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Lentil

The lentil (Lens culinaris) is a bushy annual in the family , cultivated globally for its edible, lens-shaped seeds, which serve as a nutrient-dense . Growing to 12–30 inches (30–76 cm) tall with semi-erect stems, the plant features pinnate leaves with 4–7 pairs of ovate to lanceolate leaflets terminating in tendrils, small self-pollinating flowers ranging from white to dark purple, and flat pods (0.5–0.75 inches long) that each contain 1–2 seeds (0.25–0.5 inches in diameter). The seeds vary by type, with macrosperma varieties having larger seeds and yellow cotyledons, and microsperma varieties featuring smaller seeds with red, orange, or yellow cotyledons. Lentils originated in the Near East, where wild progenitors (Lens orientalis) were gathered as early as 23,000 years before present at sites like Ohalo II in , and domestication occurred around 10,000–9,500 BCE in the (southern to ) during the period, evidenced by carbonized remains at sites such as and Yiftah'el showing non-shattering pods and larger seeds. By the 7th millennium BCE, cultivation spread to , , and the , with lentils becoming a foundational crop alongside and in early . Today, lentils are grown in over 50 countries across cool, semi-arid regions requiring at least 10 inches (25 cm) of annual rainfall, thriving in well-drained sandy soils with pH 5.5–7 and maturing in 80–110 days; major producers include , , , , and the (notably and ), with global production reaching approximately 7 million metric tons as of 2023. As a rotational crop, lentils fix 30–100 pounds of per in the , enhancing and reducing the need for synthetic fertilizers in cereal rotations. Lentils are prized for their versatility in cuisines worldwide, appearing in soups, stews, salads, , and as or alternatives, with whole seeds (e.g., green or brown varieties) retaining shape during cooking (20–30 minutes) and split seeds (e.g., red or yellow) breaking down quickly (10–15 minutes). Nutritionally, one cup of cooked lentils (198 g) delivers 18 g of protein, 16 g of fiber, and significant (358 mcg, 90% ), iron (6.6 mg, 37% ), (731 mg, 16% ), and polyphenols with properties, while being low in sodium (<5 mg) and (0.1 g). These attributes support health benefits including lowered and , improved glycemic control in diabetics, and gut health via prebiotic fibers and , as shown in human and animal studies. Varieties like French Puy (green, earthy flavor) and Beluga (black, caviar-like) add diversity, with dried lentils storable for up to a year and cooked portions lasting 3–5 days refrigerated.

Taxonomy and Description

Etymology

The English word "lentil" derives from the lentille, which in turn comes from the Latin lenticula, a form of lēns meaning "lentil," reflecting the seed's characteristic biconvex, -like shape. This Latin root traces back to a Proto-Indo-European word lent-, shared with terms in other ancient languages for similar objects. The optical term "" later borrowed from this same Latin source due to the resemblance between the seed and the curved glass used in . In , the lentil was known as phakós (φακός), a term used by classical authors like and to denote the plant and its seeds, often in medicinal or dietary contexts. Similarly, in , it is called masūra (मसूर), a name appearing in ancient texts such as the for its role in and cuisine. These historical terms highlight the lentil's early cultural significance across , where it was valued as a staple . Common names vary by region and often reflect seed color or type; for instance, in , the red variety is termed "masoor" in and related languages, derived directly from the Sanskrit masūra. In , it is "lenteja," echoing the Latin lenticula and emphasizing the shape. Such variations underscore linguistic adaptations influenced by trade and . The scientific evolved with Carl Linnaeus's system in the ; he classified the lentil as Ervum lens in (1753), drawing on the Latin lēns for the species epithet. Later, in 1787, Friedrich Kasimir Medikus established the modern name Lens culinaris, placing it in the genus to better reflect its distinct morphology and culinary use, a designation retained in contemporary .

Botanical Features

The lentil plant, Lens culinaris, is an annual herbaceous legume in the family, characterized by its slender, semi-erect stems that typically reach heights of 15 to 50 cm, though they can extend up to 75 cm under favorable conditions. The stems are often branched in a bushy form, with many ascending branches, and exhibit pubescence, featuring fine hairs that cover the surface, aiding in adaptation to various environmental stresses. The leaves are and pinnate, arranged alternately along the , with 4 to 7 pairs of leaflets (totaling 8 to 14), each leaflet ovate to lanceolate and measuring 6 to 20 mm in length. These leaves also bear stipules at the base and terminate in a , which facilitates climbing or support on nearby structures, contributing to the plant's vining growth habit. The leaflets are typically pubescent on both surfaces, enhancing the plant's resilience in dry or windy conditions. Flowers emerge in axillary clusters on short racemes, measuring 2 to 5 cm long, and are small, typically 4 to 8 mm across, with colors ranging from to or purple. These papilionaceous blooms are self-pollinating, featuring five petals in a standard butterfly-like arrangement, and they open sequentially from lower branches upward, with individual flowers lasting only 2 to 3 days before fading. The is a flat, smooth, oblong , approximately 10 to 19 mm long and 5 to 9 mm wide, containing 1 to 2 -shaped per . The vary in color, including green, brown, red, and yellow varieties, and are or flattened, with sizes ranging from 2 to 9 mm in diameter depending on the type (macrosperma or microsperma). The life cycle of L. culinaris spans 80 to 150 days from to maturity, varying by , sowing date, and environmental conditions, progressing through distinct stages beginning with , which occurs 5 to 10 days after sowing under optimal temperatures above 5°C. Vegetative growth follows emergence, involving elongation, branching, and over the next 20 to 40 days, during which the plant establishes its bushy form. Flowering initiates 40 to 70 days after sowing, starting at the lower nodes and continuing indeterminately upward if conditions permit, with pod formation beginning 3 to 4 days after petal drop. Maturity is reached when pods yellow and dry, typically 30 to 80 days post-flowering, with lower pods ripening first to allow staged harvest. This species is closely related to its wild progenitor, Lens orientalis.

Classification

The lentil (Lens culinaris) is classified within the kingdom Plantae, phylum Tracheophyta, class Magnoliopsida, order Fabales, family Fabaceae, genus Lens, and species L. culinaris. This positioning places it among the legumes, a diverse family known for nitrogen-fixing capabilities through symbiotic relationships with rhizobia bacteria. The cultivated lentil is L. culinaris, distinguished from its wild progenitor L. orientalis, which exhibits greater genetic diversity and adaptation to natural habitats in the Near East. L. orientalis is considered the direct ancestor, with archaeological and genetic evidence supporting its role in the domestication process around 10,000 years ago. The genus Lens encompasses the cultivated L. culinaris alongside six wild species, including L. orientalis, L. ervoides, L. nigricans, L. tomentosus, L. lamottei, and L. odemensis, all sharing close genetic proximity that facilitates interspecific hybridization for breeding purposes. These wild relatives form a primary with L. culinaris, enabling , while secondary and tertiary pools involve more distant species with partial cross-compatibility. Evolutionarily, the lentil belongs to the cool-season food group within , characterized by a diploid number of 2n=14, which supports its adaptation to temperate climates and short-day photoperiods. This is conserved across the genus, underscoring the phylogenetic closeness among species.

Varieties and Types

Major Cultivars

Lentils (Lens culinaris) are primarily categorized into two major groups based on seed size: macrosperma (large-seeded types, also known as Chilean lentils) with seeds exceeding 6 mm in diameter or over 50 g per 1,000 seeds, and microsperma (small-seeded types, or lentils) with seeds under 6 mm or less than 40 g per 1,000 seeds. This classification influences agronomic traits such as yield potential, which generally ranges from 1 to 3 tons per across cultivars, with large-seeded types often achieving higher yields in favorable conditions due to fewer seeds per pod but larger individual size. Macro lentils, characterized by their larger seeds, include and varieties that are typically cooked whole due to their firmer and longer cooking time of 20 to 45 minutes. These s feature yellow cotyledons and minimal pigmentation in vegetative parts, making them suitable for dehulling with moderate ease, though their thicker seed coats can require mechanical processing. Protein content in macro types varies from 20% to 30% on a dry weight basis, contributing to their nutritional value. Prominent examples include the , a large-seeded lentil originating from with seeds around 7 mm, valued for its high yield and adaptability in North American production systems. Micro lentils encompass small-seeded , , , and types, which are frequently or dehulled for rapid cooking times of 15 to 20 minutes, owing to their thinner seed coats that facilitate easy dehulling and quicker breakdown during preparation. These varieties exhibit protein levels in the 20% to 30% dry weight range, similar to macro types but with potentially higher digestibility post-processing. Yield potentials align with the broader 1 to 3 tons per , though small-seeded micros may produce more uniformly in diverse environments. Key examples include Masoor, a lentil prevalent in with seeds measuring 3 to 5 mm, noted for its bold flavor and resistance to fungal diseases in South Asian trials; the Eston , a small-seeded lentil with seeds around 4-5 mm or 34-45 g per 1,000 seeds, valued for its early maturity and suitability for whole-seed cooking; and Petite Crimson, a compact variety with seeds under 4 mm, prized for its sweet profile and quick-cooking properties in global markets.

Regional Adaptations

In , particularly in subtropical regions of , lentil breeding has emphasized heat-tolerant red lentil varieties to withstand high temperatures during reproductive stages, which can otherwise lead to flower and pod abortion. The IPL 406, released in 2007 by the Indian Institute of Pulses Research, exemplifies this with its bold-seeded, red-coated profile and demonstrated tolerance to elevated temperatures up to 40°C, where it maintains higher activity and membrane stability compared to susceptible genotypes. This variety performs well under timely sown, irrigated conditions in the North Western Plains Zone, yielding 12-14 quintals per hectare over 120-155 days, making it suitable for the region's variable subtropical climates with summer heat stresses exceeding 35°C. Mediterranean lentil has focused on drought-resistant brown lentil types to cope with semi-arid conditions and irregular rainfall patterns prevalent in and . In , the Pardina (Spanish Brown) lentil, originating from the region, is noted for its drought hardiness and adaptability to a broad range of soils and elevations, featuring small, round seeds with brown seedcoats that hold shape during cooking and thrive in low-water environments without . Turkish landraces, predominantly brown-seeded, have been characterized for agro-morphological traits suited to drought-prone western Asian areas, including deeper systems and reduced canopy size to minimize loss, supporting in rainfed systems across the country's southeastern semi-arid zones. These adaptations enable consistent yields in regions with annual rainfall as low as 250-400 mm. North American lentil production, centered in the cooler prairies of and the northern U.S., has led to the development of winter-hardy lentil cultivars optimized for extended s and frost-prone conditions. Varieties like CDC LeMay, a small-seeded type released by the Crop Development Centre, exhibit resilience to cold temperatures down to -5°C during early growth, allowing fall or early spring planting in regions with 100-150 frost-free days, and deliver high yields relative to standards like CDC Milestone at 101% under cooler, longer-season environments. These cultivars benefit from the general cool-season nature of lentils, which tolerate extreme cold better than many when sown appropriately, supporting rotations in the Canadian prairies where average temperatures range from 15-20°C. In and the , small-seeded lentil variants have been selected for arid, low-input farming systems, where minimal and use predominate in marginal lands. Ethiopian landraces and improved varieties, such as Alemaya and Teshale, feature small seeds (typically 20-30 g/1000 seeds) with enhanced escape mechanisms like early maturity (90-110 days) and compact growth, enabling cultivation on vertisols and in highlands with erratic rainfall below 600 mm annually, while contributing to in resource-poor smallholder systems. Similarly, ern types from and emphasize small-seeded, prostrate habits for in arid zones, with wild relatives providing genetic resources for to low and prolonged dry spells, sustaining yields in rainfed areas with inputs limited to and basic .

History and Domestication

Origins in the Near East

The earliest archaeological evidence of wild lentil gathering dates back to approximately 23,000 years ago at Ohalo II in , with remains of the progenitor Lens orientalis found among exploited plants. Later, evidence for lentil exploitation appears at Franchthi Cave in around 11,000 BCE, where wild lentils were gathered in a context. Domestication, however, occurred during the period (approximately 9,000–8,000 BCE) in the , with charred seeds recovered from sites such as in the and Çayönü in southeastern , indicating intentional management alongside other . These findings mark lentils' integration into sedentary communities in the and northern . Key domestication traits distinguishing cultivated lentils from their wild progenitor, Lens orientalis, include non-shattering pods that retain seeds for easier harvesting, larger seed size to enhance yield and , and reduced to facilitate uniform in agricultural settings. These adaptations emerged gradually during the transition, with archaeological records showing a slow increase in seed size from wild forms (typically under 4 mm) to domesticated varieties exceeding 4 mm by the mid-Neolithic. Genetic analyses confirm that these traits were selected through human intervention, transforming wild lentils from scattered foragers' resources into reliable crops. DNA studies using genotyping-by-sequencing on wild and domesticated accessions reveal multiple domestication events centered in the , particularly in southwest Asia from southern to , around 10,000–8,000 years ago. These analyses identify two primary gene pools in cultivated lentils, indicating a protracted process rather than a single origin, with low from wild L. orientalis populations supporting localized selection. Phylogenetic and population structure data correlate these genetic patterns with archaeological evidence from sites like Yiftah'el in the (dated 10,100–9,700 cal ), underscoring the region's role as a domestication hotspot. Lentils played a foundational role in early agriculture as one of the "founder crops" of the , complementing cereals like , wheat, and in systems that supported permanent settlements. Their high protein content and ability to fix enriched soils, making them essential for sustainable food production in the resource-scarce environment. This integration marked a shift from economies to , with lentils providing dietary diversity and storage stability alongside staple grains.

Global Spread

Following its in the during the period, lentil cultivation began to disseminate through trade and agricultural expansion in the . By around 2000 BCE, lentils had reached , where archaeological evidence from tombs confirms their use as a staple food alongside other . In the , lentils appeared during the early Harappan period circa 2800 BCE, likely introduced via overland trade routes that foreshadowed later networks like the , integrating them into early agrarian societies. During the Classical era, lentils were well-established in the Mediterranean, with cultivation documented in from the onward and referenced in texts by , who described their growth habits and culinary preparation in the 4th century BCE. The further promoted lentil farming across its territories, incorporating them into diverse agricultural systems from to the provinces, where they served as a reliable protein source for soldiers and civilians alike. Medieval expansion saw lentils solidify their role in European diets, with cultivation intensifying in regions like from Roman foundations but gaining prominence as a Lenten staple and food amid feudal . Colonial exchanges in the introduced lentils to the through and Portuguese explorers, who brought seeds to the for cultivation in colonies like and . In , while present in since , lentils spread southward through Arab trade networks during the medieval period, adapting to varied climates in East and . In the , lentils gained traction in new hemispheres as protein alternatives amid post-World War II agricultural diversification. Cultivation began in in the late 1960s, starting with small-scale trials in that evolved into major production by the 1970s, driven by demand for pulse crops. Similarly, adopted lentils in the late 1980s, initially as but quickly shifting to production in response to export opportunities and benefits in .

Cultivation Practices

Environmental Requirements

Lentils (Lens culinaris) are a cool-season well-adapted to semi-arid and Mediterranean climates, where they thrive under moderate temperatures and relatively low moisture conditions. Optimal daytime temperatures range from 15°C to 30°C, with the crop exhibiting good tolerance to cooler nights and brief periods of heat, though prolonged exposure above 27°C can induce heat stress and reduce growth rates. Lentils demonstrate notable tolerance, with seedlings able to withstand temperatures as low as -5°C (21°F), allowing for early spring planting in temperate regions. Annual rainfall requirements typically fall between 300 mm and 800 mm, with a minimum of about 250 mm sufficient in drought-tolerant varieties; the crop is moderately drought-resistant once established but benefits from even distribution of precipitation to avoid water stress during flowering and pod development. Soil conditions play a critical role in lentil productivity, favoring well-drained, deep y or sandy loam textures that prevent waterlogging and support development. The performs best in s with a to slightly alkaline pH of 6.0 to 7.5, though it can tolerate a broader range from 5.5 to 8.0; acidic soils below 5.5 often lead to nutrient deficiencies and reduced yields, while highly alkaline conditions may limit iron availability. Lentils are grown successfully at elevations from (0 m) to 3,000 m above , with some varieties extending to 3,800 m in tropical highlands, where cooler temperatures at higher altitudes align with the crop's preferences. Certain cultivars exhibit sensitivity to photoperiod, with longer day lengths (over 12-14 hours) accelerating flowering in photoperiod-sensitive types, influencing to specific latitudes. As a nitrogen-fixing , lentils enhance through symbiotic relationships with bacteria, contributing 30 to 100 kg of per to subsequent crops, which supports systems with cereals by improving overall soil cycling and reducing reliance on synthetic fertilizers. This benefit also helps suppress grass weeds and mitigate disease carryover in cereal-dominated sequences, promoting sustainable agronomic practices. Varietal adaptations can further tailor lentils to marginal environments within these parameters, such as drought-prone or higher-altitude sites.

Planting and Crop Management

Lentil crops require a well-prepared to ensure optimal and . The should be cultivated to a fine, firm , free of large clods and debris, to promote even emergence and development. Fertilizers such as and can be incorporated during seedbed preparation if soil tests indicate deficiencies, but applications should be minimized or avoided to prevent interference with nodulation. Sowing lentils involves precise timing and techniques tailored to regional climates. In temperate regions, spring sowing occurs when soil temperatures reach about 4–5°C, typically in late to early May, while in Mediterranean climates, autumn sowing in allows for winter growth. Seeds are planted at a depth of 2–5 cm to access moisture without risking poor emergence, with rates of 80–150 kg/ha depending on seed and to achieve 800,000–1,300,000 per . Row spacings of 15–30 cm facilitate mechanical operations and . Prior to sowing, seeds must be inoculated with Rhizobium leguminosarum biovar viciae to enable , particularly in fields where have not been grown recently or is below 6.0. Fertilization for lentils emphasizes soil testing to match nutrient needs with availability, given the crop's low requirements due to symbiotic . If is ineffective or residual is low, apply 20–40 kg N/ha, but avoid higher rates to prevent reduced nodulation. applications of 40–60 kg P₂O₅/ha are recommended for deficient soils (below 15 Olsen P), while rates of 40–50 kg K₂O/ha suffice for levels under 100 . These inputs should be banded or broadcast pre-planting based on test results to support early growth without excess. During the growing season, lentils demand minimal , relying on 400–500 mm of total seasonal water, primarily from rainfall in semi-arid production areas. Supplemental may be applied during flowering and pod fill if rainfall is insufficient, but overwatering risks . Weeding is essential in the early stages, as lentils are poor competitors; mechanical cultivation between rows or pre-emergent herbicides like target broadleaf weeds effectively, while post-emergence options are limited to avoid injury.

Pests and Diseases

Lentil crops are susceptible to several fungal diseases that can significantly impact yield and plant health. Ascochyta , caused by the fungus Ascochyta lentis, manifests as tan to dark brown lesions with concentric rings on leaves, stems, and pods, often leading to premature defoliation and reduced seed production during wet conditions. Root rots, primarily induced by Fusarium species such as and , result in reddish-brown to black discoloration and lesions on roots and hypocotyls, causing , yellowing of foliage, and plant stunting, particularly in poorly drained soils. General control strategies for these fungal pathogens include using certified disease-free seeds, implementing crop rotations with non-host plants like cereals, and applying foliar fungicides preventatively during prolonged wet periods. Viral infections pose another major threat to lentils, with Pea seed-borne mosaic virus (PSbMV) being particularly destructive. Transmitted through infected seeds and vectors, PSbMV causes mosaic patterns on leaves, clearing, stunting, and , leading to yield losses of up to 61% in severe cases, especially with early-season infections. Management focuses on planting virus-free certified seeds, rogueing infected plants, and controlling populations to limit spread, as no curative treatments exist for diseases. Insect pests inflict both direct feeding damage and indirect harm through disease transmission in lentil fields. Aphids, particularly the pea aphid (Acyrthosiphon pisum), feed on sap from leaves and stems, causing curling, yellowing, and stunted growth while vectoring viruses like PSbMV, resulting in substantial yield reductions. Wireworms, the larvae of click beetles (Agriotes spp.), bore into seeds and roots, leading to poor seedling emergence, stand thinning, and root shredding that exacerbates drought stress. Bruchid beetles (Bruchus spp.), such as the lentil seed weevil, lay eggs on developing pods, with larvae consuming seed contents during storage, causing weight loss, reduced germination, and quality degradation. Integrated control measures encompass seed treatments with insecticides, timely scouting for thresholds (e.g., 30-40 aphids per plant), cultural practices like tillage to disrupt wireworm habitats, and hermetic storage to suppress bruchids. Nematodes represent persistent soil-borne challenges for lentil cultivation, with root-knot nematodes (Meloidogyne spp., such as M. incognita) forming on that impair nutrient and water uptake, leading to aboveground symptoms of , , and yield declines of 20-50%. Cyst nematodes (Heterodera spp., including H. ciceri) similarly encyst on , causing stunting and patchy growth patterns without prominent . Effective relies on planting resistant lentil varieties, rotating with non-host crops like small grains for 2-3 years to reduce nematode populations, and avoiding susceptible hosts in infested fields.

Production and Economics

Global lentil production has grown steadily in recent years, reaching 6.65 million metric tons in 2022, according to (FAO) data. Estimates for the 2024/2025 marketing year project output at approximately 6.6 million tonnes, marking a 15% increase from the 5.7 million tonnes of the prior season, driven by expanded acreage in key producing regions. As of late 2025, projections for the 2025-26 season indicate further growth to 7.6 million tonnes. Over the longer term from 2010 to 2024, annual production growth has averaged 2-3%, reflecting improvements in techniques and rising global . Average yields worldwide range from 0.8 to 1.5 tons per , with the global figure standing at 1.3 tons per in 2024 despite a 10.7% decline from the previous year due to variability. In intensive systems, such as those in parts of and , yields can reach up to 2.5 tons per under optimal management. These productivity levels underscore lentils' role as an efficient pulse crop in diverse agroecosystems. International trade in lentils is robust, with exports totaling 4.31 million metric tons in 2022, representing about 65% of global production. dominates exports, accounting for roughly 50% of the world market and shipping 2 million metric tons in 2023 alone. Primary importing nations include , with 0.66 million metric tons, and , with 0.52 million metric tons in 2022, highlighting the crop's importance in addressing protein needs in densely populated regions. Emerging trends point to sustained growth in demand fueled by the global shift toward plant-based diets, where lentils serve as a versatile, nutrient-dense protein alternative. Concurrently, introduces risks, including erratic rainfall and temperature extremes that could reduce yields by up to 10-20% in vulnerable areas without adaptive measures.

Leading Producers

Canada remains the world's leading producer of lentils, with output reaching approximately 1.7 million tonnes in 2023, primarily consisting of green and large-seeded varieties grown in the prairie provinces. Saskatchewan accounts for over 87% of this production, benefiting from the region's suitable semi-arid climate and extensive research support from organizations like the Saskatchewan Pulse Growers, which provide funding for crop improvement and market development. Government initiatives, including federal agricultural programs, have bolstered yields through advanced breeding and pest-resistant varieties, enabling Canada to supply about 25-30% of global exports despite domestic consumption being relatively low. In 2024, production increased to 2.4 million tonnes. India ranks second in lentil production, harvesting around 1.56 million tonnes in , with red lentils dominating output in states like , which contributes nearly 40% of the national total due to its fertile black soils and rabi-season . High domestic demand drives nearly all production toward internal markets, supporting for over a billion , though imports supplement shortages during lean years. Challenges such as recurrent droughts in rainfed areas like have prompted government interventions, including subsidies for drought-tolerant seeds and under the National Mission, aiming to boost from the current average of about 900 kg per . Turkey and Australia are significant contributors, with Turkey producing about 0.47 million tonnes in 2023 and Australia approximately 1.4 million tonnes, focusing on and varieties for markets. Turkey's , centered in the southeastern regions, supports exports to and , with lentils comprising over 85% of output and government seed distribution programs aiding farmers in expanding acreage. Australia, with major growing areas in and , exports primarily to and , leveraging research grants from the Grains Research and Development Corporation to address challenges like deficits and seasonal labor shortages during . In 2024, Australia's rose to 1.6 million tonnes. Both countries benefit from trade agreements that facilitate access to high-demand regions, though Australia's exports faced temporary hurdles from import tariffs in prior years.

Human Uses

Processing Methods

Lentils undergo initial processing through and to remove impurities such as debris, stones, and immature seeds, ensuring product quality and safety for further handling. This is typically achieved using methods like sieving to separate by size and air classification to remove lighter foreign matter based on . Dehulling and splitting follow to prepare lentils for , particularly for varieties like red lentils that are well-suited to these processes due to their thinner seed coats. Dehulling involves , where the outer is removed using pearlers or rollers, often after a preconditioning step like tempering with to loosen the coat. Splitting then divides the dehulled cotyledons into halves, producing products like commonly used in , while soaking during this stage helps reduce anti-nutritional factors such as and . After dehulling, lentils are dried to optimal levels for and milling, typically reduced to 10-14% using sun in traditional settings or mechanical hot-air dryers in operations to prevent microbial and maintain viability. Milling then grinds the dried, dehulled lentils into , known as besan, using or pin mills, which facilitates applications in and by yielding a fine with improved . Preservation methods extend and enable diverse product forms, including where cooked lentils are sealed and heat-processed at 116-125°C in retorts to achieve sterility, freezing to halt enzymatic activity, and or barrier to minimize oxygen exposure. Additionally, involves soaking and controlled to produce nutrient-enhanced lentil sprouts, a traditional that also diminishes certain anti-nutritional compounds.

Culinary Applications

Lentils serve as a versatile staple in cuisines worldwide, valued for their ability to absorb flavors and provide hearty textures in both hot and cold preparations. They are commonly boiled to create creamy bases for soups and stews, where their quick cooking time—typically 20 to 40 minutes depending on the variety—allows for efficient meal assembly. In many cultures, lentils are simmered with aromatics like onions, , and spices to form comforting dishes that highlight their earthy taste. In , lentils form the foundation of a classic featuring green lentils cooked with carrots, , and or herbs for a robust, warming . Similarly, in cooking, dal tadka showcases split red lentils tempered with , , and in a spiced topping, resulting in a creamy, aromatic often served with or . These techniques emphasize lentils' role in building depth through gradual flavor infusion. For salads and side dishes, lentils offer a protein-rich element in cold or room-temperature presentations. The Lebanese combines brown lentils with rice and caramelized onions, creating a simple yet satisfying that balances nutty grains with sweet, crispy toppings. This preparation highlights lentils' firmness when cooked , making them ideal for textured salads. and expand lentils' applications in innovative ways. In the , red lentil variants of patties, blended with herbs, onions, and spices before baking or frying, provide a lighter alternative to chickpea-based versions, often served in wraps or as appetizers. Regionally, lentils anchor distinctive dishes across continents. Ethiopian misir wat is a spicy red lentil stew simmered with spice blend, onions, and tomatoes, traditionally eaten with flatbread for a bold, fiery meal. In , lentejas soup features brown lentils cooked with chorizo, potatoes, and cilantro in a tomato-based , offering a comforting, everyday family dish. Italian pasta e lenticchie integrates small pasta shapes directly into a lentil with soffritto vegetables and tomato, yielding a thick, one-pot that thickens as it simmers. Split lentils, often processed for quicker cooking, are frequently used in these recipes to enhance efficiency.

Nutritional Value

Lentils are a nutrient-dense , providing a balanced profile of macronutrients in their dry form. Per 100 grams of raw lentils, they contain approximately 24.6 grams of protein, which is notably rich in the compared to other plant proteins. The carbohydrate content is around 63 grams, including 10.7 grams of , predominantly soluble and insoluble types that contribute to and digestive health. Fat levels are low at about 1.1 grams, resulting in a density of 352 kcal per 100 grams. In terms of micronutrients, raw lentils are an excellent source of , offering about 479 micrograms per 100 grams, which meets approximately 120% of the daily value (). They also provide significant iron at 6.51 milligrams per 100 grams (around 36% ), along with at 955 milligrams, supporting oxygen transport and cardiovascular function. Additionally, lentils contain antioxidants such as polyphenols, including procyanidins and flavanols, which exhibit strong free radical-scavenging properties. The nutritional profile of lentils contributes to several health benefits, particularly in metabolic regulation. Their low (GI) of 22–37 for cooked varieties helps control blood sugar levels by slowing glucose absorption, making them suitable for . Soluble in lentils, such as and gums, binds to acids and in the digestive tract, promoting their and thereby reducing serum levels. Despite these advantages, lentils can cause digestive discomfort due to oligosaccharides like , which ferment in the gut and lead to and gas. This effect can be mitigated through soaking, which leaches out up to 40% of these compounds, or thorough cooking, which further breaks them down via enzymatic and thermal processes.
Nutrient (per 100g raw lentils)Amount% Daily Value*
Calories352 kcal-
Protein24.6 g49%
Carbohydrates63 g23%
10.7 g38%
1.1 g1%
479 µg120%
Iron6.5 mg36%
955 mg20%
*Based on a 2,000-calorie ; sourced from USDA FoodData Central.

Breeding and Research

Genetic Development

The genetic of lentils traces back to early human interventions during during the period, around 8,000 BCE (10,800–8,500 cal BP), in the , particularly in the regions of modern-day , , and . Farmers selectively propagated wild Lens culinaris subsp. orientalis for advantageous traits, including non-shattering pods (pod indehiscence) to prevent loss during and larger size to boost and ease of . These selections, driven by repeated harvesting and replanting of desirable plants, marked the transition from wild gathering to cultivated , with archaeological evidence from sites like Yiftah'el in confirming cultivation by 10,100–9,700 calibrated years . Hybridization techniques within the genus have been central to refining key traits and introducing variation. Breeders conducted controlled crosses between cultivated lentils and closely related or landraces to enhance pod indehiscence, ensuring better seed retention, and to modify seed color for market preferences and processing qualities. Such inter- and intra-specific hybridizations, often using and methods, successfully transferred monogenic traits while mitigating fertility barriers, resulting in stable recombinant lines that broadened the genetic base for and yield stability. Germplasm collections have underpinned these efforts by safeguarding diverse genetic resources. The International Center for Agricultural Research in the Dry Areas (ICARDA), a center established in 1977, maintains one of the world's largest lentil collections, with over 11,000 accessions of cultivated and wild forms preserved for breeding use. These repositories enable systematic evaluation and incorporation of traits from global landraces, supporting ongoing selection for in marginal environments. Wild relatives within the have occasionally been referenced in crosses to infuse novel alleles, though primary focus remains on cultivated diversity.

Modern Improvements

Since the 2010s, (MAS) has advanced lentil breeding by leveraging quantitative trait loci (QTL) mapping to enhance traits such as and resistance to Ascochyta blight. For , a 2016 study identified 18 QTLs controlling root and shoot traits like root , lateral root number, and root-shoot ratio in a recombinant inbred line under progressive stress, with key QTLs on linkage groups III, IV, and IX explaining up to 28.9% of phenotypic variance and linked to SNP markers for potential MAS deployment. Similarly, for Ascochyta blight resistance, QTL mapping in 2012 and 2016 pinpointed multiple loci on linkage groups 1, 2, 4, 5, and 9, accounting for 3-47% of variance, with SNP-based markers like AB_IH1 achieving over 85% accuracy in predicting field resistance across diverse . These MAS approaches have facilitated the of resistance genes from wild relatives into cultivars, improving overall without relying solely on phenotypic selection. The 2019 whole-genome of lentil has revolutionized genomic research, providing a high-quality reference that spans approximately 3.7 Gb across seven chromosomes and enables precise identification for targeted editing. This , developed using PacBio and technologies on the CDC Redberry , has identified 58,243 protein-coding genes, facilitating studies and the application of / for trait enhancement. Specifically, it supports edits to boost protein content by targeting genes involved in biosynthesis pathways, such as those regulating sulfur-containing , and to reduce anti-nutritional factors like and inhibitors, which hinder nutrient absorption. Although lentil-specific applications remain nascent, successes in related demonstrate the potential for knockouts that increase protein quality by 10-20% while minimizing anti-nutrients, paving the way for bioengineered varieties with improved digestibility. Recent 2025 studies have applied / to enhance lentil resistance to Ascochyta blight and develop herbicide-tolerant varieties, accelerating breeding for tolerance. Biofortification efforts in lentil focus on breeding varieties with elevated iron and zinc levels to combat micronutrient malnutrition in developing regions, where over 2 billion people face deficiencies. Through conventional breeding and association mapping, researchers have identified alleles associated with up to 20% higher grain iron (reaching 119 mg/kg) and zinc (75 mg/kg) content, as demonstrated in diverse germplasm panels from South Asia and the Middle East. Dual-biofortified lines, such as those released by the International Center for Agricultural Research in the Dry Areas (ICARDA), enhance bioavailability by co-increasing promoters of mineral transporters while reducing phytic acid. These varieties target regions like sub-Saharan Africa and South Asia, where lentils form a dietary staple, supporting global goals to alleviate hidden hunger by 2030. Sustainability research in lentils emphasizes systems and the development of climate-resilient varieties to ensure production stability amid projected 2050 climate shifts. studies show that pairing lentils with cereals like or in Mediterranean increases land equivalent ratios by 15-30%, enhances , suppresses weeds by up to 50%, and maintains yields under water-limited conditions without significant competition for resources. For instance, lentil- in sustains lentil productivity while improving and profitability, reducing reliance on synthetic inputs. Climate-resilient varieties, bred via genomics-assisted selection, incorporate traits for heat and , with projections indicating that such adaptations could offset 20-40% yield losses under RCP 8.5 scenarios by 2050, particularly in rainfed systems covering 80% of global lentil acreage. These strategies promote agroecological intensification, bolstering in vulnerable regions.

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