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Scup

Scup (Stenotomus chrysops) is a temperate marine fish belonging to the family (porgies), characterized by its compressed body, small mouth, and silvery coloration with dark bars. Native to the western , it ranges from , , to , , though it is rare south of . This demersal species inhabits sandy, muddy bottoms, beds, and areas with rocky structures or eelgrass, typically at depths from 15 to 200 meters. Adults commonly reach lengths of 25 cm, with a maximum of 46 cm and weight up to 2.1 kg, maturing at around 15.5 cm. Scup exhibit migratory behavior, schooling in inshore waters from early to fall along the U.S. East , particularly from to , before shifting to offshore habitats in winter, south of to . Juveniles favor estuaries and bays with salinities above 15 ppt and temperatures over 45°F, settling on substrates like , , and beds for and shelter. Their diet consists primarily of benthic , including amphipods, worms, , and juvenile squid. occurs from May to August in coastal waters of southern to , where eggs and larvae develop in estuarine "mixing" and "seawater" zones at temperatures of 55–73°F. Economically significant, scup supports both commercial fisheries—harvested for fresh, smoked, or frozen markets—and recreational , with management overseen by the Mid-Atlantic Fishery . Current stock assessments indicate it is not overfished, with no occurring as of the 2025 assessment. Essential habitat protections, established under Amendment 12 in 1999, encompass key areas for all life stages to sustain populations.

Taxonomy

Scientific classification

The scup, scientifically known as Stenotomus chrysops (Linnaeus, 1766), belongs to the family , which encompasses porgies and seabreams. Its full taxonomic hierarchy is as follows:
RankTaxon
KingdomAnimalia
PhylumChordata
Class
OrderSpariformes
Family
GenusStenotomus
SpeciesS. chrysops
The family comprises approximately 166 valid species across 39 genera, primarily distributed in temperate and tropical marine waters worldwide. Recent phylogenomic studies have revised the higher classification of from the traditional order to Spariformes within the , based on molecular evidence from mitochondrial and nuclear genes that better reflects evolutionary relationships among perciform fishes. The Stenotomus is restricted to two endemic to the western North Atlantic, including S. chrysops (scup) and S. caprinus (longspine porgy), distinguishing it from the more diverse Indo-Pacific genera within , such as Pagrus and Acanthopagrus, which exhibit broader biogeographic ranges and adaptations to environments.

Etymology and common names

The scientific name of the scup is Stenotomus chrysops. The genus name Stenotomus, established by Theodore Nicholas Gill in 1865, derives from the Greek words stenos (narrow) and (cutting), referring to the species' narrow incisors. The specific epithet chrysops combines the Greek chrysos (gold) and (eye or face), alluding to the golden iridescent markings around the eyes and on the head. The primary common name "scup" originates from the Narragansett language, an Algonquian dialect spoken by indigenous peoples of the ; it is a shortened form of mishcuppaug (or variants like mishcùppaûog), which translates to "thick-scaled" or "large, closely scaled," describing the fish's robust, adherent scales. This name was documented in early colonial records, such as ' 1643 A Key into the Language of America, where he referred to the fish as "mishcup" while noting its use by . In addition to "scup," the fish is widely known as "porgy" or "northern porgy," an English adaptation of the pargo (referring to sea breams in the family ), which entered through colonial trade and fishing contexts in the mid-Atlantic and southern regions. Regional market variations include "fair maid" or "maiden" in the area, "ironsides" in some fisheries due to the fish's tough body, and "sea bream" in commercial settings to evoke sparid . These names reflect the scup's broad distribution along coast and its historical role in both subsistence and market fisheries.

Physical characteristics

External morphology

The scup (Stenotomus chrysops) possesses an ovate-elliptical body that is deep and laterally compressed, with a steep profile formed by a convex and a strong depression anterior to the origin. The body coloration is predominantly silvery, with the back and upper sides exhibiting a grayish to brownish hue often tinged with bluish or greenish ; the belly is white, and faint pale vertical bars may appear along the sides, particularly in younger individuals. The head features iridescent blue-green markings, with faint spots and a patch above and behind each eye. The head is short and blunt, bearing a small terminal mouth with a narrow gape. The jaws contain small, incisiform front teeth arranged in close-set bands for grasping prey, with the outer band teeth enlarged, compressed, and narrower at the base; laterally, there are two rows of molar-like teeth for crushing hard-shelled organisms. The is long and continuous, typically with 12 spines and 12 soft rays. The anal fin includes 3 spines and 11 to 12 soft rays, the pectoral fins are elongated and extend beyond the dorsal fin origin, and the caudal fin is forked. Adult fins are mottled with dark brown, while those of juveniles may show faint barring. The body is covered with relatively large, ctenoid scales that are thick and firmly adherent.

Size, growth, and lifespan

Scup (Stenotomus chrysops) attain a maximum total length of 46 and a maximum reported weight of 2.1 kg. Individuals commonly encountered in commercial and recreational fisheries typically measure 20–30 in total length and weigh 0.2–0.5 kg. Growth in scup follows the von Bertalanffy model, derived from length-at-age data obtained through otolith analysis. For males, the asymptotic length (L) is approximately 34.3 cm with a growth coefficient (K) of 0.27 year−1; for females, L is 37.4 cm and K is 0.23 year−1. Juvenile growth is relatively rapid, with fish reaching 10–15 cm fork length by age 1, after which the rate slows in adults as energy allocation shifts toward maintenance and reproduction. The lifespan of scup extends up to 19–20 years in the wild, as determined by counting annuli in otoliths. is typically reached at age 2, when females measure about 16.3 cm and males 15.6 cm in length; females exhibit slight by attaining marginally larger sizes at maturity.

Distribution and habitat

Geographic distribution

The scup (Stenotomus chrysops) inhabits the western , with its overall range extending from , , southward to , , though records become rare south of . Occasional vagrants have been documented as far south as , but these sightings are infrequent and do not indicate established populations. The species is absent from established populations outside this native range, including failed introduction attempts in . The core population is concentrated in the Mid-Atlantic Bight, spanning from , , to , , where the species is most abundant. Highest densities occur off the coasts of and , as evidenced by commercial and recreational catch data from these regions. Scup is assessed and managed as a single stock from to , . Scup exhibit distinct seasonal migrations driven by temperature changes. In winter, they migrate offshore to deeper waters (typically 50-150 meters) from southern to , where they overwinter in cooler, stable conditions. As water temperatures rise in , scup undertake an inshore migration northward, reaching their northern range limits by summer and residing in coastal areas through the warmer months. By fall, they begin an offshore exodus southward, returning to wintering grounds as temperatures decline. These movements align with the species' preference for waters above 7°C, influencing their annual distribution patterns.

Habitat requirements

Scup primarily occupy demersal habitats on the continental shelf in temperate western North Atlantic waters, with environmental preferences centered on moderate depths, soft substrates, and stable parameters. They thrive in areas with bottom temperatures ranging from 5 to 25°C (41 to 77°F), salinities of 25 to 35 , and dissolved oxygen concentrations exceeding 4 /L, conditions that support their physiological needs across life stages. Habitat utilization differs markedly by developmental phase, reflecting adaptations to , , and . Eggs and larvae are pelagic, drifting in the upper over the inner in coastal waters typically shallower than 50 m, where temperatures span 13 to 23°C and salinities exceed 15 ppt for optimal development. Juveniles transition to shallow, nearshore environments in estuarine and inshore bays, such as , at depths under 20 m on sandy or muddy bottoms often associated with eelgrass beds or other vegetation for shelter. These young prefer high-salinity waters (30 to 35 ppt) and temperatures ranging from 5 to 27°C, with summer peaks at 16-22°C, to support growth and avoid predation. Adults are demersal, inhabiting the continental shelf at depths of 10 to 150 m (33 to 492 ft), where they associate with sandy or muddy substrates near structural features like shipwrecks, natural reefs, or beds that provide cover and prey access. Optimal conditions mirror those of juveniles but extend to cooler shelf breaks, with peak abundance in waters of 13 to 16°C.

Biology and behavior

Reproduction and life cycle

Scup engage in broadcast spawning, with occurring when females release eggs and males simultaneously release in coastal waters over sandy or bottoms. The spawning season in the Mid-Atlantic region spans May to August, peaking in June when water temperatures exceed 10°C. This activity takes place in nearshore areas, including large bays and sounds from to . Females are batch spawners, releasing eggs in multiple batches over the season, with an average annual of approximately 7,000 pelagic eggs per female. The eggs are small (0.85–1.15 diameter), transparent, spherical, and buoyant, hatching in 2–4 days. Upon hatching, scup larvae measure approximately 2 and remain pelagic in coastal waters, feeding on small within 2–3 days post-hatch and drifting toward estuarine areas for 2–4 weeks. to the juvenile stage occurs at 10–15 total length, with to inshore bottom habitats by early July at sizes of 15–30 . Juveniles recruit to adult habitats around age 1 year. is attained by about 50% of females at age 2 (approximately 20 cm total length) and by males at age 3. The overall is nearly 1:1, though females tend to dominate in older age classes due to differential mortality patterns.

Diet and feeding habits

Scup primarily consume benthic , with crustaceans such as amphipods, , and forming the largest portion of their at approximately 40-55% by weight, depending on region and size class. Mollusks, including clams and mussels, contribute around 13-30% of the , while polychaete worms account for 17-47%, alongside smaller amounts of small , , and occasional fish larvae. These proportions reflect opportunistic bottom feeding, where prey availability in sandy or muddy substrates influences composition. Feeding is predominantly bottom-oriented and occurs during daytime, relying on visual foraging to locate prey on or near the seabed. Scup utilize specialized pharyngeal teeth to crush hard-shelled items like mollusks and crustaceans, enabling efficient processing of benthic resources. Juveniles, typically under 10 cm, shift toward more planktonic prey, consuming higher proportions of mysids, copepods, and insect larvae (up to 40-55% arthropods overall), which supports their early growth in estuarine and nearshore environments. Seasonal variations occur, with increased intake (e.g., bivalves) in spring and summer, alongside sustained and consumption, aligning with higher prey densities and metabolic demands. As mesopredators, scup occupy a of approximately 3.5-3.8, determined from diet composition analyses across northwest Atlantic studies. This position underscores their role in linking primary consumers to higher trophic levels through selective predation on infaunal and epibenthic communities.

Predators and ecological interactions

Scup (Stenotomus chrysops) face predation pressure across life stages from a diverse array of marine predators, primarily elasmobranchs and bony fishes. Adults are consumed by at least 19 species, with the most frequent including smooth dogfish (Mustelus canis), spiny dogfish (Squalus acanthias), bluefish (Pomatomus saltatrix), striped bass (Morone saxatilis), black sea bass (Centropristis striata), and summer flounder (Paralichthys dentatus), as well as sharks like sandbar shark (Carcharhinus plumbeus) and dusky shark (Carcharhinus obscurus), silver hake (Merluccius bilinearis), weakfish (Cynoscion regalis), and goosefish (Lophius americanus). Juveniles experience heightened vulnerability to these piscivores, in addition to invertebrate predators such as crabs and avian predators including shorebirds. Larvae suffer intense predation from planktonic organisms like medusae, ctenophores, chaetognaths, crustaceans, and small fishes, contributing to high mortality rates during the early pelagic phase. Predation rates on scup vary by size class and predator, with larvae facing near-total attrition from diffuse planktivory, while juveniles and adults form a notable component of diets in coastal and shelf ecosystems. For instance, scup constitute 4.7% by weight in the diet of , and broader analyses indicate adults can comprise 10-20% of the diet for select predators like and weakfish in regions of overlap. This top-down pressure shapes scup , with juveniles particularly susceptible during inshore , where predation can exceed 80% annually in some cohorts due to larger fishes and birds. As a key species, scup play a vital ecological role in linking benthic —through their consumption of and small fishes—to higher trophic levels, supporting piscivores and maintaining energy flow in mid-Atlantic shelf ecosystems. Their abundance and distribution serve as indicators of continental shelf health, reflecting responses to environmental changes like shifts and prey availability. Scup occasionally exhibit cannibalistic behavior, particularly larger individuals preying on juveniles, though this is infrequent. They also compete with other sparids, such as (Tautoga onitis) and sheepshead (), for shared benthic resources like crustaceans and mollusks in estuarine and nearshore habitats. Additionally, scup host various parasites, including trematodes like those in the family Lepocreadiidae and branchiurans such as Argulus inexpectatus, but these generally exert low population-level impacts due to moderate prevalence.

Fisheries

Commercial fishing

The commercial fishery for scup (Stenotomus chrysops) is one of the oldest , with documented harvests dating back to colonial times. Landings peaked in 1960 at approximately 48.9 million pounds (22,200 metric tons), driven by high demand and unrestricted effort along coast. By the late , annual commercial landings had stabilized between 8.8 and 21.7 million pounds, but intense pressure in the led to a sharp decline, with catches dropping to a low of 2.7 million pounds by 2000 due to and reduced stock biomass. Commercial harvesting primarily employs bottom otter trawls, which account for about 84% of landings, targeting scup on bottoms at depths of 50-150 feet. Other methods include fish pots and traps (7% of catch), gillnets, and handlines, with seasonal variations favoring traps in inshore areas and trawls offshore. in these operations commonly includes , little skate, , and northern sea robin, though observer data from 2019-2023 indicate these species comprise less than 10% of total catch, mitigated by gear restrictions in sensitive areas. In 2023, U.S. commercial landings totaled 13.01 million pounds (approximately 5,900 metric tons), representing 93% of the annual quota and valued at $8.18 million ex-vessel. In 2024, landings increased to 14.50 million pounds (approximately 6,600 metric tons), representing 69% of the quota. Major landing ports include (29% of total), (19%), and (13%), reflecting the fishery's concentration in the Mid-Atlantic region. The market is predominantly domestic, with scup sold fresh or frozen as fillets or whole fish under names like porgy, though limited exports occur to where it is marketed as sea bream.

Recreational fishing

Scup represent a significant component of the recreational fishery in the Northeast , particularly valued as a gamefish by surf casters and boat anglers in southern and the Mid-Atlantic region. They are highly sought after for their abundance, schooling behavior, and fighting spirit on light tackle, contributing to their popularity among hobbyist anglers. Annual recreational harvest of scup typically ranges from 10 to 20 million , with 12.85 million landed in 2023 alone according to Marine Recreational Information Program (MRIP) estimates from to . In 2024, recreational harvest was approximately 11 million pounds, equivalent to about 11-15 million . This harvest underscores the species' importance to the sport , where party boats provide accessible opportunities for anglers to target schools over wrecks and structures. Common techniques for recreational scup involve bottom rigs with baited hooks, using natural baits such as strips, clams, or sandworms to entice bites from the schooling . Light tackle with 10- to 20-pound test line is standard, allowing for sensitive presentations, while with crushed clams or mussels helps concentrate schools around the boat or structure. Party boats are particularly popular for this , enabling groups to wrecks and reefs efficiently during half- or full-day trips. The peak inshore season for scup runs from June to August, when migratory schools move close to shore in warm waters, though they can be caught from May through October in northern areas. Prime locations include the waters around in , the coastline, and in the southern portion of their range, where structures like jetties, piers, and artificial reefs attract concentrations of fish. The International Game Fish Association (IGFA) all-tackle world record for scup stands at 2.06 kg (4 lb 8 oz), caught in , , in June 1992.

Management and conservation

Regulatory measures

The scup is jointly managed by NOAA Fisheries and the Mid-Atlantic in federal waters of the , while the Atlantic States Marine Fisheries Commission coordinates interstate in state waters, with implementation by state marine fisheries commissions in , , , , , , , , and . The , Scup, and Plan, adopted through Amendment 8 in 1996, provides the framework for these efforts, establishing catch limits, size restrictions, and gear requirements to prevent . Key regulations include a minimum size limit of 9 inches (23 cm) total length for commercial harvest and 10 inches for in federal waters, with states often aligning or adjusting slightly for consistency. Seasonal closures, such as November through April in some state waters (e.g., January–April recreational in ), help manage quotas and protect overwintering aggregations, while recreational bag limits vary by state from 30 to 50 fish per person per day, with a federal possession limit of 40 fish. For 2025, the annual catch target () is 65% of the acceptable biological catch (), with the commercial quota set at 19.54 million pounds (8,865 metric tons) after subtracting projected discards, and 35% allocated to the recreational ACT. The commercial quota is divided into three seasons: Winter I (45.11%, January-April), Summer (38.95%, May-September), and Winter II (15.94%, October-December), with state-specific shares applying only to the Summer quota—for example, receives 56.2% and 21.6%. Trip limits for federally permitted vessels are 50,000 pounds per trip in Winter I and 15,000 pounds in Winter II, while Summer harvests are capped by quotas without federal trip limits. In September 2025, the Winter II quota was increased by 1,047,723 lb (475,260 kg) to 4,162,331 lb (1,888,020 kg) due to rollover of unused Winter I quota, with the possession limit raised to 15,000 lb (6,804 kg). Enforcement is facilitated through vessel monitoring systems required on federally permitted vessels longer than 50 feet, mandatory electronic logbooks for all commercial trips to track landings against quotas, and at-sea observer programs to assess bycatch rates. NOAA Fisheries provides weekly quota updates, and states submit annual compliance reports to ASMFC by 1. Framework adjustments and addenda to the FMP, such as minimum mesh sizes of 5.5 inches for trawls, have been implemented to reduce of juvenile scup and non-target species.

Population status and threats

The scup (Stenotomus chrysops) population is classified as Near Threatened by the , with the conducted in 2011 based on observed declines due to historical and potential ongoing pressures. In the United States, the (NOAA) determined the to be overfished in 1996 following severe reductions in spawning during the late and early , when commercial landings peaked at over 15 million pounds in 1991 before plummeting to historic lows by the mid-1990s. in the late and early reduced spawning by approximately 97%, reaching historic lows in the late ; subsequent led to a roughly 30-fold increase in through the under reduced pressure. The was successfully rebuilt ahead of schedule in , and the 2023 track confirmed it remains not overfished, with spawning estimated at 193,087 metric tons in 2022—246% of the (MSY proxy of 78,593 metric tons)—while mortality remains below sustainable levels (F < FMSY). Beyond historical , current threats to scup populations include degradation from , which disturbs benthic communities in the species' preferred and substrates along the Mid-Atlantic shelf, though these habitats show relative compared to more structured environments. exacerbates vulnerability through warming waters that have driven northward shifts in scup , particularly during spring migrations on the Northeast U.S. shelf, potentially altering seasonal availability and spawning success. in non-selective trawl gears, such as those targeting squid, continues to impact juvenile and sub-legal scup, with discard rates historically contributing significantly to total removals before regulatory modifications like excluder devices reduced incidental capture. Additionally, pollution in coastal estuaries—where scup larvae settle and rear—poses risks through contaminants like and organic pollutants, which have been linked to and other sublethal effects in juvenile stages exposed via contaminated prey or direct contact. Population monitoring relies on annual stock assessments coordinated by NOAA and the Mid-Atlantic Fishery Management Council, incorporating commercial and recreational catch data, fishery-independent surveys, and age-structured models such as virtual population analysis (VPA) to estimate trends and reference points. variability, a key driver of stock fluctuations, is strongly tied to environmental factors including water temperature and prey , which influence larval survival and year-class strength, as observed in interannual shifts in catch rates and . These assessments highlight the stock's but underscore the need for ongoing vigilance against non-fishing stressors to maintain .

Culinary aspects

Preparation and cooking

Scup possesses firm, flaky white flesh with a mild, sweet flavor reminiscent of , making it versatile for various culinary applications. Its lean texture and relatively low bone content allow for easy preparation when cooked whole, though the small bones can make filleting challenging. To preserve quality, scup should be bled immediately after capture by cutting the gills or throat while the heart is still pumping, then placed on to cool rapidly and maintain freshness. Before cooking, and gut the fish thoroughly, as this removes impurities and enhances the sweet . Common preparation methods emphasize simple techniques that highlight the 's delicate . Pan-frying or deep-frying yields a crispy exterior while keeping the interior moist, often seasoned with salt and pepper. Broiling or whole scup with herbs, , or adds a charred smokiness, and baking with and creates a tender, aromatic dish. For lighter options, scup can be served raw as or in , known as "orata Americana," where thin slices showcase its subtle sweetness. In regional cuisine, scup features prominently in clam bakes, where whole fish are steamed with s, corn, and potatoes over hot rocks for a communal seaside meal. Along the coast, it is prepared Mediterranean-style as "Montauk sea bream," roasted whole with , , and to emphasize its flaky texture and shrimplike notes.

Nutritional profile

Scup (Stenotomus chrysops) is a nutrient-dense option, providing a low-calorie source of high-quality protein when cooked. Per 100 grams of scup cooked by dry heat, it contains 123 calories, 23.7 grams of protein, 2.47 grams of total fat (including 0.57 grams of ), and 0 grams of carbohydrates. This macronutrient profile makes it a protein that supports muscle maintenance and without contributing significant dietary carbohydrates or excess calories. In terms of micronutrients, scup is particularly rich in essential vitamins and minerals vital for metabolic and immune function. It offers 47.7 micrograms of (87% of the daily value), 2.07 micrograms of (86% of the daily value), and 298 milligrams of (24% of the daily value) per 100 grams cooked. Additionally, it provides 0.614 grams of omega-3 fatty acids per 100 grams, including (EPA) at 174 milligrams and (DHA) at 349 milligrams, which are long-chain polyunsaturated fats concentrated during cooking. The omega-3 fatty acids in scup contribute to cardiovascular health by reducing and supporting healthy function, as established in dietary guidelines from health authorities. With low mercury levels below 0.1 parts per million—placing it among the safest choices for regular consumption—scup serves as a sustainable, low-fat alternative to , delivering comparable protein with fewer saturated fats and added marine-derived nutrients. Nutritionally, scup aligns closely with other lean white fish like , which provides about 105 calories, 23 grams of protein, and 0.9 grams of fat per 100 grams cooked, but scup stands out with higher content (47.7 micrograms versus 38 micrograms in ). Compared to , which offers around 86 calories and 15 grams of protein per 100 grams cooked with only 33 micrograms of , scup provides a more concentrated source of this for function and protection against .
Nutrient (per 100g cooked, dry heat)Amount% Daily Value*
Calories123 kcal-
Protein23.7 g47%
Total Fat2.47 g3%
0.57 g3%
Carbohydrates0 g0%
Omega-3 Fatty Acids (total)0.614 g-
47.7 µg87%
2.07 µg86%
298 mg24%
*Based on a 2,000-calorie ; values from USDA data.

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