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Common nighthawk

The Common nighthawk (Chordeiles minor) is a medium-sized, crepuscular bird in the family Caprimulgidae, characterized by its cryptic mottled grayish-brown , long pointed wings with white wing bars, a notched , large eyes, and a wide gape for capturing flying . It measures 22–24 cm in length, weighs 65–98 g, and is renowned for its erratic, bat-like flight during dawn and dusk foraging, as well as its distinctive "peent" call and booming aerial courtship dives produced by flexing its wing primaries. Native to the , the Common nighthawk breeds across much of and —from to —and migrates to winter in the lowlands of central and southern , undertaking journeys of 4,000–11,000 km, with occasional vagrants recorded in . It inhabits diverse open landscapes, including boreal forests, grasslands, savannas, woodlands, coastal dunes, and urban areas with flat gravel roofs or bare ground for nesting, often roosting on branches, posts, or rooftops during the day. The species is primarily aerial insectivorous, feeding on flying , , moths, and other caught in mid-air, and it may drink by skimming surfaces. Breeding occurs from May to August in northern populations, with pairs laying two eggs directly on the ground without a nest; incubation lasts 18–20 days, and fledglings become independent after 45–50 days, though southern populations may raise a second brood. Males perform dramatic dives to attract females, producing a hollow booming sound audible up to a kilometer away. Solitary nesters, they are territorial during breeding but form loose flocks during . Globally listed as Least Concern by the IUCN, Common nighthawk populations have declined significantly in some regions, with a 68% drop in southern from to and a 12% decrease from 2005 to in southern , attributed to habitat loss, use reducing prey, , and urban development favoring smooth roofing materials over gravel. In , it is designated as Special Concern under the Species at Risk Act (updated from Threatened in 2007 as of ), with an estimated 270,000–900,000 adults in remaining as of , highlighting the need for ongoing monitoring and conservation efforts.

Taxonomy and nomenclature

Etymology

The scientific name of the common nighthawk, Chordeiles minor, originates from New Latin roots derived from Ancient Greek. The genus name Chordeiles is an irregular formation from chordē (string of a lyre or harp) and deilē (afternoon or evening), alluding to the bird's distinctive cry resembling a musical string sounded in the evening. The specific epithet minor is Latin for "smaller," reflecting its relatively diminutive size compared to other nightjars in the Caprimulgidae family. The common English name "nighthawk" dates to the early 17th century, first appearing in the 1611 King James Bible as a term for the before being applied to species like Chordeiles minor; it combines "night" for the bird's crepuscular and nocturnal activity with "" due to its swift, erratic flight resembling that of a hawk. Another widespread colloquial name, "bullbat," emerged in the 19th century and refers to the bird's bat-like silhouette in flight and the booming, bull-like sound produced by air rushing through specialized wing feathers during courtship dives. Regional North American names include "pisk," an onomatopoeic term of uncertain origin mimicking the bird's nasal call, particularly used in parts of the . Culturally, the common nighthawk contributed to Nebraska's early state nickname "Bugeater," adopted in the late 19th century to describe the bird's voracious insect-eating habits, which were seen as beneficial during times of agricultural hardship in the region. In North American folklore, the species was often confused with the Eastern whip-poor-will (Antrostomus vociferus) by 18th- and 19th-century naturalists and locals due to overlapping habitats, similar nocturnal behaviors, and comparable calls, leading to shared myths such as omens of death or supernatural associations.

Classification and subspecies

The common nighthawk (Chordeiles minor) is classified within the Caprimulgidae, known as nightjars or goatsuckers, which comprises crepuscular and nocturnal insectivorous birds characterized by their cryptic plumage and wide gape for aerial foraging. Within this family, it belongs to the genus Chordeiles, which includes three extant of nighthawks native to the ; the common nighthawk is the sister species to the Antillean nighthawk (C. gundlachii), with the (C. acutipennis) as the outgroup, based on molecular phylogenetic analyses. The species has been recognized as a distinct entity, Chordeiles minor, since the early 19th century, following its initial scientific description by in 1771 as Caprimulgus minor and subsequent placement in the Chordeiles by in 1827. Early taxonomic treatments often lumped or split it from other nightjars, such as the eastern whip-poor-will (Antrostomus vociferus), due to superficial similarities in nocturnal habits, but 20th-century revisions based on and vocalizations established its separation. More recent genetic studies in the , including sequencing, have confirmed the monophyly of the Chordeiles while suggesting potential within the nighthawk Chordeilinae, prompting ongoing taxonomic reviews. Nine subspecies of the common nighthawk are currently recognized, primarily distinguished by variations in body size, wing length, and tone (ranging from paler, grayer forms in arid regions to darker, more mottled ones in humid areas), as defined through morphometric analyses in the early by ornithologists like Harry C. Oberholser. These subspecies reflect clinal variation across the broad range from to southern . The following table summarizes the recognized , their primary geographic distributions, and key distinguishing traits:
SubspeciesGeographic RangeDistinguishing Traits
C. m. minorEastern (from Newfoundland to and west to )Nominate form; medium-sized (wing 200–210 mm); darker dorsal plumage with buff mottling; breast tawny.
C. m. hesperisWestern (southwestern to and )Larger (wing 215–225 mm); paler overall, with grayer tones adapted to arid habitats; reduced tawny on underparts.
C. m. sennettiSouth-central U.S. () to northeast Intermediate size (wing 205–215 mm); slightly paler than minor with more extensive white on throat; found in coastal plains.
C. m. howelliSouthwest U.S. (, ) to central Smaller (wing 190–200 mm); darker with denser barring on underparts; breeds in open woodlands.
C. m. henryiSouthwest U.S. () to north-central Similar to hesperis but smaller (wing 195–205 mm); pale grayish plumage with fine vermiculation.
C. m. aserriensis (highlands)Small (wing 185–195 mm); dark with rich buff tones on upperparts; restricted to montane areas.
C. m. panamensis (lowlands)Medium-small (wing 190–200 mm); paler ventral barring and lighter overall tone suited to Pacific slope.
C. m. neotropicalisNorthern ( to northern )Larger (wing 210–220 mm); very dark dorsum with heavy blackish mottling; adapted to humid forests.
C. m. rufescensNorthern and Medium (wing 200–210 mm); reddish-buff tones on underparts, distinguishing it from neighboring forms.

Physical characteristics

Morphology and plumage

The common nighthawk (Chordeiles minor) is a medium-sized measuring 22–24 in length, with a of 53–57 and a body mass ranging from 65–98 g. It possesses a slender body, long pointed wings that extend beyond the at rest, and a forked that aids in agile aerial maneuvers. These features contribute to its distinctive bounding flight pattern, optimized for pursuing in open airspace. The plumage is cryptically mottled in shades of gray, brown, and black, providing effective against ground substrates and bark during roosting. In flight, a prominent bar on the primaries becomes visible, serving as a flash marker, while males display a V-shaped throat patch during displays. is minimal, though males exhibit more pronounced wing patches and a subterminal tail band, whereas females have smaller patches, a throat, and lack the tail band. Juveniles appear duller overall, with less contrast in their patterning. show subtle variations, such as browner tones in eastern populations and grayer hues in northern ones. Key adaptations include large eyes equipped with a layer to enhance low-light for crepuscular . The small bill opens to a wide gape fringed by bristle-like rictal feathers, which help detect and capture flying . The adopts a horizontal perching posture, aligning parallel to branches or the ground to minimize visibility.

Molt and variation

The common nighthawk follows a complex basic molt strategy, undergoing a single annual prebasic molt without a prealternate molt. This complete molt occurs primarily on the wintering grounds in from September to February, with replacement often peaking in January and February. are replaced sequentially, typically beginning with the innermost primaries and progressing outward, which preserves aerodynamic integrity and flight ability throughout the process. Some individuals exhibit a continuous molt-migration pattern, initiating or completing portions of the molt during southward travel. Juveniles possess paler, less defined than adults, featuring reduced black markings on the scapulars, finer mottling, and an indistinct or absent patch, often with buffy tones enhancing . By the first winter, through the initial prebasic molt, they attain adult-like feather structure and patterns, including broader primaries and stronger contour feathers. Adult plumage shows seasonal wear, becoming faded and frayed toward the end of the season as feathers abrade, prompting the prebasic molt for renewal. Individual variation is minimal beyond subtle differences in wear or freshness, with no significant geographic distinctions outside of subspecies-specific tones. This molt schedule aligns with the ' migratory cycle, ensuring arrival at northern grounds with pristine feathers that support cryptic concealment and enhance the visual and acoustic impact of displays, such as the male's booming aerial dives.

Range and habitat

Geographic distribution

The common nighthawk (Chordeiles minor) has a broad breeding distribution across , extending from western and central and much of southward through the to , including local breeding populations in interior and Pacific slope regions from and , Mexico, to central . This range encompasses diverse open landscapes and has historically expanded into urban environments, such as cities with flat gravel roofs suitable for nesting, particularly since the early 1900s. The species is absent from dense forested regions but is widespread in suitable open areas within its continental limits. During the nonbreeding season, the common nighthawk winters primarily in , with its range spanning the lowlands east of the from and through the , eastern , , , central (including and regions), , , and northern , and occasionally extending to . While primarily migratory, some populations in the are year-round residents. Vagrant common nighthawks have been recorded outside their regular ranges, including in and various European locations such as , , the , the , and the , likely due to the species' extensive routes. Recent monitoring indicates relative stability in the overall breeding distribution, though population densities have shown regional variations, with no major documented shifts in range boundaries as of the latest assessments.

Habitat requirements

The common nighthawk (Chordeiles minor) prefers open or semi-open habitats for breeding, including forest clearings, woodland edges, logged or burned areas, grasslands, prairies, forests, sagebrush flats, rocky outcrops, and coastal sand dunes. These sites provide bare or sparsely vegetated ground for nesting, often near scattered cover such as logs, boulders, shrubs, or debris, which offers and protection from predators. In urban environments, the species has adapted to flat gravel rooftops, which mimic natural gravel or sandy substrates and support nesting in cities where traditional habitats are scarce. For roosting, common nighthawks select shaded, horizontal surfaces in open areas, including branches, fence posts, the ground, or flat building roofs, where their cryptic provides effective concealment during daytime rest. Males may roost communally near sites, often on snags or low branches in proximity to open areas, to minimize energy expenditure and predation risk. Foraging occurs primarily in aerial microhabitats above treetops, open fields, river valleys, marshes, or urban streets, targeting swarms of flying at and ; the avoids dense canopies that obstruct flight and insect access. These areas are typically within 500 feet of the ground or water surfaces, with concentrations near artificial lights in urban settings that attract prey. Habitat loss from and has reduced available natural edges and open grounds, contributing to localized declines in breeding success, though urban roofs have partially offset this in some regions.

Migration patterns

The common nighthawk undertakes one of the longest migrations among , traveling 4,000–11,000 (2,500–6,800 mi) between breeding grounds in and wintering areas in . Tracking studies indicate that individual birds may cover up to 10,000 kilometers one way to reach the and biomes, primarily in . These long-distance journeys highlight the species' endurance as an aerial insectivore adapted to extensive seasonal movements. Southbound migration typically begins in late to , earlier than many other North migrants, with flocks departing areas and peaking in visibility during early evening flights. Northbound return occurs from March to May, with arrivals on grounds by April in southern regions and later in northern latitudes. During , common nighthawks often fly nocturnally or at , utilizing high-altitude soaring to cover distances efficiently while foraging opportunistically on . Migration routes vary by population: eastern birds follow the Mississippi Flyway, converging in the before crossing the in a shared trans-Gulf flight, often stopping briefly in or coastal areas. Western populations tend to travel overland through and , avoiding major water crossings. Stopover sites include prairies, river valleys, marshes, and coastal dunes, where birds rest and feed in open habitats similar to their breeding preferences. Recent studies in the using GPS and geolocator tracking have revealed detailed dynamics, including non-stop flights of up to 2,500 kilometers across the and convergence of diverse breeding populations on common routes. These efforts, such as those deploying lightweight GPS-Argos tags, confirm high connectivity between breeding and wintering areas, aiding by identifying key bottlenecks.

Ecology and behavior

Activity patterns and vocalizations

The common nighthawk (Chordeiles minor) is primarily crepuscular and nocturnal, with peak activity occurring at dawn and dusk when it forages for insects in erratic, bounding flights characterized by deep wingbeats interspersed with gliding. During the day, it roosts inconspicuously on horizontal branches, flat rooftops, or the ground, often aligning its body parallel to the perch for camouflage among bark or gravel. At higher latitudes, activity may extend through much of the night due to prolonged twilight periods. The species produces a variety of vocalizations and mechanical sounds for communication. Its most distinctive call is a loud, nasal "peent" or "beernt," delivered repeatedly during flight to advertise territory and attract mates. Males uniquely generate a deep "boom" by rapidly flexing their wings during steep dives, creating a rush of air through the modified primaries; this sound serves as a territorial signal centered around nesting areas. When perched or disturbed, individuals may emit softer, throaty clucks or hisses, particularly in defensive contexts. These vocalizations and displays play key roles in territorial defense and social interaction, with the "peent" call functioning broadly for spacing individuals and signaling presence, while the male's boom delineates exclusive nesting territories. In breeding contexts, these sounds intensify to facilitate attraction and pair coordination. Adaptations for low-light activity include large eyes with a tapetum lucidum, a reflective layer that enhances vision by redirecting light onto the retina, enabling effective navigation and prey detection in dim conditions.

Foraging and diet

The common nighthawk is an aerial insectivore that forages primarily by hawking flying insects during crepuscular periods at dawn and dusk. It pursues prey in acrobatic flight, scooping insects into its wide-gaping mouth lined with stiff, bristle-like rictal bristles that function to guide and net small arthropods efficiently. This method allows the bird to capture prey mid-air without perching, often over open areas, wetlands, or near artificial lights where insects congregate. Additionally, the common nighthawk drinks on the wing by skimming low over water surfaces to sip from lakes, rivers, or ponds. The consists almost exclusively of flying , with over 50 documented, including (Coleoptera), moths and (Lepidoptera), flying and wasps (Hymenoptera), flies (Diptera), (Trichoptera), and grasshoppers. Studies indicate a selective preference for and hymenopterans relative to their availability in the environment, while dipterans are consumed less than expected. For example, flying can comprise up to 25% of the in some populations, with individual birds consuming 200–1,800 per content analysis. Overall, a single nighthawk may ingest hundreds to over 1,000 in a night, depending on prey abundance and duration. diversity varies seasonally and regionally; breeding birds in boreal habitats show reduced prey diversity compared to non-breeding periods, potentially shifting toward larger or more available taxa like wood-boring in post-fire landscapes.139[020:ADOCNCM]2.0.CO;2.full) Specialized adaptations enhance efficiency, including the enormous gape—up to one-third of the head length when open—and forward-projecting rictal bristles that act as a sensory and trapping mechanism for airborne prey. Prior to long-distance , common nighthawks increase to build reserves, often substantially elevating body mass to fuel non-stop flights southward. This hyperphagia supports their energy demands during travel, with intensifying in late summer to maximize nutritional .

Reproduction and breeding

The common nighthawk exhibits a monogamous , forming seasonal pairs that cooperate during the breeding period. rituals primarily involve males performing aerial displays, including steep dives from heights of up to 100 meters that produce a distinctive booming sound as air rushes through the modified primaries of the wings, often accompanied by repeated nasal "peent" calls to attract females and establish . Nesting occurs without any constructed nest structure, with the female selecting camouflaged sites in open, well-drained areas such as bare ground in forest clearings, gravel beaches, rocky outcrops, burned-over woodlands, or flat gravel roofs to minimize detection by predators. These sites provide nearby shade and proximity to areas, enhancing the eggs' cryptic coloration that blends with the . In northern portions of the breeding range, the season spans from late May to early August, typically producing one brood per year with a clutch of two eggs laid 1–2 days apart. requires 18–20 days, during which asynchronous hatching is uncommon due to the brief laying interval. Breeding site fidelity is generally loose, though some females return to the same or nearby locations in successive years, while males may reuse territories within approximately 1 km; overall, pairs show moderate to similar types rather than exact sites.

Parental care and development

The female common nighthawk performs the majority of duties, lasting 18-20 days, during which she leaves the eggs briefly in the evening to . The eggs, laid directly on the ground without a nest structure, feature cryptic gray and brown speckling for against predators. The female turns the eggs periodically using her bill and feet to ensure even development, shifting her position to regulate temperature by orienting away from the sun during the day. Upon hatching, the semiprecocial chicks emerge covered in sparse down—dark gray above and creamy below—with eyes partially or fully open, enabling mobility within hours of hatching in response to the female's calls. As nidicolous young, they remain dependent on the site initially but can move short distances, up to several meters per day in the first week, often guided by the female who may relocate them for better shade or protection. Both parents feed the chicks regurgitated insects, with the male taking a more active role in provisioning, especially during potential second broods. Biparental care continues through the nestling period, with the brooding the chicks during the day for the first 10-15 days while the defends the through aerial displays and chases. The young at 17-21 days, achieving initial flight capability, though they remain under parental supervision for feeding until approximately 30 days old, when they become independent. Second broods are uncommon but possible in some populations, where the cares for the first clutch during the 's re-incubation. Chick development is rapid, with pinfeathers emerging by day 7, strong hopping by day 16, and proficient flight by 25-30 days; they reach adult size and full within 45-50 days. Despite this quick growth, early mortality is high, often due to exposure from the open ground nest sites and predation, with nest survival rates through the full period estimated at around 0.50-0.79 in studied populations.

Predators and defenses

The common nighthawk faces threats from a variety of predators targeting adults, eggs, and chicks. Ground-based predators include mammals such as , raccoons, coyotes, foxes, domestic dogs, and housecats, as well as reptiles like ; corvids (e.g., and ) and also prey on eggs and young. Aerial predators consist of raptors like American kestrels and , which may target flying adults or resting birds. In specific cases, such as in habitats, corvids like Florida scrub-jays have been observed directly preying on unattended eggs by grasping and consuming them. To counter these threats, common nighthawks employ several anti-predator strategies. Their cryptic, mottled brown provides effective against ground substrates, allowing adults and chicks to blend seamlessly and remain immobile when approached. Behavioral defenses include distraction displays, where females feign injury to lure predators away from the nest, and aggressive responses by males, such as diving at intruders with booming wing sounds or hissing and beating wings. Chicks may spread their wings and hiss when threatened to appear larger or more formidable. Additionally, the species' crepuscular and nocturnal activity patterns limit encounters with diurnal predators. Ground-nesting habits heighten vulnerability, as eggs and chicks receive no structural protection and are laid directly on bare , , or sparse , exposing them to disturbance and opportunistic predators. Urban roof-nesting on flat surfaces can offer partial protection by elevating sites above ground predators, though it still leaves nests susceptible to mammalian climbers like . Predation significantly impacts , serving as the primary cause of nest failure across studied populations, with overall nest success rates varying from 43% to 93%. This results in substantial mortality, often exceeding 50% in ground-nested cohorts due to predation and related factors.

Conservation and status

The global population of the common nighthawk is estimated at 23 million mature individuals (as of ). This figure, derived from Partners in Flight assessments, reflects the breeding population primarily across , with approximately 93% occurring in the , , and . Population trends indicate a steep overall decline across , with a cumulative reduction of 58% since 1970 according to Partners in Flight data. In specifically, populations have dropped by 68% since 1970, though the rate of decline has slowed in recent decades. North American Breeding Bird Survey () analyses from 1966 to 2019 further document an average annual decline of 1.23%, resulting in a roughly 48% loss over that period. While some southern regions, such as parts of the , show relative stability or slower declines in the past decade, the species remains on a downward trajectory continent-wide. Regional variations highlight pronounced differences, with sharp declines in the and , where breeding occupancy has fallen by up to 70% in some areas, partly linked to pressures. In contrast, certain urban-adapting populations in southern latitudes exhibit slight increases or stabilization, suggesting localized resilience. Ongoing monitoring through platforms like eBird and provides insights into fluxes and abundance patterns, revealing persistent declines as of 2025 assessments. For instance, eBird trends maps from 2012 to 2022 indicate continued negative changes in relative abundance across much of the range, while data from migration watchpoints confirm reduced passage numbers in key flyways. These tools, combined with routes, underscore the ongoing population challenges without signs of reversal.

Threats and conservation measures

The common nighthawk faces several human-induced threats that contribute to its population declines. Habitat loss, primarily from and , has reduced suitable open and areas, such as grasslands and urban gravel rooftops, which the species relies on for nesting. suppression in forests further limits natural open habitats by allowing tree canopy closure, exacerbating the loss of ground-nesting sites. The widespread use of non-selective pesticides has significantly decreased aerial populations, the bird's primary food source, leading to reduced prey availability across and wintering grounds. Collisions with buildings and vehicles pose a direct mortality , particularly during when artificial lights disorient individuals at urban stopover sites, causing them to crash into structures or become exhausted. compounds these issues by altering and patterns, potentially creating mismatches in timing that affect success and food availability, as evidenced by recent studies on influences during fall migrations along key routes like . Conservation efforts for the common nighthawk focus on mitigating these threats through , enhancement, and policy measures. The is classified as Least Concern globally by the IUCN but holds Special Concern status in under COSEWIC and Species at Risk Act, and is designated a Species of Greatest Conservation Need in several U.S. states, including , reflecting regional vulnerabilities despite a stable overall population. Ongoing programs, such as Partners in Flight assessments and regional initiatives like Project Nighthawk in and Chicago's urban surveys, track population trends and nesting success, with 2025 data showing increased volunteer participation to inform . restoration efforts emphasize creating gravel nest patches on urban rooftops to replace unsuitable rubberized surfaces and restoring open grasslands through controlled burns and reduced mowing, as outlined in Canada's 2016 Recovery Strategy. Regulations on use, including restrictions on non-selective applications in agricultural areas, aim to bolster populations, while emerging 2020s initiatives target reduction at stopovers to minimize disorientation. Recent research highlights the need for further study on climate-induced mismatches, potentially leading to upgraded protections if declines accelerate.

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