the connective matrix. For instance, in reptiles, it manifests as a foldable skin flap, whereas in mammals, it appears as a more static, hanging fold enriched with loose areolar . The dewlap is distinct from similar integumentary structures like wattles, which are paired, fleshy caruncles attached laterally to the head near the in , and combs, which form raised, vascular on the crown of the head. Its defining feature is the specifically pendulous, midline positioning along the throat or neck, setting it apart from these more cranial or bilateral appendages. Historical veterinary anatomy references, such as those in early texts on bovine , describe the dewlap as a characteristic loose under the in , while reptilian descriptions note it as a throat-based extension in .

The term "dewlap" entered the in the Middle English period, with its earliest recorded use dating to 1398. It derives from the compound "dewlappe," combining "dew" (likely through ) and "lappe" or "lap," which stems from læppa, meaning a loose hanging piece, fold, or flap of skin. The "dew" component may represent an alteration of an earlier, unattested element—possibly dunn ("down" or soft hair)—as the association with moisture does not align directly with the anatomical feature; scholars note this as a common case of reshaping for phonetic or associative familiarity. Initially, "dewlap" described the pendulous fold of under the of bovine animals like oxen and , reflecting its origins in medieval agricultural terminology. By the , following the introduction of turkeys from the to around 1520–1530, the word extended to the similar fleshy on these birds. Its application to human anatomy emerged in the late 1590s, denoting sagging on the associated with age or excess. In literary usage, the adjective "dew-lapped" gained prominence in the works of , who employed it to evoke bovine or aged characteristics—for instance, in (c. 1595), Puck refers to an old woman's "withered dewlap," and in (c. 1611), Gonzalo questions the existence of "mountaineers dewlapped like bulls." These references highlight the term's metaphorical flexibility by the early . During the 19th and 20th centuries, zoological texts adopted "dewlap" for comparable structures in reptiles and , marking a shift from primarily mammalian and contexts to broader scientific .

Functions

Display and Communication

The dewlap plays a primary role in particularly in reptiles such as , where it is often larger and more colorful in males than in females, serving as an advertisement of genetic quality and physical condition. Males actively inflate or extend the dewlap during displays to signal their body size, health, and vigor, thereby attracting potential mates and influencing female choice. In territorial interactions, the dewlap functions as a visual deterrent, with individuals rapidly flashing or pulsing it to assert dominance and ward off rivals. This signaling is frequently combined with dynamic body movements, such as displays, to amplify the threat and facilitate the defense of resources like feeding or breeding areas. Dewlap characteristics, including distinct color patterns and extension sequences, aid in species recognition by providing unique visual cues that help individuals discriminate conspecifics from other species during mate selection, thereby reducing the risk of hybridization. Evolutionary analyses reveal that dewlap traits are shaped by pressures in , with morphological variations—such as increased brightness or size—linked to ecological factors like predation intensity; brighter dewlaps, for example, tend to evolve in habitats with reduced predator presence, balancing signaling efficacy against survival costs. Dewlaps enhance communication by integrating visual signals with complementary behaviors, such as postural adjustments or acoustic cues, to convey more complex messages in and reproductive contexts, thereby improving signal reliability and receiver response. In , such as turkeys and roosters, wattled dewlaps may serve similar roles in and communication, though these functions are less extensively studied.

Thermoregulation and Other Physiological Roles

The contributes to by expanding the effective surface area for convective and radiative heat loss, which is particularly advantageous in large-bodied animals facing challenges from a low surface area-to-volume ratio. Its vascular structure enables to increase blood flow near surface, promoting heat dissipation through and conduction in hot conditions, while can conserve heat in cooler climates. This mechanism aids in maintaining core body temperature across environmental fluctuations. In mammals, particularly ungulates, studies demonstrate that dewlap presence and size positively correlate with tropical habitat occupancy and body mass exceeding 400 kg, supporting a role in mitigating via enhanced evaporative cooling without reliance on behavioral adjustments. Phylogenetic analyses indicate that drives dewlap more than alternative functions like , as structures appear in species from warmer biomes regardless of levels. Beyond , the dewlap acts as a fat storage site in certain , accumulating subcutaneous reserves that serve as an energy buffer during nutritional scarcity and contribute to overall body condition assessment. It also provides structural flexibility, allowing greater range of motion and reducing strain during or by accommodating displacement. However, the pendulous nature of the dewlap predisposes it to retention from environmental or salivary , increasing vulnerability to bacterial infections such as moist , which can lead to localized if untreated. In mammals, evolutionarily, dewlaps have developed prominently in lineages inhabiting arid or regions, with size variations primarily tied to climatic demands rather than or reproductive pressures alone.

Occurrence in Mammals

Ruminants and Ungulates

In ruminants and ungulates, dewlaps occur prominently in males of large species exceeding 400 kg body mass, such as wild examples including (Alces alces) and (Taurotragus oryx), where they aid through increased surface area for heat dissipation, as shown by elevated temperatures in infrared imaging. These structures are also well-developed in tropical-adapted domestic breeds of and , forming a distinctive pendulous fold of extending from the lower to the chest. Bos indicus () cattle breeds, such as the , exhibit particularly large dewlaps in mature bulls, serving as a key morphological trait for heat dissipation in arid conditions. Similarly, Boer goats display well-developed dewlaps, often more pronounced in males, which contribute to their breed's characteristic profile alongside long, drooping ears. These structures are absent or minimal in temperate breeds like the Holstein dairy cow, highlighting a clear distinction between Bos taurus and Bos indicus lineages. Anatomically, the dewlap consists of loose, folded with underlying subcutaneous deposits that provide and elasticity, allowing it to hang pendulously without restricting . This feature is typically more extensive in males and increases with age due to accumulation and skin laxity, varying by breed; for instance, cattle show exaggerated sizes compared to humpless African Bos taurus breeds. In agricultural contexts, has amplified dewlap size in lines for enhanced thermoregulatory efficiency in hot climates, while some programs aim to reduce it in crossbreeds to improve or reduce . Evolutionarily, the dewlap in Bos indicus ruminants originated in African and Asian lineages as an adaptation to arid and subtropical environments, where the increased surface area facilitates evaporative cooling through greater blood flow and activity. Across ungulates more broadly, dewlaps have evolved convergently in large species for similar thermoregulatory benefits, preserved through in wild populations and intensified by human-mediated selection in husbandry, aiding in regions with high temperatures and low . Practically, dewlaps play a role in identification during livestock shows and auctions, where size and conformation are judged as indicators of tropical heritage. However, they can pose veterinary challenges, such as abscess formation from trauma or , which may lead to localized infections requiring lancing and antibiotic treatment.

Lagomorphs and Other Mammals

In lagomorphs, the dewlap refers to a pendulous fold of loose beneath the and , most notably developed in female rabbits (Oryctolagus cuniculus) and to a lesser extent in hares (Lepus species). This structure serves as a influenced by , becoming more prominent during reproductive maturity to facilitate nest-building; females pluck from the dewlap to line nests for their , providing insulation and warmth. In males, dewlaps are typically smaller or absent, though some intact bucks may develop minor folds due to hormonal factors. While dewlaps are minimal and often inconspicuous in wild lagomorph populations—likely an adaptation to reduce vulnerability in natural environments—they are more exaggerated in breeds, such as Flemish Giants or Angoras, due to for aesthetic or genetic traits. These enlarged dewlaps can lead to health issues, including moist or "wet dewlap," where damp conditions cause bacterial infections ( species) in the skin folds, particularly if the rabbit's drinking habits or housing exposes the area to moisture. In humid habitats, such infections pose a greater to captive or escaped domestic lagomorphs, though wild individuals with slimmer dewlaps experience lower incidence. Beyond lagomorphs, dewlaps appear in various other mammals as loose serving structural or protective roles. In canids, such as the , the dewlap consists of pendulous skin around the neck and , enhancing mobility by allowing the skin to shift without restricting underlying muscle movement during hunting or tracking. This feature is breed-specific, prominent in scent hounds for accommodating extended head-lowered postures. In felids, mature male leopards (Panthera pardus) exhibit a well-developed dewlap of loose skin, which grows with age and may aid in displays or provide padding during territorial conflicts, though its exact function remains under study. Developmentally, lagomorph dewlaps enlarge post-puberty in response to reproductive hormones, with size varying by , , and spaying status—intact females often show the most pronounced growth, while can prevent excessive accumulation. In domestic breeds, hereditary factors amplify this trait, sometimes leading to welfare concerns like chronic irritation if not managed. For wild lagomorphs in humid regions, the subtler helps mitigate infection risks, supporting overall population health amid environmental stressors like moisture-related pathogens.

Occurrence in Reptiles

Lizards

In lizards, particularly in the infraorder Iguania, such as anoles (genus , family ) and the family , dewlaps are extensible flaps of skin located beneath the throat, prominently featured in species such as lizards (anoles) and the green iguana (Iguana iguana). These structures are supported by the hyoid apparatus, specifically the second ceratobranchial , which enables rapid extension through a lever-like mechanism involving the hyoid bones and associated musculature. In Anolis equestris, for instance, contraction of the M. ceratohyoideus pulls the ceratohyals posteriorly, pivoting the ceratobranchials forward to unfurl the dewlap in phases: initial protraction of the hyoid, followed by downward pivoting and skin expansion. Retraction occurs via antagonistic muscles, including the M. sternohyoideus and M. hyoglossus, which draw the hyoid back to its resting position, allowing quick pulsing motions. The dewlap's skin is thin and highly elastic, often covered in small scales and pigmented for vibrancy, with size, shape, and coloration varying markedly across species to reflect phylogenetic and ecological differences. In the green anole (), the male dewlap is typically pink to orange, spanning up to 7 cm when extended, while females usually lack a prominent dewlap or possess a reduced, less colorful version. In Iguana iguana, the dewlap forms a pendulous, scalable flap up to 30 cm long in adults, with subtle coloration dominated by the lizard's overall green hue but expandable via the same hyoid-driven mechanics. These variations arise from differences in length, muscle fiber composition, and skin elasticity, with larger dewlaps supported by elongated ceratobranchials for greater extension range. Habitat influences dewlap morphology in anoles, where over 400 Anolis species have undergone , resulting in diverse forms adapted to island environments. Species in dense foliage habitats, such as trunk-crown ecomorphs, exhibit larger dewlaps—up to 20% of body length—to facilitate visibility amid cluttered vegetation, whereas trunk-ground ecomorphs have smaller, more subdued versions suited to open perches. This ecomorphological pattern underscores how hyoid modifications enable habitat-specific mechanics without altering core extension principles. Similar hyoid apparatuses in fossil squamates, such as those preserved in specimens like Cryptovaranoides microlanius (dated to ~205 million years ago), recently confirmed as a crown-group squamate, suggest that dewlap-like extensible throat structures may have been present in early lineages, inferred from impressions and skeletal supports analogous to modern iguanids. In living species, these mechanics integrate with behaviors like dewlap extension during , where rapid pulses enhance signal precision through precise muscle control.

Other Reptiles

In non-lizard reptiles, structures analogous to dewlaps are rare and typically manifest as subtle, non-inflatable folds of loose in the or region, serving functional roles in feeding, , or rather than display. These features lack the colorful, extendable nature seen in and are adapted to the specific lifestyles of , , and other groups. In snakes, the neck skin exhibits remarkable distensibility to facilitate the ingestion of large prey through macrostomy, a derived trait enabling whole-prey consumption. Mechanical analyses reveal that snake skin properties vary longitudinally, with the neck region capable of significant stretching—up to 25% more extensible in species like boa constrictors following large meals—to accommodate prey size without tearing. This loose, elastic skin forms temporary folds during swallowing, particularly in constrictors such as pythons, where it expands to envelop oversized items like mammals or birds. Among (chelonians), subtle throat folds appear in the , the anterior portion of the plastron, where flexible, leathery skin predominates in like softshell (). In these aquatic or semi-aquatic forms, the lacks rigid scutes and instead consists of a thinner corneous layer, allowing flexibility for extension during feeding or burrowing into . This structure supports gular pumping, a ventilatory involving expansion to assist inflation, especially in without diaphragms. Gular scutes in hard-shelled chelonians resemble reduced flaps but are bony projections integrated into the shell for protection rather than extension. Dewlap-like structures are absent in most crocodilians, whose is dominated by thick, armored osteoderms that prioritize defense and hydrodynamic efficiency over flexible throat flaps. In captive reptiles, such as and , loose gular or neck can be susceptible to bacterial if is poor, often requiring veterinary like topical antifungals. Evolutionarily, these reduced throat structures in non-lizard reptiles may represent homologous loose to that in but are minimized in or burrowing species for streamlining and reduced . The scarcity of prominent dewlaps outside correlates with diminished reliance on visual signaling, as snakes and turtles emphasize chemosensory or tactile cues in often obscured or underwater habitats.

Occurrence in Birds

In , dewlaps, commonly known as wattles, are prominent in species such as the wild turkey (Meleagris gallopavo) and various pheasants (family ), where they appear as reddish, fleshy, pendulous, and highly vascularized appendages hanging from the or . These structures are integral to the bird's head ornamentation, often connected to the —a fleshy on the crown—in a continuous expanse of bare skin that facilitates rapid physiological changes. In wild turkeys, the wattle typically measures several inches in length and can enlarge or contract in size through blood engorgement, particularly during the breeding season when males display heightened activity. Domestic breeds exhibit variations, with heritage turkeys often featuring more prominent and elongated wattles compared to their wild counterparts, a result of that emphasized exaggerated physical traits for visual appeal and robustness. The adaptive roles of these dewlaps in ground-dwelling include the wattles undergoing color shifts—from pale to vivid red or blue—correlating with states of or excitement, serving as quick visual indicators of the bird's condition. Due to their dense vascular network, these structures also contribute to by dissipating excess heat during physical exertion. Historical domestication of has prominently shaped dewlap characteristics, with early in like turkeys and chickens prioritizing larger, more conspicuous wattles to enhance breed distinctiveness and as markers of vitality in breeding stock. In modern contexts, swollen wattles serve as diagnostic indicators for diseases such as fowl pox, a viral infection that causes proliferative lesions and in the unfeathered skin of affected birds, including turkeys and chickens. Across related species like (Numida meleagris) and domestic chickens ( gallus domesticus), dewlaps exhibit similar fleshy, vascular , with size variations strongly tied to social hierarchy; dominant males typically possess larger, more pronounced wattles, reflecting their status in peck-order dynamics and influencing access to resources. In , for instance, adult males display elongated, upward-folded wattles that exceed those of females, underscoring sex- and rank-based dimorphism in these communal flocks.

Seabirds and Other Avian Groups

In seabirds, the magnificent frigatebird (Fregata magnificens) exemplifies the use of an inflatable gular pouch as a prominent dewlap-like structure. This pouch, a thin, elastic membrane connected to the lungs via an opening near the scapula, allows males to inflate it with air into a balloon-like form that can extend over the throat and foreneck, reaching sizes large enough to obscure the head during displays. The pouch features bright red coloration in breeding males, contrasting sharply with their black plumage, and its inflation is controlled seasonally by muscular action during the courtship period. This aerial adaptation enables frigatebirds to perform mate-attraction displays while soaring over oceanic habitats, where the inflated pouch serves as a visual signal visible from afar in tropical and subtropical environments. In contrast, among ground-dwelling ratites, cassowaries (Casuarius spp.) possess paired wattles—bright red or golden folds of loose throat skin hanging from the —that integrate with the casque and neck region for potential protective functions during aggressive interactions or territorial defense. Males inflate the underlying throat area to produce low-frequency calls during courtship, enhancing the structure's role in communication and physical encounters in dense settings. Variations in dewlap presence occur across groups, with such inflatable throat pouches evident in ratites like (Dromaius novaehollandiae), where a tracheal pouch in the throat inflates to generate deep booming sounds for long-distance signaling in open landscapes. These structures are largely absent in most passerines, the diverse order, but persist in select non-passerine lineages, reflecting adaptive divergences in and strategies.

References

  1. [1]
    DEWLAP Definition & Meaning - Merriam-Webster
    1. a fold or flap of skin on the neck of some animals: such as a : loose skin hanging under the neck of dogs and cattle of some breeds.
  2. [2]
    Evolution of the ungulate dewlap: thermoregulation rather than ...
    Jul 18, 2016 · According to this idea, the dewlap makes it easier for predators to get a hold on their prey, and because only 'high quality' individuals can ...
  3. [3]
    Evolution of Anolis Lizard Dewlap Diversity | PLOS One
    Dewlaps are flaps of skin beneath the chin that are extended and combined with head and body movements for visual signals and displays.
  4. [4]
    Dewlap - an overview | ScienceDirect Topics
    Dewlap is defined as the loose skin under the chin and throat of rabbits, which can become infected and sore, particularly in conditions of excessive moisture ...
  5. [5]
    Functional morphology of dewlap extension in the lizard Anolis ...
    The dewlap is an extendible flap of skin ordinarily folded under the throat. Lizards, particularly those in the genus Anolis, extend their dewlaps during ...Missing: composition | Show results with:composition
  6. [6]
    Overview of Reptiles - Exotic and Laboratory Animals
    Skin characteristics (eg, crests, spines, dewlaps) are often used for species or sex identification in those species that exhibit dimorphic variation. All ...
  7. [7]
    Caruncles, Combs and Wattles: The Features, Functions and ...
    Rating 4.8 (6) Jun 1, 2013 · It generally refers to the protuberances found on the head and neck including the combs, earlobes, snoobs and wattles. Their purposes vary, but include ...
  8. [8]
    Anatomy: Wattle - Birds Outside My Window
    Sep 17, 2010 · A wattle is a fleshy dewlap or caruncle hanging from various parts of the head or neck in several groups of birds, goats and other animals.
  9. [9]
    Dewlap - Etymology, Origin & Meaning
    Originating in mid-14c. from Old English læppa "loose piece," lappe means a hanging fold of skin, first on oxen, then turkeys, and later humans with age.
  10. [10]
    dewlap - Wiktionary, the free dictionary
    Etymology. From Middle English dewlappe. The first element may be dew (and if not, seems to have been altered by folk etymology to resemble it); the second ...
  11. [11]
    dewlap, n. meanings, etymology and more | Oxford English Dictionary
    The earliest known use of the noun dewlap is in the Middle English period (1150—1500). OED's earliest evidence for dewlap is from 1398, in ...
  12. [12]
    DEWLAP definition in American English - Collins Dictionary
    a loose fold of skin hanging from the throat of cattle and certain other animals, or a similar loose fold under the chin of a person.
  13. [13]
    The Tempest - Act 3, scene 3 | Folger Shakespeare Library
    1595 Dewlapped like bulls, whose throats had hanging at 1596 60 'em 1597 Wallets of flesh? Or that there were such men 1598 Whose heads stood in their ...
  14. [14]
    What determines dewlap diversity in Anolis lizards? An among ...
    Jan 19, 2009 · We find evidence for the role of the dewlap as a pursuit-deterrence signal in both males and females as relative dewlap size is larger on ...Missing: vertebrates | Show results with:vertebrates
  15. [15]
    [PDF] Importance of Dewlap Display in Male Mating Success in Free
    The dewlap is believed to have many functions in Anolis lizards (Losos and Chu, 1998), and there is good experimental evidence that the male dewlap plays a role ...
  16. [16]
    The brown anole dewlap revisited: do predation pressure, sexual ...
    May 8, 2018 · Several studies have shown that male brown anoles use their dewlap during territorial disputes (Scott, 1984) and/or during courtship (Simon, ...
  17. [17]
    Physiological and thermographic response to heat stress in zebu cattle
    The main body characteristics of these animals include the presence of a hump, abundant and pleated dewlap, pigmented and loose skin and shorter and thinner ...
  18. [18]
    Relationship of relative dewlap size, body condition, and number of...
    The higher dewlap size in male suggests that the dewlap can store fat reserves, especially in well-fed animals and fat stores can serve as an energy reserve ...
  19. [19]
    chapter 4 types and breeds of tropical and temperate cattle
    The hump consists of muscle, connective tissue, and variable amounts of fat. The size and shape of the hump vary by breed, sex, and age of the animal. The ...
  20. [20]
    Parts of the Goat - Cornell CALS
    Here we look at overall parts of the goat, and in particular, relevant body parts for dairy and meat animals. ... dewlap; 11. point of shoulder; 12. heart ...
  21. [21]
    African Indigenous Cattle: Unique Genetic Resources in a Rapidly ...
    Bos indicus breeds can effectively regulate their body temperature against thermal stress and are better adapted to hot weather than Bos taurus breeds ...
  22. [22]
    Genomic clues of the evolutionary history of Bos indicus cattle
    Sep 2, 2019 · The adaptation of zebu cattle to tropical and sub-tropical environments makes them more efficient in coping with heat, humidity, nutritionally ...
  23. [23]
    Cattle adapted to tropical and subtropical environments
    The carbon fixation pathway in C4 grasses allows these plants to be more adapted to warm and dry regions (Volenec and Nelson, 2007), commonly found in tropical ...
  24. [24]
    African Indigenous Cattle: Unique Genetic Resources in a Rapidly ...
    They are adapted to dry environmental conditions and high temperatures and are known to be more resistant to tick infestation compared to Bos taurus cattle ( ...
  25. [25]
  26. [26]
    Basic Anatomy, Physiology, and Husbandry | Veterian Key
    Sep 6, 2016 · A female rabbit has a large fold of skin over the throat known as a dewlap. Breeding does pull fur from this area to line their nests before ...Anatomy · Sense Organs And Nervous... · Digestive System
  27. [27]
    Why Does My Rabbit Have a Double Chin (or Dewlap)?
    Dewlaps are secondary sex characteristics tied to certain hormones produced in adult female rabbits, and to a lesser extent, some male rabbits.
  28. [28]
  29. [29]
    Dewlap: The Rabbit Double Chin - The Bunny Lady
    Sep 11, 2019 · Female rabbits develop dewlaps to help when it comes time to make a nest for babies. They have nothing to do with attracting mates and there is ...
  30. [30]
    Disorders and Diseases of Rabbits - All Other Pets
    Wet Dewlap (Moist Dermatitis)​​ Female rabbits have a heavy fold of skin called a dewlap on the front of the neck. As the rabbit drinks, this skin may become wet ...
  31. [31]
    RABBITS - PMC - PubMed Central - NIH
    Rabbits that drink from crocks may develop “blue fur,” which is a moist dermatitis of the dewlap that is associated with Pseudomonas infections.
  32. [32]
    Canine Glossary | United Kennel Club (UKC)
    Dewlap: Loose, pendulous skin under the throat and neck. Dish-faced: Slight concaveness of foreface when viewed in profile. Disqualification: A decision made by ...<|control11|><|separator|>
  33. [33]
    Dewlaps: They Have a Purpose - National Purebred Dog Day
    Sep 25, 2018 · When a dog pants, air descends directly to the lungs. The dewlap prevents the shock of cold air from being taken directly into the body.
  34. [34]
    Lions Have Manes; Leopards Have…Dewlaps? - Panthera.org
    Sep 4, 2018 · In mature male leopards, a well-developed dewlap is prominent and one of the best ways to determine age.
  35. [35]
    Where Did the Dewlap Come From? - Anole Annals
    Oct 14, 2015 · There is some evidence to support a correlation between dewlap morphology and a lizard's physiological characteristics or body condition.
  36. [36]
    Functional morphology of Dewlap extension in the lizard Anolis ...
    Aug 6, 2025 · The dewlap is an extendible flap of skin ordinarily folded under the throat. Lizards, particularly those in the genus Anolis, extend their ...<|separator|>
  37. [37]
    The mechanism of dewlap extension in Anolis carolinensis (Reptilia
    Anolis carolinensis has two aggressive displays involving movements of the hyoid apparatus: erection of the throat and extension of the dewlap.
  38. [38]
    The biomechanical basis of evolutionary change in a territorial display
    May 29, 2013 · The Dewlap Mechanism​​ Bels (1990) and Font & Rome (1990) described in detail the anatomy of the Anolis hyoid apparatus as it related to the ...
  39. [39]
    The incredible shrinking dewlap: signal size, skin elasticity ... - NIH
    Sep 19, 2015 · Activation of the paired ceratohyoid muscles at the front of the hyoid apparatus causes the ceratobranchials to extend forward, away from the ...
  40. [40]
    The Vibrant Dewlap of Green Anoles
    May 10, 2023 · The dewlap can range in color from bright red to pink, and even orange or yellow. In the Ouachita National Forest, I have been lucky enough to ...
  41. [41]
    Common Green Iguana - Animal Diversity Web
    Other distinguishing features of this species include a pendulous dewlap under the throat, a dorsal crest made up of dermal spines that run from the mid neck to ...
  42. [42]
    Fiber Type Composition of the Muscle Responsible for Throat Fan ...
    Jun 16, 2004 · Male anoles extend their dewlap in as little as 0.25 s [Bels, 1990], and will either maintain extension for the duration of the head bobbing ...
  43. [43]
    Signal size allometry in Anolis lizard dewlaps - PMC - NIH
    Jul 5, 2023 · Anolis lizards possess an elaborate retractable throat fan called a dewlap that is used for visual communication and differs greatly in size and ...
  44. [44]
    Evolution of Anolis Lizard Dewlap Diversity - PMC - PubMed Central
    Anolis dewlaps are skin flaps used for visual signals, with substantial diversity in size, patterns, and colors, though most fit into six patterns.
  45. [45]
    Female Dewlaps - Anole Annals
    May 31, 2012 · Female dewlap size differed significantly among habitat specialists, with crown-giant and twig anoles having larger dewlaps than the other ecomorphs.
  46. [46]
    [PDF] Evolution of an ornament, the dewlap, in females of the lizard genus ...
    Studies on the role of female ornaments in territorial defence are limited; in at least one species, the size of a female ornament was shown to affect the ...
  47. [47]
    Scientists Discover Oldest Modern Lizard Fossil, Shaking Up ...
    Dec 16, 2024 · A 205-million-year-old fossil, Cryptovaranoides microlanius, was confirmed as the oldest modern lizard, shifting Squamata origins back by 35 million years.
  48. [48]
    Origin and Early Evolution of Squamates and Their Kin: From Fossils ...
    Squamates (lizards, including snakes) are the most diverse group of terrestrial vertebrates on Earth today and have an evolutionary history dating back to ...
  49. [49]
    Behavioural display systems across nine Anolis lizard species
    Feb 3, 2010 · First, numerous features of the mechanical system that extends the dewlap (including muscle fibre size, cartilage length and motoneuron somata) ...
  50. [50]
    Description and Physical Characteristics of Reptiles - All Other Pets
    The males of some species have a loose flap of skin called the dewlap that can be extended to either intimidate a predator or to help attract a mate. And, as ...Missing: excluding | Show results with:excluding
  51. [51]
    Mechanical properties of snake skin vary longitudinally, following ...
    Dec 23, 2024 · ... eating large prey modifies the skin of some snakes. For example, compared with fasted snakes, a wide variety of snake species that we have ...
  52. [52]
    Mechanical properties of snake skin vary longitudinally, following ...
    Dec 15, 2024 · The ability for snakes to ingest large prey (macrostomy) is a widespread, derived trait that involves distending the skin during ingestion ...
  53. [53]
    Skin structure and cornification proteins in the soft-shelled turtle ...
    The epidermis of the carapace and plastron has a thick corneous layer composed of alpha-corneocytes surrounded by a cornified cell envelope. The softer ...
  54. [54]
    (PDF) Skin structure and cornification proteins in the soft-shelled ...
    Aug 5, 2025 · The epidermis of the carapace and plastron has a thick corneous layer composed of alpha-corneocytes surrounded by a cornified cell envelope. The ...
  55. [55]
    Crocodylians evolved scattered multi-sensory micro-organs - PMC
    Jul 2, 2013 · Crocodylians have exquisite mechanosensory abilities thanks to the presence of small integumentary sensory organs (ISOs) distributed on postcranial and/or ...
  56. [56]
    Trionychidae - an overview | ScienceDirect Topics
    Trionychidae are soft-shelled, aquatic turtles with flattened, oval shells, a proboscis, and leathery skin, found in North America, Africa, Asia, and Indo- ...
  57. [57]
    Wild Turkey Anatomy
    Here is a breakdown of some of the wild turkey's distinct anatomical features and their function, from snood to spur, if you will.
  58. [58]
    Swinhoe's Pheasant – wattles & function - Bird Ecology Study Group
    Sep 23, 2021 · Many birds have wattles (a caruncle), a fleshy structure, often brightly coloured, that adorns various parts of the head and neck, occasionally hanging.Missing: dewlaps | Show results with:dewlaps
  59. [59]
    Gamebird Anatomy - Veterian Key
    Nov 10, 2022 · The ornamental structures on the head include cere, comb, wattles, and ear lobes when present. The appearance of orifices and ornamental ...
  60. [60]
    [PDF] Galliformes - Harrison's Bird Foods
    The dewlaps of turkeys are smooth, can in- crease and decrease in size ... Silver Pheasant, eared pheasants, Golden Pheasant,. Lady Amherst's Pheasant ...
  61. [61]
    [PDF] Choosing a Variety and obtaining Poults - The Livestock Conservancy
    Heritage turkeys are naturally mating, with long lifespans. Standard varieties have approved descriptions, and non-standard varieties can be marketed as  ...
  62. [62]
    Wild Turkeys - UC IPM
    When males are excited, the wattles and the bare skin of the head and neck ... prevent their “scratching” behavior and discourage them from frequenting the area.<|control11|><|separator|>
  63. [63]
    What Are Wattles For? - National Audubon Society
    Feb 25, 2019 · That flap of bumpy red skin hanging off a male turkey's face is both functional and fashionable.
  64. [64]
    What's a turkey wattle for anyway?
    Nov 25, 2019 · A turkey wattle helps males attract hens, releases excess heat, and its color changes to show fear or illness.
  65. [65]
    Domestic Turkeys and Their Wild Ancestors | Smithsonian Institution
    Oct 9, 2012 · The domestic turkey that ends up on the dinner table exhibits less genetic variation than not only its ancestral wild counterparts, which were first ...
  66. [66]
    Fowlpox in Chickens and Turkeys - Poultry - Merck Veterinary Manual
    Fowlpox is a viral infection in chickens and turkeys, causing skin lesions (cutaneous form) and lesions in the upper GI and respiratory tracts (diphtheritic ...Etiology Of Fowlpox · Clinical Findings Of Fowlpox · Diagnosis Of Fowlpox
  67. [67]
    The relationship between the comb and social behaviour in laying ...
    Dec 31, 2011 · This study examined the relationship between dominance behaviour and the size as well as colour of domestic laying hens' combs.
  68. [68]
    [PDF] Observations on the Wattles of Adult Helmeted Guinea Fowls ...
    The differences in the shape and size observed in wattles of heavier adult males were that they appeared elongated, large and folded upwards at their margins.
  69. [69]
    Age related differences in wattle length between male and female ...
    Three hundred and sixty guinea fowls (180 birds per sex) were used in determining a more accurate and farmer friendly sexing technique.Missing: hierarchy | Show results with:hierarchy
  70. [70]
    Magnificent Frigatebird Identification - All About Birds
    Adult male. Male's gular sac becomes bright red and is inflated like a large balloon during courtship displays and early incubation period, but becomes pale ...
  71. [71]
    Plumages, Molts, and Structure - Great Frigatebird - Fregata minor
    Gular Sac. Red gular sac of male inflated from lungs through opening anterior to humeral articulation of scapula; extends over throat, foreneck when expanded ...Missing: anatomy | Show results with:anatomy
  72. [72]
    Morphology and ornamentation in male magnificent frigatebirds
    Male magnificent frigatebird (Fregata magnificens) ornamentation includes bright iridescent plumage and a red inflatable gular pouch.
  73. [73]
    Gular Pouch Used to Attract Mate — Biological Strategy - AskNature
    Aug 18, 2016 · A male frigatebird or Man-o-war bird has selected a suitable nest site and is advertising for a mate by inflating its crimson throat pouch.Missing: display | Show results with:display
  74. [74]
    Cassowary | San Diego Zoo Animals & Plants
    Its casque is larger and more flared than the southern cassowary's, and the throat skin and wattle are either red or golden, depending on where the cassowary is ...
  75. [75]
    Casuarius casuarius (southern cassowary) - Animal Diversity Web
    Southern cassowaries are the largest of the three species of cassowary and the only species to have have two bright red flaps of skin, called wattles, hanging ...Scientific Classification · Physical Description · Reproduction
  76. [76]
    Emu | San Diego Zoo Animals & Plants
    The emu has a pouch in its throat that is part of the bird's windpipe and is used for communication. When the pouch is inflated, the emu can make deep booming, ...
  77. [77]
    The evolution of closed‐mouth vocal behavior in birds
    Closed-mouth vocalizations are unlikely to be ancestral in birds and have evolved independently at least 16 times within Aves, predominantly in large-bodied ...
  78. [78]
    Magnificent Frigatebird | Audubon Field Guide
    Climate Threats Facing the Magnificent Frigatebird. Choose a temperature scenario below to see which threats will affect this species as warming increases. The ...
  79. [79]
    The effects of climate change on wildlife biodiversity of the ...
    These species are especially vulnerable to changing climatic conditions because they are restricted to a particular geographical location and are often unable ...