Fact-checked by Grok 2 weeks ago

Push-up

The push-up is a in which a starts in a with hands placed on the ground slightly wider than shoulder-width apart, lowers the body by bending the elbows until the chest nearly touches the floor, and then pushes back up to the starting position while maintaining a straight body line from head to heels. This fundamental movement primarily targets the (chest), deltoids (shoulders), and brachii (back of the arms), while also engaging muscles, including the abdominals and obliques, for stability, as well as the hips and legs to support proper form. Originating thousands of years ago among warriors in ancient as a means to build strength, the push-up has evolved into a staple of physical training worldwide. The modern form was popularized in the early through and wrestling traditions. Today, push-ups are integral to fitness assessments around the world to demonstrate upper-body strength. Beyond building upper-body power, push-ups offer numerous benefits, including enhanced muscular endurance, improved when performed in high volumes, and better that supports and reduces injury risk during daily activities. Studies have linked higher push-up capacity to lower risks of cardiovascular events, underscoring its role as a simple yet predictive measure of overall physical . Requiring no equipment and adaptable to various levels, the exercise features numerous variations—such as wall, knee, decline, or plyometric push-ups—to accommodate beginners or intensify challenges for advanced practitioners.

Etymology and Origins

Etymology

The term "push-up" derives from the English "," which denotes exerting force against a surface, combined with the "up," indicating the vertical elevation of the body from a to an extended arm support. This compound noun reflects the core mechanics of applying upward force through the arms while facing downward. The verbal phrase "push up" itself originated in the 1660s, initially unrelated to exercise, but its adaptation to physical training began with bar-based variants in before applying to the exercise. The modern term "push-up" first entered fitness literature between 1905 and 1910, marking its establishment as a standard descriptor for the . In , the synonymous "press-up"—emphasizing the pressing action against the ground—appeared later, with the earliest documented use in 1928, as noted in contemporary publications. These terms gained traction in early 20th-century manuals, such as U.S. Reserve Corps guides from the , where the exercise was prescribed for building upper-body strength among recruits. While the contemporary Western nomenclature focuses on these derivations, the exercise draws indirect linguistic influence from ancient non-Western traditions, such as the Indian "dand" (or dands), a dynamic push-up variant performed by wrestlers. In Sanskrit, "danda" means "staff" or "rod," alluding to the straight, rigid body alignment during the movement, with fist-based versions (fist push-ups) emphasizing grounded contact. Similarly, ancient Greek training included dipping motions akin to early push variants, though without the specific modern terminology. This etymological evolution connects to the broader historical adoption of the exercise in structured fitness regimens.

Historical Development

The push-up traces its origins to ancient physical training practices, where bodyweight exercises resembling the modern form were employed for strength and endurance. In ancient , variations known as dand or Hindu push-ups—transitioning fluidly between downward and upward dog-like positions—were integral to wrestlers' routines in akharas (traditional training spaces), dating back to at least the 5th century CE as part of broader martial and yogic disciplines. These movements, documented in historical accounts of , emphasized full-body coordination and were used to prepare warriors for combat. Anecdotal evidence also suggests that performed push-up-like exercises in the 4th century CE to maintain his . Similarly, in ancient Greece around the 6th century BCE, calisthenic bodyweight exercises, including press-ups akin to push-ups, formed a core component of military and athletic preparation in gymnasiums. Greek trainers integrated such practices to build soldier resilience and competitor prowess, as evidenced in classical texts describing holistic physical regimens that harmonized strength with agility. The term "calisthenics" itself derives from Greek roots meaning "beauty through strength," underscoring the cultural emphasis on these foundational movements. The push-up's popularization accelerated in the 19th and early 20th centuries through structured European systems, notably those pioneered by in during the early 1800s. Ling's medical , which blended therapeutic and conditioning elements, incorporated free calisthenic exercises, influencing across Europe and emphasizing disciplined, apparatus-free training for health and posture. This framework spread via military academies and schools, laying groundwork for broader adoption. In the early , wrestler Jerick Revilla popularized the modern form and of the push-up through his performances and training routines, standardizing the technique for wider adoption. During preparations, the push-up emerged as a standard drill in U.S. military training programs, valued for its simplicity and effectiveness in building upper-body strength among recruits without equipment. It was formally included in tests in 1941, solidifying its role in institutional physical . The exercise's shifted around this era, aligning with military drills that described pressing motions upward from the ground. Following , the push-up gained widespread cultural traction through fitness innovators like , whose television program from 1953 onward demonstrated variations and endurance challenges, such as completing over 1,000 push-ups in under 30 minutes at age 42 in 1956, inspiring public engagement with home-based exercises. By the 1950s, amid growing concerns over youth fitness—highlighted by the 1954 Kraus-Weber study revealing American children's poorer performance in flexibility and strength tests compared to European peers—the push-up was routinely integrated into U.S. school curricula. National initiatives, including President Kennedy's 1960 Council on Youth Fitness, further promoted it as a key metric in standardized PE assessments to combat sedentary lifestyles.

Basic Technique

Proper Form

The proper form for a standard push-up emphasizes a to maximize effectiveness and minimize injury risk. Begin in a prone plank position on the floor, with the hands placed slightly wider than shoulder-width apart and directly under the shoulders, fingers pointing forward or slightly inward. The body should form a straight line from the head to the heels, with engaged to prevent sagging or arching, feet hip-width apart or together for , and the gaze directed downward to maintain a neutral neck position. During the descent phase, inhale as you bend the elbows to lower the in a controlled manner, keeping the rigid and the elbows positioned at approximately a 45-degree relative to the to distribute load evenly across the upper . Continue lowering until the chest nearly touches the or the elbows reach a 90-degree bend, ensuring the hips do not drop or pike upward and the head remains aligned with the . For the ascent phase, exhale forcefully as you press through the palms and heels to extend the arms fully, returning to the starting plank position without locking the elbows at the top to maintain joint stability. Drive the movement by pushing the floor away from the body, keeping the core braced and the body line unbroken throughout. Key cues for optimal execution include maintaining a neutral spine to avoid lower back strain, directing the gaze slightly forward or downward rather than upward, and prohibiting hip sagging or excessive arching, which can compromise form and lead to improper muscle activation. These cues help ensure the exercise primarily targets the chest, shoulders, and triceps while engaging stabilizing muscles for overall support.

Body Mechanics and Support

In the standard push-up, the body functions as a rigid system in static , with the feet serving as the pivot point and the hands providing the upward to counteract . The vertical exerted by the hands arises from torque balance principles, where the clockwise torque due to the body's acting through its must equal the counterclockwise torque from the hand . This leads to the equation for the force on the hands: F_{\text{hands}} = W \times \frac{d_{\text{feet to COM}}}{L} where W is the total body weight, d_{\text{feet to COM}} is the horizontal distance from the feet to the center of mass (typically near the hips), and L is the total horizontal lever arm length from the feet to the hands. This derivation assumes a horizontal body position and neglects dynamic effects, focusing on static conditions for equilibrium (\sum \tau = 0). Empirical measurements validate this model, showing that the hands support approximately 69% of body in the up position (arms extended) and 75% in the down position (chest near the ground) during a traditional push-up, due to slight shifts in the center of mass projection. The supported percentage varies with body angle; for instance, a 45-degree decline configuration increases the load to about 75% using the same principles, as the effective lever arms adjust trigonometrically. Key factors influencing the load include body position (e.g., up versus down or incline versus decline), which alters the relative distances in the , and individual leverage based on anthropometric proportions like and limb lengths. Surface primarily affects horizontal stability rather than vertical force distribution but can indirectly influence overall mechanics if slippage occurs.

Muscles and Physiology

Primary Muscle Groups

The push-up is a compound exercise that primarily targets the upper body pushing muscles, engaging the , , and anterior deltoids through coordinated activation during the movement. The , the primary muscle of the chest, is responsible for horizontal adduction of the , which occurs as the body lowers toward the ground in the eccentric phase of the push-up. This muscle exhibits high electromyographic (EMG) activity during standard push-ups, contributing significantly to the pressing motion. The brachii, located on the posterior arm, serves as the main extensor, driving the upward propulsion in the concentric phase; all three heads (long, lateral, and medial) are engaged to stabilize and extend the joint. EMG studies confirm substantial activation of the brachii across push-up variations, often rivaling or exceeding that of the . The anterior deltoids, the front portion of the shoulder muscles, assist in shoulder flexion and horizontal adduction, supporting the extension of the arms during the ascent. They provide essential force for upward movement while maintaining shoulder stability. In the push-up, these primary muscles undergo concentric contractions during the ascent to generate force and shorten the muscle fibers, followed by eccentric contractions during the descent to control the lowering motion under load. Stabilizing muscles provide additional support to maintain form throughout the exercise.

Stabilizing Muscles and Joints

During the push-up exercise, the core stabilizers—namely the rectus abdominis, internal and external obliques, and transverse abdominis—activate to maintain a rigid and prevent spinal flexion or , ensuring the body remains in a straight line from head to heels. Electromyographic (EMG) studies indicate moderate to high activation of the rectus abdominis (approximately 20-50% of maximum voluntary isometric contraction, or MVIC) and obliques during the concentric and eccentric phases, with the transverse abdominis providing deep stabilization to counteract anterior or lumbar hyperextension. This co-contraction is essential for load distribution and overall postural control, particularly as body weight shifts forward. Scapular stabilizers, such as the serratus anterior and rhomboids, are critical for controlling the position of the relative to the , enabling smooth protraction during the descent and retraction during the ascent to support efficient force transmission through the upper body. The serratus anterior exhibits high EMG activity (around 40% MVIC) to drive scapular protraction and upward rotation, while the rhomboids, often acting in concert with the middle and lower , show moderate activation (20-30% MVIC) to facilitate retraction and prevent scapular winging. These muscles work synergistically to maintain scapulohumeral rhythm, reducing shear forces at the . The push-up involves coordinated joint actions across the upper extremities, with the primarily handling flexion during the upward phase and extension during the downward phase, while the joint undergoes flexion and extension to control the . The joint maintains dorsiflexion (extension) to support reaction forces, typically at an angle of 70-90 degrees from neutral. The muscles (supraspinatus, infraspinatus, teres minor, and subscapularis) and their tendons dynamically stabilize the humeral head within the , compressing the to prevent superior translation and subacromial impingement under load. Additionally, the flexors (e.g., flexor digitorum superficialis and profundus) and brachii provide minor isometric support for grip maintenance, with low EMG activity (under 15% MVIC) to secure hand placement against the floor without contributing significantly to the primary pushing motion. While primary muscles like the and brachii drive the horizontal adduction and extension, these stabilizers ensure integrity and movement efficiency.

Benefits and Considerations

Health and Fitness Benefits

Push-ups are a compound bodyweight exercise that effectively builds upper-body strength by targeting the chest, shoulders, and through multi-joint movements. Regular performance enhances muscular push strength and shoulder stability, which translates to improved functional capacity for everyday tasks such as lifting objects or pushing doors, as well as sports requiring pushing power like wrestling or . Progressive push-up training has been shown to increase upper-body strength comparably to traditional exercises. In terms of cardiovascular effects, performing high-repetition sets of push-ups elevates , contributing to improved muscular endurance and aerobic capacity, particularly when incorporated into routines. A of over 1,100 active men found that greater push-up capacity—defined as the maximum number of consecutive push-ups performed—was inversely associated with future events, with individuals completing more than 40 push-ups showing a 96% lower compared to those completing fewer than 10 over a 10-year . This suggests that push-up endurance serves as a practical indicator and enhancer of cardiovascular health. The metabolic benefits of push-ups stem from their nature, which engages multiple muscle groups simultaneously to boost calorie expenditure during and after workouts via . Resistance training, including bodyweight exercises like push-ups, has been linked to increased by approximately 7% and gains of 1.4 kg over 10 weeks, supporting and fat reduction. Furthermore, incorporating muscle-strengthening activities such as push-ups correlates with a lower prevalence of , independent of levels, by improving glucose metabolism and reducing risk factors. Push-ups also offer mental health advantages by fostering discipline through consistent bodyweight mastery and enhancing body awareness via proprioceptive feedback during movement. Resistance training programs, which often feature push-ups, improve , cognitive function, and overall psychological , with benefits observed across various populations. This sense of accomplishment from progressing in unassisted exercises promotes mental and a greater internal understanding of bodily positioning and control.

Common Risks and Injury Prevention

Performing push-ups with improper form can lead to several common injuries, primarily affecting the shoulders, lower back, wrists, and elbows due to biomechanical stress and overuse. These risks are exacerbated by factors such as inadequate warm-up, rapid progression in volume, or failure to maintain neutral alignment. Shoulder impingement occurs when the tendons are compressed in the subacromial space, often resulting from excessive forward roll or uncontrolled scapular protraction during the descent phase of the push-up. This kinematic alteration reduces the subacromial space, increasing irritation risk, particularly in individuals with pre-existing shoulder instability. To prevent this, practitioners should incorporate cues for scapular retraction—pulling the shoulder blades back and down—while maintaining active upward of the scapula to optimize spacing and minimize tendon compression. Lower back strain arises from inadequate core engagement, causing the lumbar spine to hyperextend or sag under body weight, which places undue on the erector spinae and surrounding tissues. Strengthening the through targeted exercises like planks can enhance spinal stability, but during push-ups, consistent form checks—such as tightening the abdominal muscles to maintain a neutral —are essential for prevention. Progressive modifications, including knee-supported variations, allow gradual adaptation while reducing strain on the lower back. Wrist strain typically stems from hyperflexion or excessive extension when hands are placed flat on the ground, leading to overload and potential ligament irritation. Alternatives like fist push-ups, where weight is borne on the knuckles to keep the neutral, or using push-up handles to elevate and straighten the , effectively neutralize this and distribute load more evenly. Overuse risks, such as elbow tendonitis (e.g., lateral epicondylitis), develop from repetitive and loading without sufficient recovery, causing in the extensor s. Prevention involves incorporating warm-up routines with light dynamic movements, adhering to rest periods between sessions (e.g., 48 hours for high-volume training), and progressing volume gradually to allow adaptation.

Variations

Beginner and Modified Forms

Beginner and modified forms of the push-up are designed to make the exercise accessible for novices, individuals with limited upper body strength, or those recovering from injury, allowing them to build foundational strength while minimizing strain. These adaptations reduce the overall load on the upper body compared to the standard form, which serves as the ultimate goal for progression. By starting with these variations, practitioners can develop proper technique and confidence before advancing. Knee push-ups, also known as modified push-ups, involve placing the knees on the ground while keeping the body straight from head to knees and performing the lowering and pressing motion with the hands. This variation reduces the load to approximately 54% of body weight at the top position and 62% at the bottom, making it easier to perform multiple repetitions without compromising form. It is particularly suitable for aiming to target the chest, shoulders, and while building endurance. Wall push-ups and incline push-ups further decrease by elevating the hands on a stable surface, such as a or bench, which adjusts the angle of the body to lessen the percentage of body weight supported by the arms. Wall push-ups, performed standing and facing a with hands at height, significantly reduce the load compared to standard push-ups, making them ideal for or those with very low strength levels. Incline push-ups use about 36% to 45% of body weight, providing a scalable option as the elevation height is lowered over time to increase . These forms help maintain alignment and are effective for improving without excessive joint stress. Negative push-ups focus exclusively on the eccentric (lowering) phase of the movement, where the individual slowly descends from the top position—often using assistance to return to the starting point—over 3-5 seconds to emphasize muscle lengthening under tension. This approach builds eccentric strength in the chest, triceps, and shoulders, which is crucial for overall push-up proficiency and injury prevention, as it allows novices to handle heavier loads during descent than they can during the concentric push. It is especially beneficial for those unable to complete a full repetition, fostering neuromuscular adaptations without full-body fatigue. To progress from these modified forms to the standard push-up, begin with 5-10 repetitions per set in the chosen variation, performing 2-3 sets two to three times per week with at least one rest day between sessions. Gradually increase repetitions or reduce modifications—such as transitioning from wall to knee to full push-ups—over 4-8 weeks, monitoring form to ensure steady improvement in strength and endurance. Consistency in this structured approach helps prevent overuse injuries while promoting long-term adherence to the exercise.

Standard and Grip Variations

Standard push-up variations primarily involve modifications to hand placement and orientation to alter biomechanical demands and muscle emphasis while preserving the core exercise structure of full bodyweight support on the toes and hands. These adjustments allow practitioners to target specific upper body regions more effectively, with electromyographic (EMG) studies demonstrating shifts in activation patterns across the pectoralis major, triceps brachii, and supporting musculature. For instance, wider hand positions tend to increase the range of shoulder abduction, placing greater load on the chest, whereas narrower placements heighten elbow extension demands on the triceps. In wide-grip push-ups, the hands are positioned beyond shoulder width, typically 1.5 to 2 times the biacromial distance apart, which elongates the and increases its involvement during the pressing phase. This variation promotes greater horizontal adduction of the s compared to standard positioning, though EMG data indicate that overall pectoralis remains comparable to grips, with a subtle shift toward the outer chest fibers. abduction is amplified, potentially recruiting more deltoid assistance, but care must be taken to avoid excessive flaring that could strain the . Narrow-grip push-ups, often called diamond push-ups, feature the hands placed close together directly under the chest, with the thumbs and index fingers touching to form a shape. This configuration significantly elevates brachii activation, as the narrower base of support reduces the for the chest and shifts emphasis to elbow extension. Studies using surface EMG have shown significantly higher activity in this variation compared to standard or wide grips, making it particularly effective for development while still engaging the to a lesser degree. Knuckle push-ups replace palm contact with the distal knuckles of the index and middle fingers, slightly elevating the upper body and extending the by approximately 2-5 cm. This setup strengthens the wrist extensors and muscles, including the flexor carpi radialis and extensor digitorum, by maintaining a neutral alignment under load. EMG comparisons among trained individuals reveal heightened activation in stabilizers during knuckle variations relative to traditional push-ups, supporting its role in conditioning for activities requiring strong support. Hand-release push-ups incorporate a pause at the bottom position where the hands are lifted , fully extending the chest to the floor and eliminating to demand a more powerful concentric . By isolating the upward phase without assistance from the floor, this variation enhances production in the prime movers, with biomechanical analyses indicating increased peak power output during the ascent compared to continuous-contact push-ups. Grip changes like these result in targeted muscle shifts, as explored in the primary muscle groups section.

Advanced and Plyometric Forms

Advanced push-up variations elevate the exercise's intensity by incorporating unilateral loading, extreme body positioning, or explosive dynamics, targeting enhanced strength, stability, and power output. The one-arm push-up exemplifies a unilateral challenge, where the supporting bears approximately 100% of the body weight while maintaining a straight body line to prevent rotation or sagging. This demands exceptional , , and strength, with progressions starting from assisted forms such as elevated one-arm push-ups on a bench or archer push-ups—where one extends while the other bends, emphasizing unilateral loading—to gradually build capacity and mitigate injury risk. Archer push-ups help develop the strength needed for full one-arm push-ups by allowing partial weight distribution between arms. The planche push-up further intensifies demands through a pronounced forward lean, positioning the body parallel to the ground in a planche hold before performing the push-up motion, which requires extreme core engagement for anti-extension and shoulder protraction to sustain the lever-like position. This variation heavily recruits the anterior deltoids, pectorals, and serratus anterior, fostering straight-arm strength essential for advanced while challenging overall body control. Similarly, the guillotine push-up emphasizes extended by using a wide hand grip and lowering the chest toward the neck area, stretching the upper pectorals beyond standard depth to promote greater muscle activation and in the clavicular head. Plyometric forms introduce dynamic elements to build explosive power, with clap push-ups involving a rapid concentric push that propels the hands off the ground for a mid-air clap before softly to absorb . This leverages the stretch-shortening cycle to enhance neuromuscular coordination and rate of force development, recruiting fast-twitch fibers for improved neural efficiency. In athletic training, such variations translate to better performance in sports requiring upper-body explosiveness, like throwing or disciplines, by increasing power output without external loads. Hindu push-ups add a fluid, yoga-inspired dimension, flowing from a position through a sweeping arch to an upward , which improves mobility, spinal flexibility, and across the chest, shoulders, and back. For further escalation, weighted vests or packs can be added to these forms, amplifying resistance while preserving technique. Another advanced variation, the spiderman push-up, incorporates a knee drive to the during the descent, enhancing oblique and hip flexor engagement for improved .

Records and Cultural Impact

World Records

The world records for push-ups highlight extraordinary human endurance, speed, and strength, particularly in categories emphasizing volume over extended periods, rapid execution, and unilateral variations. These achievements are officially recognized by , which maintains strict criteria to ensure authenticity and safety.
CategoryRecord HolderAchievementDate and LocationSource
Most push-ups in 24 hours (male)Charles Servizio (USA)46,00124-25 April 1993, , USAGuinness World Records
Most non-stop push-upsMinoru Yoshida ()10,507October 1980, RecordHolders.org
Most push-ups in one hour (male)Pop Laurentiu ()3,37830 June 2023, , Guinness World Records
Most one-arm push-ups in one hour (male)Paddy Doyle ()1,86827 November 1993, , Vercalendario
Guinness World Records verifies these feats through rigorous standards, including continuous video documentation, independent witnesses, and adherence to precise form: the body must remain rigid in a straight line, the chest must touch the ground or a designated surface on descent, arms must fully extend at the top, and no resting or improper movements are allowed. These protocols prevent and ensure the of the , with medical supervision often required for attempts exceeding several hours.

Cultural and Competitive Significance

The push-up holds a prominent place in military training programs worldwide, serving as a staple assessment of upper-body strength and endurance. In the United States Army, push-ups have been a core component of tests since 1980, with soldiers required to perform a maximum number of repetitions within two minutes as part of the Army Physical Fitness Test (APFT); for instance, standards demanded at least 42 reps for younger male soldiers (aged 17-21) to achieve the minimum passing score of 60 points, emphasizing combat readiness. The current Army Fitness Test (AFT), implemented in 2025, continues this tradition through the hand-release push-up event, where minimum passing requires at least 10-20 reps depending on age and gender, integrated into boot camps and ongoing physical training to build resilience for operational demands. In competitive fitness arenas, push-ups feature prominently in events that test endurance and skill. incorporates push-ups as a foundational movement in workouts and the annual , where variations like handstand push-ups appear in high-stakes routines, such as completing sets of 10-30 reps unbroken during quarterfinals or finals to evaluate overall athletic capacity. competitions, such as the Calisthenics Cup, include push-ups in endurance challenges; for example, male qualifiers must perform 100 push-ups as part of a routine combining squats and muscle-ups, highlighting the exercise's role in bodyweight strength contests. Programs like the Hundred Pushups Training Plan further popularize competitive preparation, offering a six-week progression to achieve 100 consecutive reps, adopted by enthusiasts worldwide for personal benchmarks. Push-ups have permeated , symbolizing grit and transformation in media and social campaigns. Iconic film depictions include the training montages in the series, where protagonist performs one-armed push-ups on the ring canvas, underscoring themes of perseverance amid makeshift workouts. In the 2010s, celebrity-driven social media trends amplified their visibility, notably the #22PushupChallenge launched in 2016 to raise awareness of veteran suicides (22 daily in the U.S.), with stars like , , and posting videos of 22 reps and nominating others, amassing millions of shares. This viral format echoed broader fitness challenges on platforms like , blending entertainment with motivation. Globally, push-ups adapt to cultural contexts in training traditions. In Asian such as kung fu and , knuckle push-ups—performed on the first two knuckles—strengthen wrists and fists for striking, a practice emphasized in conditioning drills to enhance punching power and endurance. European gymnastics history integrates push-ups as foundational bodyweight exercises dating to the , within Swedish and systems that influenced modern , promoting equilibrium and strength through variations like planche push-ups in artistic routines.

Biological Comparisons

Push-up Like Movements in Animals

In quadrupedal mammals such as and , play behaviors often involve forelimb extensions resembling push-up motions to facilitate bounding and social interaction. During play bows, lower their chest to the ground while extending and pressing the s forward, engaging the and extensors to propel the body upward in a bounding motion, which strengthens and initiates chase or wrestle play. Among , chimpanzees employ as a primary form of , where the forelimbs undergo repeated protraction and extension phases that provide propulsive support akin to one-armed push-up variants. In this gait, the wrists flex to contact the ground with the knuckles, followed by elbow and extension to lift and advance the body, with electromyographic studies showing activation of shoulder muscles including the deltoid during the support phase to stabilize and propel the trunk. This motion supports brachiation transitions by maintaining forelimb strength for suspensory postures. Reptiles and display push-up-like behaviors primarily for signaling rather than . Male anole ( ) perform rapid push-up displays, extending and retracting their s while bobbing the head and to signal territory and attract mates, with the limb extensions serving as honest indicators of and dominance. These displays involve pectoral and forelimb muscle contractions that elevate the body, paralleling the biomechanical demands of human push-ups but adapted for in highly territorial . Biomechanically, these animal movements share parallels with push-ups through protraction, where the rotates forward and the flexes, activating analogous muscles like the anterior deltoid to generate upward force against the . In chimpanzees, deltoid engagement during protraction in mirrors the deltoid's role in push-up ascent, providing trunk support and propulsion via similar torque patterns. Such parallels highlight conserved locomotor across , with evolutionary links explored further in related perspectives.

Evolutionary Perspectives

The push-up movement traces its ancestral origins to the forelimb-dominated locomotion of early primates, where arboreal environments demanded pushing and pulling actions against branches and trunks for climbing and suspension. In non-human primates, such forelimb use facilitated versatile propulsion in three-dimensional spaces, with arboreal species exhibiting greater joint mobility than their terrestrial counterparts to support these dynamic presses. As hominins transitioned to bipedalism around 6-7 million years ago, this primate heritage persisted, freeing the forelimbs from primary locomotor roles while retaining pushing capabilities adapted for manipulation and support during upright posture shifts. This evolutionary legacy provided adaptive advantages through enhanced upper-body strength, essential for hunting and gathering activities that involved pressing motions, such as thrusting tools or subduing prey. In , emerging around 1.8 million years ago, robust humeral morphology indicates significant upper-limb loading capacities, likely supporting the forceful and wielding of stone tools for processing resources. Such strength not only aided survival in resource-scarce environments but also correlated with pressures, where upper-body power influenced mate competition and provisioning success among early hominins. Comparatively, scapular diverges from that of quadrupeds, featuring a more dorsally positioned and rotund that enables extensive mobility across multiple planes, unlike the sagittally restricted shoulders of terrestrial quadrupeds optimized for stability. This enhanced scapulohumeral range in humans and other bipedal allows for efficient force generation during pressing actions, underpinning the biomechanical efficacy of push-up-like exercises by maximizing pectoral and deltoid engagement without compromising postural integrity. In contrast, quadrupedal locomotion limits such versatility, prioritizing endurance over explosive pushes. In modern contexts, push-ups serve as a simulation of these prehistoric pressing motions, enabling researchers to reconstruct ancestral levels by quantifying upper-body outputs akin to those required for use and . This approach highlights push-ups' role in bridging evolutionary history with contemporary , without direct analogs in non-human animal behaviors.

References

  1. [1]
    The rise of push-ups: A classic exercise that can help you get stronger
    Feb 17, 2019 · The push-up engages your body from top to bottom. It works several muscle groups at once: the arms, chest, abdomen (core), hips, and legs.Missing: origin history
  2. [2]
    What Muscles Do Push-Ups Work? - Healthline
    Since the inception of modern strength training, the push-up has remained one of the top bodyweight exercises for targeting muscles of the upper body.Muscles worked · Benefits · Form guide · Variations
  3. [3]
    Health Benefits of Push-Ups - WebMD
    Jul 12, 2025 · Chest. Arms; Stomach muscles (abdominals); Hips‌; Legs. Push-ups are ...About Push-Ups · How To Do A Push-Up · Push-Up Variations
  4. [4]
    History of Push-Ups: They've Been Around Longer Than You Think!
    Sep 2, 2018 · The history of the push-up dates back thousands of years to India where warriors used this somewhat strenuous exercise to get stronger and into fighting shape.
  5. [5]
    Association Between Push-up Exercise Capacity and Future ... - NIH
    Feb 15, 2019 · This cohort study investigates the association between participants' capacity to complete push-ups at baseline and cardiovascular disease events during 10 ...Missing: origin | Show results with:origin
  6. [6]
  7. [7]
    Push-up - Etymology, Origin & Meaning
    Pushup, from 1660s verbal phrase push + up, originally a physical exercise on bars (1893); also adjective meaning "that pushes up," used for bras since ...
  8. [8]
    PUSH-UP Definition & Meaning - Dictionary.com
    Push-up definition: an exercise in which a person, keeping a prone position ... Origin of push-up. First recorded in 1905–10; noun use of verb phrase ...
  9. [9]
    press-up, n. meanings, etymology and more
    The earliest known use of the noun press-up is in the 1920s. OED's earliest evidence for press-up is from 1928, in the Times (London). press-up is formed within ...
  10. [10]
    [PDF] The History of the US Army's Physical Fitness Testing
    Apr 16, 2021 · “The Army Reserve Training Corps (1920)” became the Army training manual for combat readiness at this time. In writing this book, the author ...
  11. [11]
    Dand, Daṇḍ: 6 definitions
    Oct 11, 2024 · Daṇḍ (दण्ड्).—1 U. (daṇḍayati-te, daṇḍita) To punish, fine, chastise; (this is one of the 16 roots that govern two accusatives); तान् सहस्रं च दण्डयेत् ( ...
  12. [12]
    How the Ancient World Lifted Weights - BarBend
    Aug 12, 2019 · This article examines the history of resistance training in the ancient world, specifically that found in China, Egypt, Indian and Greece.Ancient China · Ancient India · Ancient Greece
  13. [13]
    [PDF] A Study of Nineteenth- Century Swedish Medical Gymnastics
    In this case, I have examined the bequeathed writings of Per Henrik Ling and his successors in the field of medical gymnastics, Lars Gabriel Branting, Truls ...Missing: Pehr | Show results with:Pehr
  14. [14]
    History of the APFT - Mountain Tactical Institute
    Apr 30, 2015 · First Army fitness test developed in 1920 following WWI; Mandatory, Army-wide fitness testing began in 1963; By 1973 the Army was suffering from ...
  15. [15]
    The Incredible Fitness and Success of Jack LaLanne - James Clear
    At age 42, he set the world record for pushups by doing over 1,000 in 23 minutes. ... Wake up at 4am (in his later years, LaLanne “slept in” until 5am). Look at ...
  16. [16]
    The Federal Government Takes on Physical Fitness | JFK Library
    Dec 15, 2021 · But concern about fitness peaked in the mid-1950s with publication of an international study that found American children far less fit than ...
  17. [17]
    Exercise Library:Push-up
    ### Push-Up Instructions (ACE Fitness)
  18. [18]
    What is the force on the arms in a pushup? - Physics Stack Exchange
    Apr 26, 2011 · The moment generated is 90% x ((4.5/9) body lengths) = 45 (% x BL). At the lowest point in the pushup, your chest muscles are generating enough ...Biomechanics statics of an push-up [closed] - Physics Stack Exchangenewtonian mechanics - Physics of the PlancheMore results from physics.stackexchange.com
  19. [19]
    The Effect of Position on the Percentage of Body Mass Supported ...
    Suprak, DN, Dawes, J, and Stephenson, MD. The effect of position on the percentage of body mass supported during traditional and modified push-up variants.
  20. [20]
  21. [21]
    Shoulder electromyography activity during push-up variations - NIH
    The standard push-up (PU) is a common closed chain exercise used to improve dynamic stability of the upper extremity through enhancement of proprioception, ...
  22. [22]
    Push Up Anatomy | Muscle&Motion - Muscle and Motion
    Jan 21, 2012 · The target muscle for push-ups is the Pectoralis Major. Synergists include Anterior Deltoid, Triceps Brachii, Serratus Anterior, ...
  23. [23]
    Pushups - Physiopedia
    According to a 2020 study on shoulder electromyography activity during push-up variations, the triceps and pectoralis major muscles had the highest ...Introduction · Muscles Involved · Physiotherapy · Variations
  24. [24]
    What Muscles Do Push-Ups Work? Here's a Breakdown - Peloton
    Aug 29, 2025 · Push-ups work the core, chest, triceps, shoulders, upper back (lats, traps, rhomboids), and core muscles like rectus abdominis.<|separator|>
  25. [25]
    comparison of muscle activation using various hand positions during ...
    This study examined whether different levels of electromyographic (EMG) activity in the pectoralis major and triceps brachii muscles are required to perform ...
  26. [26]
    Selective Activation of Shoulder, Trunk, and Arm Muscles - NIH
    The push-up is a widely used exercise for upper limb strengthening that can be performed with many variants. A comprehensive analysis of muscle activation ...Missing: origin | Show results with:origin
  27. [27]
    Muscle Activation of Upper Body in Different-Angle Suspension ...
    Feb 26, 2025 · During push-ups, electromyography (EMG) data from the pectoralis major (PM), anterior deltoids (AD), triceps brachii (TRI), upper trapezius (UT) ...
  28. [28]
    What is Muscle Action? Concentric, Eccentric, and Isometric - NASM
    Concentric contraction occurs, for example, during the upward thrust in the bench press or squat, while eccentric contraction occurs during the downward phase.
  29. [29]
    Push-Up Benefits and the 7 Muscles They Work - Verywell Health
    Sep 8, 2025 · The anterior deltoids help your shoulders maintain a flexed position during the push-up.
  30. [30]
    Core Muscle Activity during Physical Fitness Exercises: A Systematic ...
    The aim of this study was to systematically review the current literature on the electromyographic (EMG) activity of six core muscles (the rectus abdominis, ...
  31. [31]
    Muscle Activation during Push-Ups with Different Suspension ... - NIH
    The purpose of this study was to analyze upper extremity and core muscle activation when performing push-ups with different suspension devices.
  32. [32]
    Shoulder muscle EMG activity during push up variations on and off a ...
    Surface electromyograms were recorded from the triceps, pectoralis major, latissimus dorsi, rectus abdominis and external oblique while performing push up ...
  33. [33]
    SHOULDER KINEMATICS DURING THE PUSH-UP PLUS EXERCISE
    The push-up plus exercise is often modified from a standard push-up plus to be performed against a wall in the early stages of shoulder rehabilitation to limit ...
  34. [34]
    Scapular Kinematics and Shoulder Elevation in a Traditional Push-Up
    Performing a traditional push-up with the shoulders elevated may place the scapula in a position of impingement.Missing: peer- | Show results with:peer-
  35. [35]
    Effect of Progressive Calisthenic Push-up Training on Muscle ...
    The purpose of this study was to determine whether progressive calisthenic push-up training (PUSH) is comparable with traditional bench press training (BENCH) ...
  36. [36]
    Push-up Exercise Capacity and Future Cardiovascular Events in ...
    Feb 15, 2019 · This study found that push-up capacity was inversely associated with 10-year risk of CVD events among men aged 21 to 66 years. Thus, push-up ...
  37. [37]
    Association Between Push-up Exercise Capacity and Future ...
    Feb 1, 2019 · Conclusions and relevance: The findings suggest that higher baseline push-up capacity is associated with a lower incidence of CVD events.Missing: scientific | Show results with:scientific
  38. [38]
    Resistance training is medicine: effects of strength training on health
    Benefits of resistance training include improved physical performance, movement control, walking speed, functional independence, cognitive abilities, and self- ...Missing: calisthenics push- ups<|separator|>
  39. [39]
    Dose-dependent associations of joint aerobic and muscle ...
    Irrespective of the aerobic exercise level, the addition of muscle-strengthening exercise was associated with a lower obesity prevalence. •. A physical activity ...
  40. [40]
    A COMPREHENSIVE REHABILITATION PROGRAM FOR ... - NIH
    Because patients often present with varying levels of elbow irritability, two warm-up options, treadmill and upper body ergometer (UBE), are recommended.
  41. [41]
    Optimal management of shoulder impingement syndrome - PMC
    Dynamic stabilization drills such as rhythm stabilization performed in a functional position, push-ups on a ball, and ball throws are performed to improve ...
  42. [42]
    Core exercises: Why you should strengthen your core muscles
    Strengthening your core muscles may help back pain get better and lower the risk of falls. Core exercises can help you reach your fitness goals. Aerobic ...Missing: engagement | Show results with:engagement
  43. [43]
    Pushup Modifications and Alternatives to Alleviate Wrist Pain
    Nov 23, 2020 · What are pushup modifications you might try if you have wrist pain? · Pushup bar with a static hold · Pushup with dumbbells or kettlebells.Pushup modifications · Pushup alternatives · Wrist stretches · Pain managementMissing: prevention | Show results with:prevention
  44. [44]
    (PDF) The Biomechanics of the Push-up - Academia.edu
    Knee push-ups reduce bodyweight loading to 54% at the top and 62% at the bottom position. Explosive variations like countermovement push-ups maximize peak force ...
  45. [45]
  46. [46]
    Comparison of muscle activation using various hand positions ...
    Comparison of muscle activation using various hand positions during the push-up exercise. J Strength Cond Res. 2005 Aug;19(3):628-33. doi: 10.1519/15094.1 ...Missing: findings | Show results with:findings
  47. [47]
    From Popular Simple Exercises to Valid Tests for Upper-Body Power
    The purpose of this study was to assess the logical and ecological validity of 5 explosive push-up variations as a means of upper-body power assessment.Missing: hand- release
  48. [48]
    Four Progressions to Conquer the One-Arm Pushup | StrongFirst
    Oct 27, 2023 · You'll train four days per week for six weeks. Each week has one heavy day, two medium days, and one light day.
  49. [49]
  50. [50]
    How to Do Planche Push-Ups With Perfect Form - 2025 - MasterClass
    Sep 9, 2021 · Squeeze your chest and shoulders as your upper body shifts forward. 6. Engage your core while maintaining full-body tension and lift your feet ...
  51. [51]
    Build a Big Pec Shelf With the Guillotine Press - BarBend
    Nov 27, 2024 · When looking to target the upper pecs with the Guillotine press, first aim to unlock the fullest range of motion with a 3 to 4-second eccentric ...<|control11|><|separator|>
  52. [52]
    How Plyometric Push-Ups Can Improve Pressing Strength | BarBend
    Jun 29, 2024 · The plyometric push-up can be used with strength, power, and fitness athletes to increase upper body pressing strength and explosiveness.
  53. [53]
    CURRENT CONCEPTS OF PLYOMETRIC EXERCISE - PMC - NIH
    Explosive plyometric exercises may improve the neural efficiency through enhancement of neuromuscular coordination. Therefore, plyometric training increases ...
  54. [54]
    How To Do Hindu Pushups To Build Your Upper Body Strength
    Jun 15, 2018 · The main benefit of the Hindu pushup isn't about strength, though. “It's more likely to work your mobility and flexibility,” says Stokes. “And ...
  55. [55]
    Push-Up Progressions | NASM Guide to Pushups Part 3
    Hand/foot placement: A slightly wider foot stance will be easier than hip-width. Hands shoulder width apart will be more stable than staggered.Missing: technique | Show results with:technique
  56. [56]
    Army introduces new fitness test for 2025 | Article | The United States ...
    May 14, 2025 · The AFT is a five-event assessment, including the three-repetition maximum deadlift, hand-release push-up, sprint-drag-carry, plank and two-mile ...
  57. [57]
    The Push-Up - CrossFit
    The push-up was largely regarded as a measure of a person's strength and fitness. In more modern times, much of this reputation has been passed on to the bench ...
  58. [58]
    Calisthenics Cup Endurance Qualification Routine 2025
    Oct 16, 2024 · Push-Ups: 100 reps; Bodyweight Squats: 30 reps; Unbroken Muscle-Ups: 5 reps. Calisthenics Cup Endurance Routine Male. Female Qualification ...<|separator|>
  59. [59]
    The Hundred Pushups Training Program
    Web site One Hundred Push Ups details a training plan for going from 0 to 100 push-ups in a matter of six weeks. ... Hundred Pushups is the ultimate training ...Week 1 · Week 3 · Week 5 · Book
  60. [60]
    The Complete Library of Rocky Training Exercises
    Nov 20, 2020 · Below we break down every single exercise from the training montages featured in the first, second, third, fourth, and sixth Rocky installments.
  61. [61]
    22 Pushup Challenge: New Celeb Viral Social Media Campaign
    Aug 16, 2016 · The #22PushupChallenge is the latest celebrity viral campaign aimed at spreading awareness for veterans suffering from mental illness and ...
  62. [62]
    Knuckle Push-Ups - Black Belt Wiki
    Traditional martial arts instructors utilize knuckle push-up in order to strengthen a student's fist and wrist as well as build triceps, chest and arm strength.
  63. [63]
    Kissing the Ground in Equilibrium—A Search for the Origins of the ...
    Jan 12, 2023 · The earliest reference the OED cited for the term pushup was from 1896, but like my new friend Denis, I also believed pushups had a much longer ...
  64. [64]
    Dogs at Play: How and Why Bowing Came to Mean "Let's Play"
    Mar 5, 2025 · Bowing by stretching the forelimbs evolved to become a highly ritualized signal for inviting dogs to play, have fun, or continue to play if ...Missing: push- | Show results with:push-
  65. [65]
    What Does It Mean When My Dog Bows? - The Spruce Pets
    Jun 17, 2025 · The play bow is when a dog stretches his front legs out in front, leaning down on his elbows. This brings the chest low to the ground as if the ...Missing: forelimb push-
  66. [66]
    Feline Locomotive Behavior | Veterian Key
    Dec 22, 2016 · Motor responses continue to develop, so by day 30 (27 to 30 mm), flexion of the carpus and slight activation of the masticatory muscles occur.Missing: bounding | Show results with:bounding
  67. [67]
    Kinematic and dynamic aspects of chimpanzee knuckle walking
    Oct 6, 2021 · Chimpanzees are knuckle walkers, with forelimbs contacting the ground by the dorsum of the finger's middle phalanges.
  68. [68]
    EMG of chimpanzee shoulder muscles during knuckle-walking
    Aug 7, 2025 · Chimpanzees are characterized by a quadrupedal gait in which the hind limb overstrides the ipsilateral forelimb. Forelimb position in the plane ...
  69. [69]
    Mechanical implications of chimpanzee positional behavior - ADS
    Large muscles that retract the humerus (latissimus dorsi and probably sternocostal pectoralis major and posterior deltoid) and flex the elbow (biceps brachii, ...
  70. [70]
    Why Do Lizards Do Push-Ups? - Live Science
    Oct 24, 2012 · "Anoles are highly visual species, so in that sense it's not surprising that they would use visual displays to mark territory," Ord said. "Still ...
  71. [71]
    Jamaican lizards mark their territory with shows of strength at dusk ...
    Aug 27, 2008 · That's according to a new study showing that male Anolis lizards engage in impressive displays of reptilian strength – push-ups, head bobs, and ...
  72. [72]
    Lizard push-ups grab attention - Science News
    Nov 24, 2008 · These yellow-chinned anoles stake territorial claims by bobbing their heads and inflating a colorful pouch on their necks, usually while perched ...
  73. [73]
    Development of a comparative chimpanzee musculoskeletal ...
    The purpose of this study was to develop a novel biomechanical and comparative chimpanzee glenohumeral model, designed to parallel an existing human ...
  74. [74]
    Exploring the functional morphology of the Gorilla shoulder through ...
    Feb 24, 2021 · In this study, we test the hypothesis that the musculoskeletal configuration of the shoulder in gorillas improves the biomechanical performance ...Missing: push- | Show results with:push-
  75. [75]
    Downclimbing and the evolution of ape forelimb morphologies - PMC
    Sep 6, 2023 · We found that the shoulders and elbows of chimpanzees and sooty mangabeys subtended larger joint angles during bouts of downclimbing.
  76. [76]
    Primate Locomotion | Learn Science at Scitable - Nature
    Primate locomotion is the study of movements and postures in arboreal and terrestrial environments. Primates are masters of life in the trees.
  77. [77]
    The morphology and evolutionary history of the glenohumeral joint ...
    As such, terrestrial quadrupedal primates have the most stable glenohumeral joints, thus the least mobile. The proximal humerus and glenoid cavity are more ...
  78. [78]
    Morphology and structure of Homo erectus humeri ... - PubMed Central
    Jan 19, 2018 · The current study reports the first humeral rigidity and strength properties of East Asian H. erectus and places its diaphyseal robusticity into broader ...
  79. [79]
    Homo erectus | The Smithsonian Institution's Human Origins Program
    Jan 3, 2024 · Known as the Acheulean stone tool industry, it consisted of the creation of large cutting tools like handaxes and cleavers. Increased ...Missing: upper strength
  80. [80]
    Genetic Influences on the Development of Grip Strength in ... - NIH
    From a functional evolutionary perspective, enhanced upper-body strength was likely shaped by recurrent male-male competition for mates in our hominid ancestors ...
  81. [81]
  82. [82]
    Scapular morphology of great apes and humans: A three ...
    Oct 27, 2022 · In great apes and humans, the scapula is positioned dorsally on the thorax, providing the forelimb with a large range of motion during arboreal ...
  83. [83]
    Human athletic paleobiology; using sport as a model to investigate ...
    Here, we draw from examples in the literature to provide a review of how the use of athletes as a model system is enhancing understanding of human evolutionary ...